Hoya acanthominima Kloppenburg, Mendoza & Ferreras 2013 ISSN 10554564 Described in New Hoya Volume 1-1, rare-hoyas.com

Calyx with ovaries on a small section of the pedicel enlarged ca. 27x. The pedicels are curved of various lengths 1.0 cm -1.9 cm long and 0.3 cm diameter, sepals are sub triangular, granulate outside, edges ciliate, with a basal overlap of 1/2 , they are 0.07 cm long and 0.07 cm at the widest. Ovaries are columnar glabrous 0.08 cm long and base pair 0.03 cm wide.

143

Hoya alwitriana Kloppenburg, Siar, Guevarra & Carandang 2012 Asklepios 114

Calyx: and ovaries: sepals are narow triangular 0.15 cm long 0.09 cm at widest, ligules are present, outside are granulose, edges ciliate, inside glabrous. Ovaries: narrowly columnar 0.15 cm tall and 0.09 cm at base, glabrous.

144

Hoya andalensis Kloppenburg 2005 Described in Fraterna 18 #1: 2005, 1-5

Pedicels: curved, glabrous, terete, lt. green, 0.09 cm. in diameter. Side view of the pedicel, calyx and ovaries enlarged about 16X. All green in color as shown. Cluster with pedicels of various lengths form a geotropic concave cluster.

Another close-up of the calyx and ovaries enlarged about 16X. The area near the calyx is granulose as is the outer surface of the sepals. The ovaries are cupped on top 0.11 cm. tall and the base pair are 0.10 cm. wide, surface is glabrous. Color as shown clear lime green.

Top view of the calyx enlarged about 16X. Calyx is small. Sepals are membranous with the central area thickened a little. Edges with an occasional cilia, rudimentary ligules present. The sepals do not reach the corolla sinuses. 0.10 cm. long and 0.16 cm to the center. 0.10 cm. at the widest.

145

Hoya anncajanoae Kloppenburg & Siar 2007 Described in Asia Life Sciences 17(1) 62-66, 2008.

Calyx side view enlarged as above, outside hirsute, ovate, 0.15 cm. long and 0.17 cm. widest, inside slick, glabrous. Ovaries bottle shaped 0.15 cm. tall and base pair 0.12 cm. wide.

Top inside view of the calyx and ovaries again enlarged about 8X. sepals overlap about 1/3 are ciliate and slick glabrous inside, ligules are present. apex to corolla sinus 0.10 cm. so they are short of reaching the corolla sinuses. Shape sub-ovate.

146

Hoya apoensis Kloppenburg & Siar Described in Asia Life Sciences 2010.

View of the calyx and ovaries enlarged about 8X. Sepals are broadly triangular with narrowing rounded apex. Glabrous on both surfaces, ligules present. 0.11 cm. long and 0.14 cm. at the widest above the basal overlap, edges are membranous with ragged edges. Ovaries are columnar, glabrous with darker bases, 0.15 cm. tall and base pair 0.10 cm. wide.

View of the flower back with pedicel and calyx attached. Sepals do not reach the corolla sinuses. Base of calyx in expanded, surface rough granulate 0.18 cm. in diameter. Corolla is revolute and outside glabrous.

147

Hoya apoensis subsp. sagittarius Kloppenburg & Siar 2010

Published in Fraterna 23/4 2010

Picture enlarge about 11X, bottom of calyx bulbous.

Calyx: small triangular sepals, triangular to sub-obtuse at apex. 0.11 cm long, 0.09 cm widest at base, glabrous no overlap, ligules. Enlarged about 12X. Ovaries: dome shape tapering to apex, glabrous, 0.15 cm long, 0.10 cm wide at the base.

Outside view of the flower showing pedicel and calyx attached.

148

Hoya archboldiana Norman 1937 Described in Brittonia 2 (1937) 328. C. Norman. Hoya archboldiana C. Norman. "Calyx equally divided, small, lobes ovate acute around 4 mm."

Top view of the calyx and Ovaries. Calyx apex is rounded overlap slightly at base and ligules are present. the base ring of ovaries is pronounced and thickened.

Pedicel: large fleshy green. 8.0 cm. long x 0.15 cm. diameter. glabrous. Calyx: Sepals convex, rolled back, rounded but abruptly acute, with a few cilia on the edges. 0.37 cm. long, overlapping 1/3 at base. 0.27 cm. at the widest, base 0.20 cm. no glands observed (auricles). Ovaries: 0.35 cm. long, base 0.20 cm. widest.

149

Hoya attenuata Christophersen 1935 Described In Bulletin of the Bishop Museum “Samoan Flowering Plants” 128 (1935) 187. Christophersen.

Outside view of the flower enlarged about 8X. The pedicel is very fine, 2.3 cm. long, terete, glabrous 0.04 cm. in diameter. Calyx is small and the sepal apexes do not reach the corolla sinuses. Corolla outer surface is glabrous.

Calyx (and pedicel) side inside view enlarged about 8X. Sepals are ciliate, surfaces are glabrous. 0.10 cm. long 0.02 cm. to the center from the outer apex. There are small dark ligules present at the bases. Apex is fairly acute to slightly rounded. Ovaries are domed 0.10 cm. tall and the base pair 0.08 cm. wide.

150

Hoya aurigueana Kloppenburg, Siar & Cajano ISSN #1655-3179

Side view of the pedicel, calyx and ovaries enlarged about 10x. Sepals do not reach the corolla sinuses, outside granulate, inside slick and glabrous with dark ligules, 0.20 cm long and 0.12 cm wide at the base, apex is sub acute. Ovaries: domed 0.10 cm tall and base pair 0.13 cm wide, glabrous, apical area notched rounded.

151

Hoya bandongii Kloppenburg & Ferreras 2011 Published in Fraterna 24 #4:9-13, 2011

Calyx: side view enlarged 15X. Sepals are ovateelliptic, ciliate otherwise glabrous 0.19 cm long x 0.12 cm at the widest. Apex is narrowly obtuse.

152

Hoya bebsguevarrae Kloppenburg & Carandang 2013 ISSN # 2329-7336

Calyx: outside surface enlarged ca. 22x. Surface is granulate, thickened centrally, pedicel not expanded at base. 0.15 cm long x 0.10 cm. widest, apex sub-acute.

Calyx inside view enlarged as above. Inner surface is slick and glabrous, base with very small ligules, overlap ¼ at base, outer apex narrowly obtuse.

Ovaries: Columnar, glabrous, 0.12 cm tall and base pair 0.10 cm wide.

153

Hoya benguetensis Schlechter 1906 Described in Philippine Journal of Science Philippine Journal of Science. 1 (1906) 301. Dr. Schlechter. "with the segments of the calyx ovate obtuse, glabrous, barely 0.2 cm. long."

154

Hoya benstoneana Kloppenburg, Siar, Guevera & Carandang Described in Journ. Nat. Studies 11(1-2):36 30 Jan 2013 e-published

Side view of the calyx enlarged about 13X. Base a little enlarged, outer surface granulate glabrous. Sepals small.

Top view of corolla enlarged about 16X, sepals are triangular, overlap a little at the base, 0.12 cm. long, 0.12 cm. at the widest, edges ragged, no ligules. Ovaries short dome shaped, glabrous, 0.13 cm. tall and base pair 0.12 cm. wide.

Outside view of a flower enlarged about 14X, to show the calyx (below center) the sepals are very small, outside granulate inside slick glabrous, they do not quite reach the corolla sinuses.

155

Hoya bifunda Kloppenburg, Siar, Cajano, Guevarra & Carandang 2013

ISSN # 2329-7336

Calyx: side view enlarged ca. 20X. The sepals are ovate triangular, ciliate with a sub obtuse outer apex, small overlap at the base.

Calyx inside view enlarged ca. 30X, the surface is glabrous, ligules are present at the base sinuses. Sepals are 0.11 cm long and 0.08 cm at the widest, with a few cilia. Sepals do not reach the corolla sinuses. Ovaries: are dome shaped, 0.09 cm tall and base pair is 0.05 cm wide; glabrous.

156

Hoya bifunda subsp. integra Kloppenburg, Siar, Cajano, Guevarra & Carandang 2013

Calyx: side view enlarged ca. 40X. Sepals are granulate outside slick glabrous inside, triangular in shape with a small basal overlap apices are sub acute.

Calyx side view enlarged ca. 34X. Sepals are 0.11 cm long and 0.07 cm at the widest, ligules are present at the sinuses inside. The ovaries are small dome shaped, glabrous, 0.08 cm tall and base pair 0.05 cm wide.

157

Hoya bilobata Schlechter 1906 Described in Philippine Journal of Science, i, Supplement Schlechter.

(1906) 301-302. R.

"with the segments of the calyx oblong obtuse, glabrous, longitudinally scarcely attaining 0.1 cm."

Top view of the calyx. Two from the same flower cluster show some variation. The one on the left seems more like Schlechter's drawing, longer, more elliptic. There are ligules at the sinus bases. The sepals overlap slightly. Sepala 0.07 cm. long, 0.05 widest.

The side view of the calyx and ovaries; a small portion of the pedicel. Compare all these photos with Dr. Schlechter's drawing and with the description. The outer surface and pedicel are granulose-punctate, inside smooth and glabrous. Ovaries tapering to the apex but do not seem as pointed as in Schlechter's drawing.

158

Hoya bilobata from Herbarium sheet (UC) #30003.

Hoya bilobata from Herbarium sheet (UC) # 4936.

159

Hoya bicolensis Kloppenburg, Siar & Cajano Described in Jour. Nature Studies 11 (1-2):39, 2013e-published.

Outside view of the corolla with pedicel and calyx attached about 8X. The calyx is linear and reaches the corolla sinuses. Corolla outside glabrous, at anthesis flat to slightly cupped. Sinus - sinus 0.20 cm. Sinus - apex 0.32 cm. Sinus - center 0.20 cm. Apex - center 0.44 cm. Widest 0.28 cm. Pedicel and calyx enlarged about 8X. Pedicel is fine, strict, glabrous, terete 0.06 cm. in diameter, 1.3 cm. long, uniform yellow color, bronze rose at base. Calyx Apex - base 0.12 cm. Apex - center 0.20 cm. Widest 0.12 cm.

Inside view of the calyx enlarged about 8X. 1 ligule detected and hard to see. Ovaries are dome shaped 0.07 cm. tall and base pair 0.07 cm. broad, glabrous.

160

Hoya bicolor Kloppenburg 2003 Described in Fraterna 16/1: 2003.

Outside view of the corolla, pedicel and calyx. The pedicel is straight, terete, puberulent, short 1.45 cm. long. The sepals do not reach the corolla sinuses, are short triangular, with ciliate edges, base somewhat granulose. The corolla outside surface is glabrous with some punctations, deeply cut. Sepal apex to the sinus 0.05 cm.

Pedicel, calyx and ovaries, side view enlarged about 8X. The ovaries narrowly domed, glabrous and waxy, 0.09 cm. tall and the base pair is 0.08 cm. wide. Calyx lays flat and outspread.

Calyx top view. Sepals are ciliate, glabrous inside wide near the base and tapering to a narrow obtuse apex, barely overlapping, no ligules are present. Sepals 0.14 cm. long, 0.11 cm. widest. from apex to center 0.18 cm.

161

Hoya blashernaezii Kloppenburg 1999 Described Fraterna 12/1:1999, 9-13. Top vied of the calyx. Sepal bases overlapped a little, edges ciliate narrowing to rounded apices.

Calyx: sepals elongate triangular 0.15 cm. long, widest 0.10 cm., thin membranous, when dry yellowish in color, ciliate, ligules present at the base, glabrous, shiny inside. Ovaries: 0.10 cm. tall base 0.08 cm wide, glossy yellow, glabrous.

Pedicel and calyx on the outside of the corolla. Sepal apices do not reach the corolla sinuses. Corolla campanulate, this surface granulose punctate. Pedicels: 2-2.5 cm. long, filiform, glabrous, terete, pale yellow. Umbel of 24 flowers, open only one day. In this respect it is similar to H. camphorifolia Warburg, also a Philippine hoya species.

Outside surface of the corolla flattened. Calyx shows up well as a different color, sepals are short, triangular, and do not reach the sinuses of the corolla. Coronal lobes are deeply cut broadening outward to an apiculately dented round sided apex.

162

Hoya brittonii Kloppenburg 1992 Described in Fraterna 1992/4: I. "Calycis segmenta ovata. margine ciliata quam corolla paulo breviora."

Pedicel, calyx and corolla. The sepals do not reach the corolla sinuses, are ciliate, overlapped at the base and with rounded apices. Pedicel: curved variable in length (forming flat topped umbels), about 1 cm. long, 0.05 cm. in diameter enlarging toward the calyx, minutely puberulous or densely granulose (actually with very broad based very short sharp apexed cells. Calyx segments: ovate to somewhat broadly triangular, margins ciliate 0.08 x 0.08 cm. with ligules at the sinuses. Top inside view of the calyx. Here again note the ciliate edges and rounded apices also the overlap at the base and the presence of ligules, 2-2-1-1-1.

163

Hoya buotii Kloppenburg 2002 Described in Fraterna 15/4: 2002, 1-5.

Calyx top view. Sepals broad based overlap about 10%, ligules present 0.02 - 0.03 cm. long, apices acute, inside waxy glabrous, edges ciliate, apex obtuse. Center to apex 0.40 cm. sepals 0.30 cm. long and 0.20 cm. wide.

Pedicel attached to the corolla. The sepals do not reach the corolla sinuses, are glabrous but edges ciliate and central thickened surface granulose. Corolla surface outside glabrous with some punctations, waxy.

164

Hoya burtoniae Kloppenburg 1990 Described in Hoyan 12(1:2):I (1990). "Calycis indivisis destituta ligulae; segmenti vix 0.12 cm longi, ovati, obtusi, hirsuti."

Calyx segments individual, ovate, obtuse velutinous, about 0.12 cm. long with no ligules.

Pedicels 1.2 to 2.2 cm long, hirsute, curved.

165

Hoya butleriana Kloppenburg, Siar, Guevarra & Carandang Asklepios 2013

Calyx: inner view, sepals 0.20 cm long, 0.14 cm at the widest, are narrowly triangular, slight basal overlap, apices sub obtuse, surfaces glabrous, center slightly thickened, outside granulose. This and picture below enlarged about 16x.

Side view of the calyx with ovaries. Ovaries shortly domed, glabrous, 0.12 cm tall and base pair 0.13 cm wide.

166

Hoya cagayanensis Burton 1987 Described in The Hoyan 8/2 (1987) b.

Calyx including ovaries and portion of pedicel. Ovaries: dome shaped 0.15 cm. Tall 0.13 cm wide at base of the pair. Calyx: with ligules one at each sinus. Sepals do not reach the corolla sinuses, apex obtuse, shape ovate, edges ciliate.

Top view of the calyx with ovaries attached. Sepals: overlap at base about 1/3. 0.23 cm. long, 0.18 at the widest. Base to sinus 0.10 cm.

167

Hoya cardiophylla Merrill 1920 Described in Philippine Journal of Science 17 (1920) 310. E. D. Merrill. "Calyx-lobes triangular-ovate, acute or obtuse, 1 mm long."

Top view of the calyx (one sepal missing). Edges a little ciliate, small ligules near sinuses. Ovaries small dome shaped 0.08 cm. tall and base pair 0.11 cm. at the widest

Pedicel, calyx & ovaries side view. Pedicels 2.0 cm. x 0.10 cm. in diameter, terete, glabrous, flowers pale yellow. Calyx stubby, granulose outside, glabrous inside, small ligules 0.13 cm. long, widest 0.15 cm.

168

Hoya carmeleae Kloppenburg, Siar & Ferreras Published in Fraterna 23 #4 2010

Pedicel: and calyx enlarged ca. 31X, pedicel terete, glabrous, strict, 1.6 cm long and diameter 0.06 cm. Calyx: lobes near linear, do not nearly reach the corolla sinuses, glabrous in and out, outside surface granulate, sepals 0.10 cm long 0.06 cm widest, slight basal overlap, no ligules seen. Base bulbous. Ovaries: columnar, glabrous 0.10 cm tall base pair 0.07 cm. wide.

169

Hoya carnosa R. Brown 1817 Described in Equitis Systema Vegetabilum (1817) 50-51. Hortus Kewensis ed. alt. 2 (1811) 84. Asclepias carnosa. "Calyx minutus." "Calyx rotate of 5 parts of the same reddish color as the peduncle, appressed to the corolla."

Calyx: 0.28 cm. long, 0.28 at widest. Sparsely ciliate with ligules present.

Pedicel: 0.13 cm. in diameter strict, terete, with a few sparse hairs pointing apically, most bent toward pedicel.

170

Hoya caudata J. D. Hooker 1883 Described In Flora of British India 4 (1883) 60.i

Bock view of the calyx with pedicel attached, enlarged about 8X. Sepals are narrowly triangular. Outer surface granulose, glabrous. Inside slick smooth.

Front view, with ovaries showing. Sepals; Apex - base 0.20 cm. Apex - center 0.27 cm. Widest 0.11 cm. Ovaries: Column shaped 0.14 cm. tall and 0.11 cm. wide at the base.

Calyx on outside surface of the corolla enlarged about 8X. The sepals are triangular 1/2 way the corolla sinuses. There are bumps between each sepal on the corolla, which is an unusual structure I have not observed on any other hoya species. Here the tow in the lower right hand area are bronze bumps clearly visible.

171

Hoya celata Kloppenburg, Siar, Mendoza, Cajano, Guevarra & Carandang Described in Asklepios 116:42. 2013

Pedicel with Calyx and ovaries: Enlarged about 10X. Pedicel terete, glabrous, curved 4.0 cm. long 0.07 cm. in diameter, very slight enlargement at calyx base. Calyx: sepals do not reach the corolla sinuses, tapered to rounded apex sides undulant with an occasional cilia, base 1.3 overlap, ligules present. Sepals 0.33 cm. long and 0.20 cm. at the widest.

172

Hoya celsa Kloppenburg, Siar, Guevarra, Cajano & Carandang 2012

Flower outside view with pedicel and calyx, enlarged ca 18x. Corolla outside is glabrous, sepals are elliptic and ¾ way to the corolla sinuses.

Calyx: side view enlarged ca. 20x, ligules are present, 2-1 pattern. Sepals glabrous, 0.11 cm long and 012 cm at the widest, apex narrowly obtuse, a little basal overlap. Ovaries: here shriveled, columnar 0.14 cm long and base pair 0.07 cm, glabrous.

173

Hoya chlorantha Rechinger 1908 Described in Repertorium Specierum Novarum 5 (1908) 131. (Fedde) “Plantae Nova Pacificae II.” Dr. K. Rechinger. 16.

Outside view left and inside view right of the calyx enlarged about 8X. Sepals are ciliate otherwise glabrous Apex - base 0.20 cm. Apex - center 0.25 cm. Widest 0.15 cm. Ovaries are glabrous, domed 0.15 cm. wide at the base.

174

Hoya chunii Li 1984 Described in Botanical Research Bulletin 4 (1984) 121. P. T. Li. "Calycis segmenta ovalia obtusissima, subverruculoso-punctata, quam corolla multo breviora." Top view of the calyx here. The sepals do not quite each the sinus of the corolla. The apical areas essentially rounded although as in all species there are slight variations present in the length, width and shape. I did not make a notation on ligules at the sepal bases. Calyx: cupped, lobes not quite reaching the corolla sinus, stubby, obtuse, thick appearance. diameter 0.42 cm.

Side view of the pedicel and calyx including the ovaries. The sepals are relatively small with rounded apex, also the ovaries are short. Pedicel is round, with a few scattered hairs but essentially glabrous, enlarging slightly toward the calyx, and straight. Pedicel: glabrous, terete, strict. diameter 0.95 cm. Ovaries: very short 0.17 cm. tall, 0.11 cm. at base.

175

Hoya citrina Ridley 1922 Described in Journal of the Royal Asiatic Society, Straits Branch 86 (1922) 300. H. Ridley. Calyx not mentioned in the type description.

Top view of the calyx. A not very clear photo. Note that the sepal apex is broadly rounded with about 1/3 overlap with very large ligules at the sinus.

Side view of the pedicel and calyx. All parts are glabrous. The outer surface of the calyx is granular and the pedicel is lenticulate. Sepals are relatively short with outer apex obtuse.

176

Hoya clemensiorum Green 2001 Described in Fraterna 14/3:2001,12-15. "calyx 4 mm. in diameter."

Calyx is short outside coarsely granular inside glossy glabrous diameter 0.48 cm.; sepals have very little overlap 0.15 cm. long and 0.10 cm. at the widest, base glands not visible. Ovaries are short domed pair, glabrous 0.07 cm. tall and base of pair 0.10 cm. Top view of calyx. The sepals essentially ovals, fairly flat and relatively small.

the are lay are

Side view of the pedicel, calyx and ovaries. The pedicel is very short, pubescent, 2.10 cm. long and 0.10 cm. in diameter, pale yellow.

177

Hoya cominsii Hemsley 1890 Described in Annales of Botany (1890) 505-506. W. B. Hemsley. "Calyx of many segments ovate oblong very obtuse."

Side view of H. sp. DAV 819. The sepals here and on IML 457. Both are membranous with linear vessels visible, with rounded obtusa apex and ligules present at the sinus. Calyx: lobes 0.16 cm. long, widest ca. 0.17 cm. Ligules long 2-1-2-1-1.

Side view of the pedicel, calyx and ovaries of H. sp. DAV 819. Here the ovaries are slightly shorter than in IML 457 the apex more rounded not somewhat blunt as in the later. The whole structure is glabrous but the pedicel and outside calyx surfaces are a little granulose and the round pedicels are lenticular. Pedicel : DAV 819 0.10 cm in diameter glabrous, terete, straight, IML 457 same but 0.12 cm. diam. Ovaries: 0.16 cm. tall, base 0.11 cm.; IML 419, 0.19 cm. tall, base 0.15 cm.

178

Hoya corazoniae Kloppenburg, Siar & Ferreras Published in Fraterna 23 #4, 2010

Calyx and ovaries enlarged ca. 31X. The sepals are small do not reach the corolla sinuses and are membranous, 0.09 cm long and wide, overlap about 1/3, glabrous on both surfaces, ovate apex obtuse. Dark ligules are present. Ovaries: short dome shaped, glabrous 0.06 cm tall and base pair 0.08 cm wide.

179

Hoya cumingiana Decaisne 1844 Described in Decandolle, Prodramus Syst. Veg. 8 (1844) 636. Decaisne. Calyx not mentioned in the type description.

Top view of the calyx. Ligules at the base of sepals are plainly visible (lighter color). The red ovary apices are also shown. Calyx: outside granulose with some scattered hair cells, edges jagged 0.25 cm. long, 0.13 cm. wide. Apex acute, ligules present.

Calyx on back of calyx Sepals do not reach the corolla sinuses. There are scattered hair cells here on this outside surface. Ovaries: long columnar, red tipped. 0.20 cm. tall and 0.11 cm. wide base pair.

180

Hoya cystiantha Schlechter 1856 Described in Flora of Indiae Batavae 2 (1856) 516. H. Zollinger & F. A. W. Miquel. Calyx not mentioned in the type description.

Top view of the calyx. The sepals have little or no overlap at the base, are rounded at the apex. Notice the fine even ciliation. The ligules at the base are very visible and easily observed one at each sinus. The ovary base is symmetrically circular. Calyx: lobes finely and evenly ciliate, cupped, rather thick in the central portion. Lobes not fused or overlapped at the base with definite easily observed ligules at each sinus, 0.03 cm. long.

Side view of the pedicel, calyx, and ovaries. Sepals are cupped inward, rather thick and finely ciliate. The pedicel (only a portion shown here) is curved enlarging slightly as it nears the calyx, with parallel lenticular longitudinal striations. The ovaries are columnar bottle shaped.

181

Hoya dennisii P. I. Forster et D. J. Liddle 1993 Described in Austrobalia 4(1) (1993) 51-52. P. I. Forster & D. Liddle. "Sepals lanceolate, c. 0.12 cm long and 0.06 cm wide, glabrous."

Photomicrograph of calyx attached to the corolla. Sepals are very small, narrowly elongated with rounded apices and not overlapped much at the base. Outside granulose; inside glabrous. Lobes 0.9 cm. long. Pedicel is very filiform, curved, glabrous 3 mm. in diameter, 1.9 cm. long expanding slightly close to the calyx, rose brown in color.

182

Hoya desvoeuxensis T. Green & Kloppenburg Asklepios 112, 2011

Calyx enlarged about 16X. Sepals are small in relation to the corolla. With occasional cilia, triangular with rounded apex, glabrous on both surfaces, 0.20 cm long and 0.16 at the widest, little basal overlap, ligules present. Ovaries columnar 0.25 cm tall and base pair 0.13 cm wide.

183

Hoya dischorensis Schlechter 1913 Described in Botanische Jahrbücher 50 (1913) 116. R. Schlechter. "Calyx segments ovate, somewhat obtuse, glabrous, much smaller than the corolla."

Top view of the calyx. Calyx is definitely cupped around the ovaries. Surface is glabrous. Sepals only slightly overlapped at the base, small ligules present. 0.24 cm. long and widest 0.17 cm.

Side view calyx and ovaries.

of the

Ovaries are domed 0.21 cm. long, base pair 0.16 cm. wide; with little nipple like apical ends. Calyx lobes outspread. Sepals outside glabrous except for a few cilia at margins.

184

Hoya eburnea Kloppenburg, Guevara & Carandang 2013 ISSN # 2329-7336

Calyx: side view enlarged ca. 20x, sepals are sublinear do not reach the corolla sinuses, glabrous granular outside inside slick glabrous; 0.15 cm long and 0.06 cm wide at the base, overlap slightly, no ligules observed. Ovaries: columnar, cm wide, glabrous, 0.11 cm tall and base pair 0.06 cm wide.

185

Hoya eitapensis Schlechter 1909 Described in Botanische Jahrbücher 50:109. R. Schlechter. "Calyx segments ovate-triangular, somewhat obtuse, minutely and sparingly puberulous and ciliate, much smaller then the corolla."

Pedicel and calyx. The sepals are small triangular but with rounder apices. The pedicel relatively stout, round, glabrous, pale lemon yellow in color. Pedicel: glabrous 1-2 cm. long, terete, 0.10 cm. in diameter, curved to form a flat faced cluster. Calyx: small, triangular with rounded apices, 0.30 cm. in diameter, sepals 0.12 cm. x 0.10 cm. single and double ligules present. Ovaries: 0.12 cm. tall, base pair 0.10 cm wide, tapering dome shaped.

Outside of flower with the pedicel and calyx intact. The sepals are small and do not nearly reach the corolla sinus. The pedicel is glabrous, and round not expanding much as it approaches the calyx.

186

Hoya erythrina Rintz 1978 Described in Malayan Nature Journal 30 (1978) 501. R. L. Rintz. Calyx not mentioned in the type description.

Top view of the calyx. The calyx is glabrous, granulose outside, with a rough edge, membranous 0.46 cm. in diameter. Sepals are 0.21 cm. long and 0.13 cm. at widest portion; they are narrowly triangular with an acute apex. One large ligule at the base of each sinus, they are large and pronounced.

Side view of the pedicel, calyx and ovaries. The pedicel is rough (silicate), glabrous, filamentous, green ca. 2 cm. long and 0.08 cm. in diameter.

187

Hoya estrellaensis T. Green & Kloppenburg Asklepios 114, 2012

Calyx: top view enlarged about 19X. Sepals are ovate-elliptic. Outer apex obtuse, ¼ overlap at the base, ligules are present, 0.19 cm long. Widest 0.15 cm Outside rough granulate. Sepals concave inside, do not quite reach the corolla sinuses, inside glabrous. Ovaries: glabrous, short columnar 0.14 cm tall and 0.11 cm wide at base pair.

188

Hoya faoensis Kloppenburg & Siar 2008 Described in Fraterna 12/4:18-21, Oct-Dec 2008.

View of the corolla outside with pedicel attached enlarged about 8X. The pedicel is short (broken off ?), terete, glabrous, shriveled longitudinally, 1 cm. long and 0.06 cm. in diameter Corolla outside is granulose and glabrous, deeply cut. Calyx is very small and sepal apexes do not come near the corolla sinuses.

Pedicel and calyx enlarged about 16X. The ovaries stayed with the corolla when this structure was removed. Calyx is ciliate, otherwise glabrous. sepals 0.24 cm long from center - apex; 0.15 cm from base - apex and 0.11 cm. at the widest just out from base. Apexes are narrowly rounded to subacute. Outside surface is finely granulose.

189

Hoya ferrerasii Kloppenburg & Siar Published in Fraterna 32 #4, 2010.

Calyx: sepals very small, does not reach the corolla sinuses, triangular, apex sub acute, 0.09 cm long, widest 0.07 cm outside puberulous, inside glabrous, very small ligules. Enlarged about 10X.

Ovaries: short domed, glabrous, 0.06 cm tall and 0.05 cm wide at pair base. Enlarged about 22X.

190

Hoya fetuana Kloppenburg 2008

Described in Fraterna 16(2): 1 2003

Pedicel, calyx and ovaries side view enlarged about 8X. Pedicle curved, terete, glabrous, 3.1 cm. long, translucent 0.06 cm. in diameter. Calyx glabrous, ligules present, an occasional cilia, 0.17 cm. long, 0.16 cm. at the widest. apex narrowly obtuse, overlap ca. 1/4. Ovaries short domed, 0.10 cm. tall and base pair 0.10 cm. wide, glabrous.

Top view of the calyx and ovaries enlarged about 16X. rather membranous, ovaries very stubby; occasional cilia.

191

Hoya finlaysonii Wight 1834 Described in Contributions to the Botany of India (1834) 38. R. Wight & Wallich. Calyx not mentioned in the type description.

Top view of the calyx. The sepals have a broad base and are overlapped about 1/3, ligules are present, edges are ciliate with apices rounded, there is much mahogany pigmentation. Ovaries are short and small, color waxy pale-yellow. Calyx: triangular, outside spotted with deep mahogany color, from a green calyx base, some hairs pointing toward apex, edges ciliate, overlap about 1/3, 0.18 cm. long and 0.20 wide at base. Long narrow ligules present that reach the sinus. Side view of a removed pedicel and calyx, with short ovaries, as above scattered hair cells and rough surface spotted a deep mahogany color on the outside. Pedicels: with scattered hairs and lenticels, green-pink 0.25 cm. long 0.13 cm. in diameter, terete straight.

Ovaries: domed, 0.10 cm. tall and base of pair 0.16 cm. glabrous and waxy.

192

View of the calyx attached to the corolla. Note: the sepals do not reach the sinus of the corolla, the outside surface of the sepals are covered with scattered hair cells and granulose colored protrusions, as with the pedicel. The corolla on this surface is glabrous.

193

Hoya fischeriana Warburg 1904

Described in Fragmentia Florae Philippinae 1 (1904) 129. (Perkins) R. Schlechter & O. Warburg. 3.

Pedicel, calyx with Ovaries. Pedicel: 2.0 cm. long, 0.05 cm. in diameter, glabrous, curved, filiform. Calyx: 0.18 cm. long 0.12 cm at the widest, triangular, membranous.

194

Hoya fitchii Kloppenburg 2009

Described in Fraterna 2009

Side view of the pedicel, calyx and ovaries enlarged about 8X Pedicel: strict, terete, glabrous. 1.5 cm. long, 0.08 cm. in diameter. Calyx: sepals nearly reaches the corolla sinuses, edges hyaline, centrally thickened, outside granulose, glabrous.

Top view of the calyx. Inside surface glabrous, shiny, sepals overlap at the base about 1/3, ligules are present, actually double one on top of the other, lower one larger than the one on top. Edges ciliate. Apex – base Apex – center Widest Base – sinus

0.18 cm. 0.22 cm. 0.17 cm. 0.06- 0.07 cm.

Ovaries: short dome shaped. 0.11 cm. tall and 0.11 cm. wide at base of pair, glabrous.

Outside view of the flower. Difficult to differentiate but sepals nearly reach the corolla sinuses but still short a little. Sepal outside surface granulose. Corolla surface glabrous with fine granulations.

195

Hoya forbesii King & Gamble 1903

Described in Flora of the Malay Peninsula (1903) 574. King & Gamble. 16.

Pedicel, calyx, and ovaries side view enlarged about 16X. Sepals are triangular 0.14 cm. long and at base 0.10 cm. wide with little overlap, ligules are present, surfaces glabrous, outside surface granulate inside shiny slick. edges ciliate. Ovaries columnar, glabrous 0.15 cm. tall and 0.10 wide at base of pair.

Outside surface of the flower and calyx attached enlarged about 16X. The sepals are small and do not even approach the corolla sinuses, the surface is granulose. Corolla here is glabrous.

196

Hoya fungii Merrill 1934 Described in Lingnan Science Journal 13 (1934) 68-9. E. D. Merrill. "sepals oblong, acute, about 2.5 mm. long, 0.15 cm. widest; outside lightly dispersed, somewhat depressed, near rusty pubescence."

Top view (skewed a little) of the calyx. Outside with long stellate hairs but edges do not appear ciliate.

Side view of the calyx. The sepals are hairy on the outer surface, long and triangular shaped glabrous within. Pedicels pubescent.

197

View from back side of flower (outside) with calyx attached to the corolla. Note the long hairs near and on the calyx outside surface, also the white pubescent inner surface of the corolla where the sinus turns under slightly. This surface of the corolla is glabrous.

198

Hoya fuscomarginata N. E. Brown 1910 Described in Kew Bulletin:10910. 577, 278. "Sepals minute, 1 mm. long, ovate, acute."

Top view of the calyx with parts of the corolla still attached. Inner surface here is pubescent which would be unusual. ?.

Side view of the pedicel, calyx and ovaries. Outer surface rough and a few hair cells. Sepal apex acute. Description says pedicels and peduncles sparsely and minutely puberulous.

199

Hoya gigantangensis Kloppenburg 1992 Described in Fraterna 4th Quarter (1992) 2. "Calyx segments ovate, glabrous, margins ciliate, much smaller then the corolla." Calyx top view. Sepals ciliate ovately triangular 0.21 cm. long and 0.17 cm. widest. Ligules are present at the bases. Overlapped about ¼. Pedicel: terete, glabrous, various lengths, curved, 1.5 cm. – 2.8 cm. long, 0.05 cm. in diameter. Ovaries: dome shaped 0.12 cm. tall base pair 0.13 cm. across. Photo from PNH 36787.

200

golamcoiana Kloppenburg 1991 Described in Fraterna 3 (1991) II. R. D. Kloppenburg. "Calycis foliola, lanceolato-ovata, subacute, margine ciliate, quam corolla paulo breviore."

Top view of the calyx and ovaries. The edges have a very few cilia, there are ligules present at the base, there is a small overlap of the Calyx: outside granular, scattered hair cells; inside glabrous, apex of sepals narrowing, rounded, with slight basal overlap, ligules present at each sinus, 0.25 cm. long, base 0.18 cm. wide. View of pedicel, and pedicel, calyx and ovaries. The pedicel is strict, terete and with sparse stellate like hair cells which increase in density on the calyx base. The sepals are glabrous inside, cupped slightly, and lay rather flat; they do not reach the corolla sinus. Ovaries are bottle shaped. Pedicle: terete, strict, with fairly long stellate hairs slightly more dense and longer toward the calyx. Ovaries: shaped 0.19 cm. tall; 0.13 cm wide at base of pair.

201

bottle

Hoya graveolens Kerr 1939 Described in Kew Bulletin (1939) 461 A. F. G. Kerr. "Calyx 5 lobed, outside sparsely hirsute, inside provided with minute glands; lobes narrowly ovate, ciliate, nearly 2 mm. long, base 1 mm. wide." Top view of the calyx. Edges ciliate, ligules present near at least two sinuses. Apices narrowly rounded . Calyx: sepal apex acute, does not reach the sinus of the corolla, outside as with the pedicel, inside glabrous. 0.14 cm. long, 0.12 cm at the widest. Edges ciliate, with 1/3 overlap.

Pedicel, calyx and ovaries. The pedicel is terete, straight with lenticels and a occasional hair cell hooked pointing apically and with short fine pubescence, 0.17 cm. long and 0.10 cm. in diameter.

Ovaries: domed, 0.11 cm. tall, base pair 0.12 cm. wide, glabrous.

202

Hoya greenii Kloppenburg 1995 Described in Fraterna1995/2, 12-13.

Top view of the calyx and ovaries. Note how distinctly the ligules at the sepal bases stand out. Sepal ends are obtuse, edges ciliate; inside glabrous waxy. Calyx: ciliate (as opposed to H. darwinii Loher) cupped inward with slightly overlapping sepals, 0.12 cm. long, widest 0.11 cm., apex rounded, with distinct ligules present.

Pedicel: straight, glabrous, terete, 2.7 cm. long x 0.10 cm. in diameter, glabrous with fine granular lenticels, orange colored. Ovaries: bottle shaped with red ends. 0.22 cm. tall base pair 0.12 cm. wide.

203

Hoya halconensis Kloppenburg 1990

Described in Fraterna 3rd Quarter 1990.

Outside surface of the calyx enlarged about 16X. Sepals are thickened triangular-linear, ciliate with a few cilia on the immediate surface, apex narrowly rounded, little base overlap.

Inside view of the calyx, surface is shiny glabrous, a little overlap at the base. Sepals: 0.30 cm. long, 0.19 cm. wide near the base, very small ligules present. Ovaries: columnar 0.24 cm. tall and base of pair 0.14 cm. wide.

Outside surface of the flower at the calyx area and corolla sinus enlarged about 8X. The sepals do not extend to the corolla sinus. Corolla is fleshy-thick. and the coronal outer lobe projects beyond the sinus. Sinus – sinus Sinus – center Sinus – apex Apex – center Widest

204

0.50 cm. 0.40 cm. 0.95 cm. 1.20 cm. 0.60 cm.

A second clone, below.

Pedicel and calyx enlarged about 8X. Scales are linear, ciliate, outside q few hair cells, base barely overlap 0.35 cm. long and 0.20 cm wide near base, ligules are present. Ovaries: narrow domes 0.16 cm. tall and base pair 0.16 cm. wide.

Top (inside) view of the calyx enlarged about 16X. Note lobe apex is rounded, edges ciliate, base a little overlapped.

205

Hoya heuschkeliana Kloppenburg 1989 Described Hoyan 11/1 part2:i, 1989. "Calyx parvus fere ad basin in lobos 5 anguste triangulares partitus, lobis circ. 0.15 cm, punctata, auriclis 1 ab inter se basi lobis."

Calyx: 0.13 cm. long, 0.10 cm at widest. Glabrous with small ligules present.

Pedicels also rigid 0.4 cm. long, expanding as they approach the calyx.

206

Hoya heuschkeliana subsp. cajanoae Kloppenburg & Siar 2007 Described in Fraterna 20/4 2007.

Top view of the calyx. The ring at base of the ovaries with its vascular bundles is visible and also the small ligules. Below this is the short pedicel which is round, green (actually as is the calyx) and it has a few scattered hairs. Calyx: 0.12 cm. long, ciliate, outside surface granulose base 0.10 cm. wide, small ligules present, little overlap of sepals.

Side view of the calyx and a portion of the pedicel. The color is actually an apple green. The outer surface has some scattered hairs (as does the pedicel) inside is waxy, smooth, the apex is narrowing rounded. The ovaries are tapering cones with a flat skewed apex. Pedicels: short, 0.35-0.38 cm. long, round, glabrous but scaly, 0.10 cm. in diameter. Ovaries: long and narrow dome narrowing columnar, 0.16 cm. long base of pair 0.09 cm wide.

207

Hoya ilagiorum Kloppenburg, Siar & Cajano Described in Asia Life Sci. 20(2) : 264. 2011.

Calyx on a flower enlarged ca. 10X. The sepals outer surface is granulate, triangular and does not reach the corolla sinuses.

Inside view of a calyx and ovaries enlarged ca. 16X. Sepals have fine cilia, ligules at the sinuses. Outside granulate, inside slick glabrous, slightly overlapped at the base. Shape triangular apex sub-acute. Apex – base Widest at base

0.15 cm long 0.13 cm wide

Ovaries: columnar, glabrous, 0.12 cm tall and base pair 0.10 cm wide.

208

Hoya incrassata Warburg 1904 Described in Perkins Frag. Fl. Philippineae (1904) 130. "Sepal 0.1 cm. long , broadly triangular rounded, obtuse tip, outside granulose; inside concave minutely ciliate, two adjacent sepals have 1 ligule each others have 2 each, edge of sepal is membranaceous and with a few ciliate cells."

209

Hoya smithii Kloppenburg 2010 (Hoya intermedia A. C. Smith) Described in Fraterna (2010). Described in Sargentia (1942) 111-112. Fijian Plant Studies II. A. C. Smith. "calyx lobes membranous deltoid-lanceolate, 3 to 3.5 mm. long, 1.8 to 2 mm. wide, on the back conspicuous fine lines".

Hoya intermedia A.C.Smith Fiji Vanalevu

Type 399

1932 Smith UC A

Photomicrographs from they type sheet flower at (UC). Photos 1/6/1990: This species belongs in the Subsection Angusticarinata Kloppenburg. Calyx: Sepals 0.30 cm. long. very pubescent outside, glabrous inside 0.60 cm in diameter. No ligules noticed. Calyx on the back of the flower enlarged about 8X. Sepal raised by a thin sewing needle long and narrow. Surface is actually hirsute. Apex just reaches the corolla sinus.

Calyx outside surface enlarged about 8X. One sepal torn free. Very hirsute on this surface, lobes ;long triangular apices acute.

Inside view of the calyx enlarged about 8X. This surface is slightly cupped in the center under the ovaries (here deteriorated) This surface is glabrous but the hirsute cells show through the thin sepals.

210

Dr. Schlechter's Hoya intermedia (invalid & unpublished) Not the same species as above, not even from the same area. An Acanthostemma species

UC 75000

211

Hoya kanlaonensis Kloppenburg, Siar & Ferreras

Published in Fraterna 23 #4, 2010

Calyx: with ovaries on the pedicel, inside view enlarged ca. 12X. The sepals are membranous, broadly ovate, apex obtuse, glabrous, sub-ciliate to ragged edged. ½ overlap, ligules present, 0.18 cm long x 0.15 cm at the widest. Ovaries: columnar, glabrous 0.10 cm. tall and base pair 0.08 cm. wide.

212

Hoya kanyakumariana Henry & Swaminathan 1978 Described in Journal, Bombay Natural History Society 75 (1978) 462-463. A. H. Henry & M. S. Swaminathan.

Pedicel, calyx and ovaries enlarged about 8X. Pedicel is nearly straight, 1.5 cm. long, terete, glabrous, 0.08 cm. in diameter. Calyx outside is covered with a microscopic fuzz. Sepals reach less than 1/2 way to the corolla sinuses. Inside glabrous, slick. Ovaries short, glabrous domed.

Top view of a calyx enlarged about 8X. The calyx is small. Sepals are 0.13 cm. long and 0.18 cm from apex to the center. The widest is 0.10 cm. and small ligules are present.

213

Hoya kastbergii Kloppenburg 2003

Described in Fraterna 16/4 1003:1-4.

Side view of the pedicel, calyx and ovaries. Calyx: outside glabrous from a bulbous base. Center apex 0.15 cm. Base - apex 0.10 cm. Very slight overlap at the base.

Ovaries: very short, glabrous, 0.15 cm. tall, 0.14 cm. at base pair.

Top view of the calyx and ovaries. There is a piece of the corolla left attached at the lower Right. Calyx is very membranous, apex rounded to narrowly rounded. I did not note any ligules present.

214

Hoya kenejiana Schlechter 1913 Described in Botanische Jahrbücher 50 (1913) 114. “Die Asclepiadaceen von Deutsch Neu-Guinea” R. Schlechter. 12.

Pedicel and calyx side view enlarged about 16X. Pedicel is terete, strict, rough but glabrous. Calyx base is granulose with a few hair cells, sepals with incised edges 0.18 cm. long and 0.12 cm. at the widest.

Tope view of the calyx, inside shiny and glabrous, dark ligules are present. Sepal bases overlap only slightly, apices are obtuse.

Outside surface of the corolla with pedicel attached enlarged about 8X. Sepals do not reach the corolla sinuses. Corolla outside glabrous, finely granulose, inside densely pubescent except for small apical tip which turns under as do the corolla lobe edges.

215

Hoya krohniana Kloppenburg & Siar 2010 Described in Fraterna 2010.

Outside view of the pedicel and calyx enlarged about 8X, Sepals are thin but centrally thickened, triangular. Apex narrowly rounded 0.16 cm. long, 0.05 cm. at the widest, pedicel base rounded and 0.18 cm. across, all parts glabrous, surface of sepals finely lenticular-granulose, base granulose.

Inside view of the calyx and ovaries enlarged as above. There are ligules present, ovaries are bottle shaped, 0.15 cm. tall, glabrous, apical area narrower, base pair 0.09 cm. wide.

Outside of flower enlarged about 8X, pedicel, calyx and corolla, sepals reach a little over ½ way to the corolla sinuses, corolla outside glabrous and finely granulate, pedicel glabrous and smooth except at the enlarged base that is roughly granulate.

216

Hoya lambii Green 2000 Described in Fraterna 13/2:2000, 2-3. "Calyx 1mm x 3mm, cuneate."

Top view of the calyx. Small ligules are present in pattern 2-1-1-1-1. Edges are very ciliate centrally thickened, edges membranous, centrally cupped and glabrous. Calyx: rather narrow, linear far from reaching the corolla sinuses; from a round swollen base, punctate with a few scattered hair cells, ciliate, inside cupped and glabrous. 0.29 cm. long to the center, otherwise 0.22 cm.; 0.15 cm widest, apex narrowly rounded.

Pedicel: terete, hirsute, hairs pointing apically with many punctations 2.65 cm. long, 0.11 cm. in diameter.

Side view of the calyx. See dimensions above. Ovaries are domed 0.12 cm. tall and the base pair 0.12 cm. broad, glabrous.

View of pedicel and calyx attached to the. The sepals do not come near the corolla sinuses. Corolla on this outside surface is glabrous and shiny.

217

Hoya landgrantensis Kloppenburg & Siar 2009

Described in Asia Life Sciences 18(1)2009, 147-150.

Calyx: sepals nearly to the corolla sinuses, membranaceous, glabrous on both surfaces, ovate elliptic apex blunt, widest at the overlap that is ca. ¼.; 0.22 cm. long and 0.13 at the widest. Ovaries: short, columnar, and glabrous. 0.10 cm. tall and as wide at pair base. Enlarged about 8X.

Side view of outside of flower enlarged ca. 8X. the sepals nearly reach the corolla sinuses. The coronal lobe outer apices exceed the corolla sinuses. corolla outer surface is glabrous and slick. All parts are yellow in color.

218

Hoya latifolia G. Don 1838 Described in General System of Gardening and Botany 4/23 (1837) 127. Leaflets of corona elongated, triangular. 21.

G. Don.

Side view of the Pedicel and calyx with ovaries on left enlarged about 16X. The picture on the right is a portion of the pedicel enlarged about 48X. The pedicel is granulose with a fine pubescence.

Top view of the calyx enlarged about 16X. Note that the lobes are triangular with slight overlapping at the base.

219

View of the outside of a flower with the calyx attached to show that the triangular shaped sepals do not reach the sinus of the corolla. It appears that the coronal outer apex just reaches the sinus on the other surface. Outer surface is glabrous.

220

Hoya leytensis Elmer ex Burton 1988 Described in The Hoyan Vol. 10 Part 2 July 1988 p.i. "Calyx very small, segments hyaline, spreading, oblong with obtuse tips, 0.15 cm. long or shorter."

Calyx very small, the 5 hyaline segments spreading, oblong with obtuse tips, 1.5 mm. long or even shorter.

221

Hoya litoralis Schlechter 1905 Described in Nachtrage zur Flora der Deutschen Schutzgebeite in der Südsee (1905) 363. K. Schumann & K Lauterbach. "with the segments of the calyx ovate-oblong, obtuse, glabrous, longitudinally barely exceeding by 0.1 cm." Photomicrographs and data from flowers sent by Ted Green, IML 708 collected by G. Hardy 25 November 1986 on Moa Island, Torres Str., Queensland, Australia:

Side view of pedicel, calyx and ovaries. I believe the important thing here is the shape of the ovaries, they are just as Dr. Schlechter drew them, narrowing near the apex. Pedicel here curved. Calyx: sepals 0.14 cm. long by 0.14 cm. at the widest, apex somewhat round, ciliate, with very small ligules, inside glabrous.

222

Hoya loheri Kloppenburg 1991 Described in Fraterna 1991/3, Sect.2,III.

From herbarium material: Calyx: lobes convex, membranaceous, ligules present at base, ovate, 0.11 cm. long, 0.07 cm. wide below the middle, apex subrounded, edges ragged. From live material: Calyx lobes: 0.10 cm. tall 0.10 at widest with relatively long yellow ligules. Edge membranaceous, somewhat irregular in outline, 1/3 overlap.

Pedicel: 1.2 cm. long, threadlike, round, flexible, glabrous. Pedicel: 0.06 cm. in diameter at calyx attachment.

Ovaries: bottle shaped 0.18 cm. tall, the pair 0.07 cm. in diameter.

223

Hoya lucardenasiana Kloppenburg & Siar 2009

Described in Asia Life Sciences 18(1): 144-147.

Calyx on pedicel enlarged as above, Pedicel enlarges only slightly at calyx base, sepals short ovate, 0.15 cm. long and 0.10 cm. at the widest, reach less than half way to the corolla sinuses, apex obtuse. Ovaries are tapered dome shaped 0.15 c, tall and base pair 0.13 cm. wide, glabrous.

Top view of the calyx enlarged about 8X. Ligules are present, sepals with very slight overlap. Both surfaces are glabrous.

224

Hoya lucyae Kloppenburg & Siar 2006

Described in Fraterna 19/4: 2006, 8-10.

Side view of the pedicel, calyx and ovaries enlarged about 16X. Pedicel: curved, terete, filiform, glabrous. Enlarged toward the calyx base. 0.8 – 2.5 cm. long. Calyx: triangular. 0.10 cm. long and 0.08 cm. wide at the base. Glabrous. Ovaries: columnar 0.13 cm. long and base pair 0.07 cm. wide, glabrous.

225

Hoya marginata Schlechter 1905 Described in Nachtrage zur Flora der Deutschen Schutzgebiete in der Südsee (1905) 364. K. Schumann & K Lauterbach. Calyx not mentioned in the type description.

From plant I collected at Tenerau Falls on Guadalcanal, Solomon Islands.

226

Hoya matavanuensis Kloppenburg 2009 Described in Asklepios 111:22. 2011

Pedicel, calyx and ovaries enlarged about 16X.The pedicel is filiform, 2.4 cm. long with a few scattered very fine hair cells, terete, shriveled on drying longitudinally, 0.05 cm. in diameter. Sepals are small, ciliate, not much centrally thickened, membranous, overlapped at the base, tapering toward the rounded outer apex, no ligules. Inside moist or waxy, glabrous. 0.18 cm. x 0.10 cm. Ovaries narrowly dome shaped 0.12 cm. tall and base pair 0.10 cm. wide, glabrous, waxy.

Top view of the calyx enlarged about 16X, sepals ciliate, apexes mostly rounded, surfaces glabrous. Only a slight basal overlap.

Side view of the flower, enlarged about 8X. the calyx at the corolla base is small and the apexes do not nearly reach the corolla sinuses. Corolla outer surface is glabrous and finely granulose.

227

Hoya megalaster Warburg 1901 Described in Flora de Deutschen Schutzgebeite in der Südsee (1901) 513. Calyx not mentioned in the type description.

228

Hoya marananiae Kloppenburg & Siar

Calyx: on pedicel enlarged about 20X. Sepals do not quite reach the corolla sinuses, are triangular ciliate, membranous apex sub-acute 0.12 cm. long x 0.06 cm at the widest; with a small basal overlap, ligules not observed. Ovaries: domed, glabrous, 0.07 cm. tall and base pair 0.07 cm. wide.

229

Hoya meredithii Green 1989

Described in Phytologia 64/4 (1989) 304. Ted Green. "calyx lobes 5, 1 ½ mm, acutely triangular, glabrous." Photomicrographs from flowers sent to me by Ted Green as 80-05 his ascension number for this species. Top view of the calyx. The sepals here are irregular in size and shape, edges are ciliate. Calyx diameter 0.42 cm. Sepals 0.18 cm. long, widest 0.14 cm. Only 2 ligules on 5 positions.

Side view of the pedicel, calyx and ovaries. Picture doubled due to shaking ?) Ovaries columnar domed 0.13 cm. tall.

230

Hoya merrillii Schlechter 1904 Described in Fragmentia Florae Philippinae 1 (1904) 131. R. Schlechter & O. Warburg. "calyx segments ovate-lanceolate, obtuse glabrous 0.1 cm long barely exceeding." Side and Top views of the calyx and ovaries. Sepals are triangular with jagged edges with small ligule like glands at the base (inside). Ovaries domed sub columnar. Calyx: triangular sepals, base pinnate, apex narrowly rounded 0.135 x 0.105 cm., inner surface moist, glabrous, outside granulose, edges jagged. Small v-shaped gland at base.

Side view of pedicels and calyx. They are glabrous but with outer granulations, Calyx base is enlarged, sepals relatively small, apex narrowly rounded and acute. Pedicel: glabrous 0.08 diameter.

231

straight, cm in

Hoya micrantha Hooker 1883 Described in The Flora of British India 4 (1883) 55. J. D. Hooker. "Sepals ovate, obtuse." Photomicrographs: flowers from plant at Fresno, California 19 April 1990. Roll 69. Calyx top view. There is some overlap of the sepals maybe 1/4, shape is long triangular, edges are ragged but not ciliate. I could not find any ligules. Calyx: 0.15 cm. long, 0.10 cm. at base, no ligules observed, margines rough, shape of sepals triangular. Outer surface rough; inner smooth and glossy. Apex does not reach the sinus.

Pedicel, calyx and ovaries. The ovaries ate very long, rather narrow and bottle shaped. The pedicel is round and rather thick with some scattered hairs pointing apically. The sepals are small and rough surfaced outside. Pedicel: terete, glabrous, with fine raised lenticels, curved variable in length mostly 2 cm. long and 0.12 cm. in diameter. This forms a flat to concave flower cluster.

232

Hoya sp. Minibella Cult. Supposedly a cultivar or hybrid.

Calyx: sepals with a few cilia, ligules at the sinuses 0.28 cm. long and 0.15 cm. widest, outside glabrous.

233

Hoya mitrata Kerr 1940 Described in Icones Plantarum 35 (1940) 3406. A. F. G. Kerr. Hoya mitrata Kerr. Asclepiadaceae. Tribus Marsdenieae. "Sepals broadly ovate, obtuse, about 2.5 mm long, shortly ciliate, outside minutely puberulous." Photo of this species sent to me by Chanin Thorut of plant collected by him in Thailand.

Pedicels: glabrous, punctate, longitudinally striate (on drying) 2.5 cm. long, curved, 0.06 cm. in diameter. Calyx: 0.28 cm. long, widest 0.18 cm. slightly ciliate, outer apex rounded, with hyaline ligules present; very little overlap at base. Ovaries: columnar, tapering narrowly upward, 0.20 cm. tall and with a base of 0.15 cm.

234

Hoya monetteae Green 2004 Described in Fraterna 17/2:2004, 10-15.

Calyx on the back of the corolla enlarged about 8X. The sepals reach about half way to the corolla sinuses. The sepals are diamond shaped with an acute apex. The corolla outer surface is glabrous.

Calyx with ovaries enlarged about 8X. The ovaries are glabrous, columnar shaped 0.20 cm. tall and 0.20 wide at the base pair.

Top view of a calyx enlarged about 16X. The sepals are overlapped at least 1/3, slick glabrous inside edges ciliate. Sepals are 0.20 cm. long, 0.25 from the apex to the center. 0.25 cm. at the widest. Note the multiple ligules at the sinus, here the pattern is 1,3,2,3,1. some ligules are as long as 0.05 cm.

235

Hoya motoskei Teijsmann & Binnendijk 1826 Described in Annales Musci Botanici Lugduno Batavi 2(1826) 33, P. A. Miquel. Calyx not mentioned in the type description.

236

Hoya mucronulata Warburg 1901 Described in Nachtrage zur Flora Deutschen Schutzgebeite in der Südsee (1901) 513. K. Schumann & K. Lauterbach. "sepals 1 mm. long, ovate-lanceolate, somewhat recurved, glabrous." in 1907 pub.

Photomicrographs: from Herbarium sheet #18329 (UC).150 m altitude, 20 flow/cluster Leaves 13.5 cm. long – 7.5 cm wide, 10 nerves/side, reticulately netted.

Pedicel: glabrous. 0.04 cm. in diameter. Top view of the calyx. Diameter 0.42 cm. and 0.10 cm. wide at base, a few scattered hairs some cilia especially near base.

237

Hoya myrmecopa subsp. kapatalanensis Kloppenburg, Siar, Cajano, Guevarra & Carrandang

Pedicel, calyx and ovaries enlarged about 8X. Pedicel: is terete, glabrous, 1.10 cm. long, 0.08 cm. in diameter sepals with little overlap. Sepals: narrow elliptic, glabrous, little basal overlap. 0.14 cm. long and 0.06 cm. at the widest at base, glabrous, apex sub-acute dark ligules present. Ovaries: columnar 0.16 cm. long and base pair 0.07 cm. wide, glabrous.

238

Hoya nakarensis Kloppenburg, Mendoza & Ferreras 2013 ISSN 1055-4564 Described in New Hoya Volume 1-1, rare-hoyas.com.

Calyx: enlarged ca. 20x, the sepals are triangular, granulate outside, slick glabrous inside, 0.11 cm long and 0.07 at the widest, I did not see any ligules. Edges slightly ragged and an occasional cilia.

Ovaries: enlarged ca. 42x, the ovaries are short dome shaped, glabrous, 0.06 cm tall and base pair 0.06 cm wide; apex slightly modified.

239

Hoya naumanii Schlechter 1908 Described in Beiblatt zu den Botanischen Jahrbücher #92 40 (1908) 15. R. Schlechter. "calyx segments oblong obtuse, minutely puberulous, margines minutely ciliate about 2.5 mm. long."

Top view of the calyx. Edges with small cilia and a few hairs on outside, laying flat, inside glabrous, with small basal ligules. Sepals 0.30 cm. long and 0.18 cm. at the widest, diameter of calyx 0.72 cm. Ovaries short columnar, glabrous all over. Pedicel thick with a few cilia, 0.15 cm. in diameter, round.

240

Hoya nuuuliensis Kloppenburg & Siar 2008 Described in Fraterna 21 #4 Oct. Dec 2008, 14-17.

Calyx on outside of the flower corolla enlarged about 8X. The Pedicel is terete, glabrous, 1.5 cm. long, filiform. The sepals are large and nearly reach the corolla sinuses. Corolla outer surface is granulose, glabrous, lobes are deeply cut with acute apexes and wide lobes just above the sinuses.

Pedicel and inside surface of the pedicel enlarged about 8X. Sepals relatively large, membranous, ligules present of same color as the sepals, difficult to detect. Apexes acute. Center to apex Apex - apex Widest

0.20 cm. 0.18 cm. 0.10 cm. wide at the base.

241

Hoya obovata Decaisne 1844 Described in Prodromus Syst. Veg. 8 (1844) 635. DeCandolle. Calyx not mentioned in the type description.

Photo of pedicel Calyx and ovaries. The sepals reach the corolla sinus, are ciliate and hairy on the outer surface as is the pedicel (where the hairs point distally. Inner surface of calyx is granulose and extremely fine pubescent. It appears glabrous except under high magnification. Pedicels: 0.25 cm. long, terete, pubescent, dull green, 0.3 cm. in diameter. Calyx: 0.30 cm. long, widest 0.21 cm., less than ¼ overlap; outside hirsute, inside glabrous, apex narrowing rounded to somewhat acute. Ligules present and prominent one at each base.

Ovaries: paired upright, bottle shaped, base and height 0.15 cm.

242

Hoya obtusifolia Wight 1834 Described in Contributions to the Botany of India (1834) 38. R. Wight. Calyx not mentioned in the type description.

Calyx and ovaries. Apex of sepals narrowing, rounded, ciliate. Inside glabrous. Pedicel: 3 cm. long 0.25 cm. in diameter with very stubby granules with some hair cells, short and hooked.

Calyx and ovaries with portion of pedicel. Outside of sepals and pedicel granulose with pubescence. Calyx: cupped inside glabrous edges ciliate, outside granulose with scattered hair calls. 0.41 cm. long base 0.37 cm. wide, no ligules seen.

Ovaries: glabrous, dome shaped, 0.24 cm. tall and 0.20 wide for base pair.

243

Hoya odorata Schlechter 1906

Described in Philippine Journal of Science 1 (1906) 303. R. Schlechter.

Pedicel: 2.5 cm. long, 0,12 cm in diameter, terete, glabrous, lenticulate. Sepals: 0.20 cm. long, pale yellow, slight overlap, blunt obtuse, 1 ligule at each sinus. Base of calyx 0.30 cm. wide.

2nd clone

Pedicle calyx and ovaries side view. Enlarged about 8X Pedicel: 1.9 cm. long 0.06 cm. in diameter. Surface with sparse pubescence enlarging toward the calyx. Calyx: sepals triangular, 0.16 cm. long and widest 0.16 cm. edge wit a few cilia, edges hyaline, Inside glabrous and slick., no overlap at base, no ligules observed. Ovaries: very stubby 0.11 cm. tall and abase pair 0.16 cm. wide.

244

Hoya odettiae Kloppenburg 1988 Described in Fraterna 11/2:1998, 8-11.

"Calyx: triangular, membranous with very slight basal overlap, glabrous and shinny inside, outside glabrous punctate, small triangular ligules present inside at base, apex rounded or tapered, edges incised."

Top view of the calyx. Note: the sepal shape and that the edges are irregular. There is about a ¼ overlap at the base. The calyx is flat.

Pedicel: persistent, slightly curved, round, glabrous, 4-6 cm long, 0.06 cm. diameter.

245

Hoya onychoides P. I. Forster & Liddle 1995

Described in Austrobalia 4 (3) (1995) 402-404. “Taxonomic studies of the Genus Hoya R. Br. (Asclepiadaceae: Marsdenieae) in Papuasia” P. I. Forster, D. J. Liddle & Iris M. Liddle.

View of the calyx outer surface enlarged about 8X. Sepals are elliptic to triangular and surface is sub-convex finely granular, with base about1/3 overlapped.

Inside view of a calyx lobe enlarged as above, surface is glabrous and slick. Sepals are 0.52 cm. long to the ovary base and 0.35 – 0.40 cm at the widest.

246

Hoya opposita G. Don 1837

Described in General System of Gardening and Botany 4 (1837) 128. G. Don. 37.

Side view of the pedicel and calyx enlarged about 8X. Pedicel: terete, filiform, 1.65 cm. long x 0.04 cm. in diameter, glabrous, yellow, curved. Ovaries: domed 0.12 cm. tall. Calyx: broadly ovate with a few cilia, membranous; at base overlapped about 1/3 (0.05 cm.) length 0.15 cm., base 0.12 cm., dark ligules present.

Flower folded enlarged about 8X. Coronal lobes extend from the sinuses of the corolla. Calyx does not hug the corolla. This surface glabrous.

247

Hoya pachyclada Kerr 1939 Described in Kew Bulletin of Miscellaneous Information (1939) 462. A. F. G. Kerr. "Calyx deeply 5-lobed, outside shortly puberulous, lobes ovate, obtuse, shortly ciliate about 2 mm. long." Top view of the calyx. The sepals are with a considerable overlap (over 1/3); the edges are finely ciliate and membranous. There are multiple ligules at the sepal base, (not at the sinus area); the apices are rounded. Calyx: sepals almost triangular, edges finely ciliate, and membranous, multiple ligules at the sinuses, overlap over 1/3. Length of sepal 0.25 cm., widest 0.21 cm.; overlap 0.10 cm.

Side view of a portion of the pedicel with the calyx and ovaries. The pedicel is straight or slightly curved, punctategranulose rough with a few short hairs. The calyx is outspread, sepals short thickened in the center also granulaterough on the outside. Ovaries are dome shaped, very short. Pedicel: straight or slightly curved 2.8 Cm. long, punctategranulate, rough with a few short hairs; 0.27 cm. in diameter. Ovaries: 0.25 cm. tall, base of the two 0.11 cm. wide, dome shaped and short.

248

Hoya palawanensis Kloppenburg, Siar, Mendoza, Guevarra & Carandang 2012

Calyx: enlarged about 12x. The sepals are long triangular membranous, overlap about ½ at the base, apex sub acute, 0.25 cm long and 0.15 cm at the widest. Granular outside, slick glabrous inside. Small dark ligules present. Ovaries: short dome shaped, glabrous, 0.10 cm tall and 0.10 cm at pair base.

Corolla: ventral surface of a flower enlarged about 16x. The sepals barely reach the corolla sinuses, the apex of the coronal lobes exceed the corolla sinuses. Corolla is normally rotate, here folded in over the corona and this surface is granulate.

249

Hoya pallilimba Kleijn & van Donkelaar 2001 Described in Blumea August: 2001. Calyx not mentioned in the type description.

Calyx top view. Sepal bases overlap about ¼, veined on inside view, Ligules are present, 0.14 cm. in length and 0.10 cm. widest.

Side view of the pedicel, calyx and ovaries. Sepals lay essentially flat outspread. Outside there are a few scattered hair cells and the edges are sparsely ciliate. Inside is glistening, glabrous slightly thickened in the central portions. Ovaries are bottle shaped, glabrous, upright 0.10 cm. tall and base pare are 0.10 cm. wide.

250

Hoya panayensis Kloppenburg & Siar 2010

Inside view of the calyx enlarged about 8X. Sepals are thin, 0.12 cm. long, 0.09 cm. wide near the base, a little overlap, glabrous on all surfaces, apex obtuse, no ligules noted. Ovaries: 0.12 cm. tall and wide at the base of pair.

Bottom view of the flower enlarged about 8X. Surface is glabrous-granulate, sepals here darker and do not reach the corolla sinuses. Sinus – sinus Sinus – center Sinus – apex Apex – center Widest

251

0.16 cm. 0.27 cm. 0.35 cm. 0.69 cm. 0.30 cm.

Hoya panchoi Kloppenburg 1988 Described in Hoyan Vol. 10 #1, part II July 1, 1988 p. 1. Syn. H. bulusanensis Elmer in Leafl. Philipp. Bot. 10:3577 1938. Type: A.D.E. Elmer. "Calyx spreading, lobes glabrous and obtuse to acute, coriaceous 0.15 cm. in length, united at the base."

Pedicels bearing rather rigid and lividus colored flowers. Glabrous, terete, enlarged only a little as it approaches the calyx.

252

Hoya parvifolia Schlechter 1908

Described in Botanische Jahrbücher 40:15(1908) Beibl. Schlechter. "segments of the calyx ovate obtuse, margins minutely ciliate, at the center glabrous 0.2 cm. long." Calyx top view much enlarged, base of sepals overlap about 1/4. Outside finely pubescent, inside glabrous edges with a few cilia, ligules are present. Sepals 0.20 cm. long , 0.15 cm. widest.

Pedicel, terete, granulose and with some fine pubescence.

Sepals do not reach the corolla sinuses.

253

Hoya pauciflora Wight 1848 Described in Icones Plantarum Indiae Orientalis 4 (1850) 1269. Robert Wight. Calyx not mentioned in the type description. Photomicrographs from flowers sent to me by Jerry Williams, Rainbow Gardens, Vista California: 27 Sept. 1993.

Top view of the calyx. Sepals are irregular in shape. Ligules are present at two adjacent sinuses and absent at the three others. Calyx: rather flat, granulose outside, a few cilia, sepals (rose colored) rather thick in the central region, edges membranous with colorless edges, widest 0.15 cm., 0.20 cm. long, 1/8-1/4 overlap; apex narrowly rounded, with base ligules. .

Side view of the pedicel, calyx and ovaries. Pedicels: strict, glabrous, terete, with raised lenticels with reddish color tones; 2.4 cm. long. Ovaries are bottle shaped, glabrous

254

Hoya paulshirleyi Kloppenburg & Green 2011 Described in Fraterna 2011

A closed flower with calyx at base enlarged about 8X. The calyx outer surface is granulose, the sepals do not reach the corolla sinuses, Outer corolla is glabrous, lobes widest above the sinuses.

Side view of the calyx enlarged about 8X. Sepals are ciliate, broad based triangular but apex is rounded, overlap at the base 1/3. 0.14-0.25 cm. long and 0.15 – 0.19 cm. at the widest.

Calyx inside view enlarged about 8X. Inner surface is slick, glabrous, thin basal ligules are present. Texture of sepals sub-membranous.

255

Hoya paziae Kloppenburg 1990 Described in Fraterna 3/4:2,VI, 1990. "calyx segments linear, narrowly oblong, ciliate, ligules present."

Calyx: edges membranaceous, 0.10 long, here deeply cut with little or no overlap, 0.07 cm. at base, edges ciliate. Top view of the calyx and the cut off pedicel beside the calyx. Note: the shape of the sepals (triangular), the amount of overlap (about ¼), the vascular ring around the base of the ovaries. The sepals are somewhat ciliate and there are ligules at the sinus, two at one point on the lower central side. The pedicel is terete and enlarges as it approaches the calyx. Pedicle: may be broken off so shorter than in vivo, length here 0.80 cm. long, terete, glabrous, dark, linearly grooved on drying. 0.06 cm. in diameter. Ovaries: 0.09 cm. tall; base of 0.11 cm. Side view of the calyx enlarged about 16X. This is from the clone IML 518, flowers from Ted Green, Kaaawa, Hawaii, USA. The pedicel enlarges slightly as it approaches the calyx, it has occasional hair calls and lenticels, is terete. The sepals are short, somewhat triangular, granulose on the outside, edges with a few cilia. The ovaries are short and bottle shaped.

256

View of the corolla with its attached calyx (red) enlarged about 16X. The sepal does not reach the corolla sinus. The apex of the sepal always points to the corolla sinus.

257

Hoya pentaphlebia Merrill 1918 In Philippine Journal of Science 13 (1918) 330-331. E. D. Merrill. "Sepals membranaceous, oblong-ovate, somewhat acuminate, 1.5 mm long."

Top view of calyx. apical ends are turned inward, ligules are present. Base overlap approx.1/4. Ovaries dome-shaped Calyx: Triangular with a few cilia, apex rounded and overlapped at the base approximately ¼. Ligules present, 0.12 cm. tall, base 0.10 cm. Does not extend to corolla sinus, short by 0.20 cm.

Pedicels: 2.5 cm. long, 0.07 cm. in diameter, glabrous, terete, of equal length, curved, 10-15 in number. Ovaries: domed shaped 0.1 cm. tall base 0.07 cm. Staminal crown apiculate.

258

View of calyx on back of corolla. The apex of the calyx always points to the sinus of the corolla. Here the pedicel is enlarged as it approaches the calyx. The calyx here appears more narrow than in the previous view, possibly due to the translucent edges which are barely visible. 16X.

259

Hoya picta Miquel 1856

Described in Rumphia 4 (1849) 29. K. L. Blume. 6. A. pictum: As a Hoya: In Flora Indiae Bataviae 2 (1856) 524-525. Zollinger & F. A. W. Miquel. Calyx not mentioned in either description. Flowers sent to me by Ted Green, Kaaawa, Hawaii. Cuts originally from Ruurd van Donkelaar, The Netherlands.

Pedicel and flower. Pedicel is curved, terete, glabrous 1.05 cm. long and 0.07 cm in diameter. Corolla is revolute and deeply cut.

Pedicel, calyx and ovaries enlarged about 8X. The flower is geotropic so the pedicel would actually be hanging down not as here upright. Calyx: is flat very small overlapping triangles, glabrous and membranous. 0.08 cm. long with base 0.08 cm. also. No ligules seen. Ovaries: 0.08 cm. tall, bottle shaped, base pair 0.09 cm. wide.

260

Hoya pimenteliana Kloppenburg 1999 Described in Fraterna 12/3:1999, 7-12. "Calyx: lobes narrowly triangular, sparse fine cilia, rusty colored, overlapping ¼ at base, 0.28 cm. in diameter, lobes 0.1 cm. long ca. 0.1 cm. wide, from a short thick base, lobes thick stubby, ligules not noted."

Pedicels: filiform, glabrous, light creambuff colored with varying amounts of purpling. 1.5 cm. long with fine lenticels, glabrous, terete, 4-18 in number. Ovaries: domed, 0.14 cm. tall, base pair 0.1 cm. wide, buttery yellow in color.

261

Hoya plicata King &. Gamble 1908 Described in Journal of the Royal Asiatic Society, Bengal Branch 74/2 (1908) 578. “Flora of the Malayan Peninsula King” & Gamble. "Calyx small; lobes ovate, acuminate, hyaline; scales subulate." It is possible the pictures below are not of this species.

Side view of the pedicel, calyx, and ovaries. The pedicel is curved, glabrous but punctate. Sepals are small and essentially triangular with narrowly rounded apices. Ovaries taped in toward the apex slightly, smooth and glabrous, yellow colored. Calyx: very small, with sepal overlap about 1/3. Sepals 0.07-0.15 cm. long, widest 0.10 -0.15 cm., narrow linear white ligules, center thickened, surface punctate and inside granulose.

Side view of the pedicel, calyx, and ovaries. The pedicel is curved, glabrous but punctate. Sepals are small and essentially triangular with narrowly rounded apices. Ovaries taped in toward the apex slightly, smooth and glabrous, yellow colored. Pedicel: curved about 2.8 cm. long but variable in length longest 3.0 cm. shortest 1.8 cm., 0.12 cm in diameter, terete, lenticeled and spotted black with a few short curved white celled hairs.

262

View of the calyx still on the back of the flower. The sepals are very small, triangular, dark purple here. The apex does not reach the corolla sinus.

263

Hoya polilloensis Kloppenburg, Mendoza & Ferreras ISSN 2329-7336 In Jour. Nat, Studies 12(1): 19, 2013.

Calyx: enlarged ca. 30x, sepals are ciliate, granulate outside, inside slick glabrous. Sepals 0.10 cm long with obtuse apex, 0.07 cm at the widest, ligules are present. Ovaries: columnar glabrous, apex with small protrusions, 0.10 cm tall and base pair 0.09 cm wide.

264

Hoya polystachya Blume 1849 Described in Museum Botanicum Lugduno-Batavum 1 (1849) 45. C. L. Blume. 106.

Photo of outside of corolla with attached Calyx enlarged about 16X. This small calyx conforms to Blume’s drawing (side view). None of descriptions mention the calyx.

Side view of the calyx and portion of the pedicel enlarged about 16X. Note the short sepal lobes with hirsute indumentum, also the enlarged knob below the calyx. The pedicel is terete and lenticeled otherwise it seems glabrous. (some hairs on swelling).

265

A rather dark photo, top view of the calyx and ovaries. The sepals are triangular and very small, ligules are present. The apex is rounded.

266

Hoya poolei White & Francis 1928? Described in Proceedings of the Royal Society of Queensland 39 (1928 ?) 69. “Plants of the Mandated Territory of New Guinea” C. T. White & W. D. Francis (t.5). "Calyx about 4 mm. in diameter; lobes lanceolate."

Top view of the calyx (one natural one dyed). The sepals are very short, triangular, translucent edges, with broad flat ligules at the sinus areas.

Calyx: small triangular sepals, outspread, glabrous, membranous; wide flat ligules (look like precursors to doubles) they have a little dent on the outer end. Pedicel and calyx with ovaries enlarged about 16X. This is very similar to the calyx of Hoya eitapensis Schlechter. The pedicel is stout, sepals lay out nearly horizontal, ovaries are dome shaped. Here the pedicel is a little rough with scattered hairs. Here again there seems to be a conflict with the type description. Pedicel: Terete, a few scattered hair cells 0.13 cm. long, 0.10 cm. in diameter. Ovaries: 0.16 cm. tall base of pair 0.10 cm.

267

Hoya pseudoleytensis Kloppenburg, Mendoza, Guevarra & Carandang 2013 ISSN #1055-4564

Calyx: side view enlarged ca. 22x, sepals are triangular, outer surface fine granulate and glabrous, inside is slick glabrous, ligules prominent exceeding the sinuses in some instances, 0.09 cm long and 0.06 cm at the widest. Ovaries: columnar, glabrous, 0.11 cm tall and base pair 0.06 cm wide.

268

Hoya pseudolittoralis Norman 1937 Described in Brittonia 2 (1937) 328. C. Norman. British Museum of Natural History, London. "Calyx lobes minute linear." Photomicrographs of mine follow taken from flowers sent to me by Ann Wayman, Central Point, Oregon. 5/20/90:

Pedicel and calyx enlarged about 16X. I do not have measurements for this species. Note, however the pedicle is relatively large compared to the calyx. Pedicel is glabrous, terete with a few lenticular ridges. The sepals are ciliate, triangular with outer apex acute.

Side view of the calyx showing the domed ovule pair. The calyx lays flat and the sepals are ciliate.

269

Hoya pubicalyx Merrill 1918 Wrong as this is incorrectly determined: Now Hoya pubicorolla Kloppenburg, Mendoza & Fereras. Published in 2013 New Hoya e-pub. Described in Philippine Journal of Science Vol. 13 C5 (1918) p.131.

Hoya pubicorolla Kloppenburg, Mendoza & Ferreras ISSN #2329-7336 Published in New Hoya Volume 1-2, 2013, rare-hoyas.com "Calyx-segments oblong-ovate to ovate-lanceolate, acute or subacute, externally somewhat pubescent, about 4 mm long." Calyx. Inside glabrous, bases of sepals overlap, ligules present. Ligules in this species variants are very variable, from long slender exceeding the sinuses, to short as here and in this case with 2 missing. Edges ciliate apices narrowly obtuse. Ovaries, dome shaped with separation on the inside surfaces near the apex. 0.25 cm. tall and base pair 0.20 cm.

Calyx, same as above. Outside pubescent, ciliate; inside glabrous, overlap about 1/3. Calyx: apex reaches the corolla sinus. Sepals 0.48 cm. long and 0.18 cm. at the widest. Diameter about 1.0 cm. flattened.

270

Hoya pubicorola subsp. anthracina Kloppenburg, Ferreras & Mendoza ISSN #2329-7336 Published in New Hoya Volume 1-2, 2013, rare-hoyas.com.

Calyx: side view enlarged ca. 15X, the base is pubescent, as is the sepal outer surfaces, also ciliate.

Calyx top view enlarged ca. 12X, the inner surface is slick glabrous, sepals 0.20 cm long and 0.20 cm at the widest, ligules are at the sinuses 1-11-2-2 pattern Ovaries: are columnar, glabrous, 0.24 cm tall and base pair 0.15 cm wide.

271

Hoya pubicenta Kloppenburg, Mendoza & Ferreras ISSN #2329-7336

Published in New Hoya Volume 1-2, 2013, rare-hoyas.com.

Calyx: side view on pedicle with ovaries enlarged ca. 10x. The sepals are triangular with a narrow rounded apex, 0.35 cm long and 0.20 cm at the widest, base overlap ca. 1/3, ligules are present. Outside with scattered indumentum, inside slick and glabrous. Ovaries: domed with narrowing apices, glabrous 0.13 cm tall and base pair 0.18 cm wide Pedicel: curved, terete a few scattered hair cells mostly near the calyx. 2.9 cm long and 0.11 cm in diameter.

Two sepals enlarged ca. 14x, details as above. Here showing the basal overlap and the slick dorsal surface.

272

Hoya purificacioniae Kloppenburg & Siar

Pedicel and calyx side view enlarged about 24X. The sepals a sub- linear 0.22 cm. long and 0.13 cm. at the widest, near the base. Outside granulate inside slick, glabrous.

Inside view of the calyx enlarged about 24 X. Sepals are thin, translucent taper to a narrow sub acute outer apex, ligules are present.

273

Hoya purpureofusca Hooker 1850 Described in Botanical Magazine (1850) Tab. 4520. (Curtis’s). "Calyx of five deep, almost subulate, segments." Synonymous with Hoya cinnamomifolia Hooker.

Top view of the calyx enlarged approximately 16X.

Side view of the calyx and a portion of the pedicel. Enlarged about 16X. Here again as with Hoya cinnamomifolia Hooker the outer surfaces are granular, however here the calyx lobe is more pointed, not as rounded.

274

Hoya querinoensis Kloppenburg & Siar 2007 Published in Fraterna 20/4. This sp. collected by Maximo Wayet at Baguio Village, Querino 15 Sept. 1988, 150 m elevation, flower white.

Outside view of the calyx enlarged about 8X. Sepals are long and narrow, ciliate, centrally a little thickened. Below center is the cut off pedicel. Apex is acute, a slight overlap at the base.

Inside view of the calyx, again the cut off pedicel is in the center below. Only a slight overlap at the base of the sepals, ligules are present, all glabrous. Sepal: apex to base 0.15 - 0.17 cm. long and at base 0.05 cm.0.09 cm. wide. Edge with a few cilia. Ovaries: 0.07 cm. tall, base pair 0.06 cm. wide, glabrous.

275

Hoya quinquenervia Warburg 1904 Described in Fragmentia Florae Philippinae (1904) 132 R. Schlechter & 0. Warburg. "Sepals 0.1 cm. long, ovate acute, glabrous, margins very minutely ciliate."

Top view of calyx. Note: the translucent lobes with broad bases overlapping at least 1/3. The apex is rounded but narrowed from the base. Calyx: No ligules seen. Base 0.12 cm. wide, length 0.123, Ovoid. Widest 0.13 cm. Diameter 0.32 cm.

Top view of calyx and pedicel. Note: the sepals are relatively small arising from the expanded end of the pedicel which is round (terete) and rough from small vertical lenticels. Color pale yellow. Pedicel: Terete, filiform, curved, glabrous, granulose. 2 cm. long and 0.1 cm. in diameter

Ovaries: Base 0.11 cm. and 0.07 cm. tall.

276

Hoya quisumbingii Kloppenburg, 1992 Data from Herbarium Type sheet PNH # 79412.

Described in Fraterna

Pedicel, calyx and ovaries enlarged about 16X. Sepals glabrous with a few cilia, broadly triangular with bases overlapped about 1/3, 0.15 cm. widest 0.18 cm. long. Apex narrowly rounded, membranous. 1-2 ligules at each sinus.

277

Hoya recurvula Kloppenburg 2000 Described in Fraterna 13/4:2000, 9-12.

Top view of the calyx. The edges are sparsely ciliate, bases of sepals overlap very little, texture is membranous. Calyx small, short, granulate outer surface with a few hair cells, inside glabrous, membranous. Sepals barely visible (more like protuberances), 0.15 cm. long widest 0.19 cm., variable, 1/3 overlap, ligules at base of

Pedicels straight, terete, glabrous 2.7 cm. long 0.11 cm. in diameter. Ovaries short 0.15 cm. high and 0.11 wide at the base of pair.

Side view of the pedicel, calyx and ovaries. Sepals are flat, granulose outside with a few scattered hairs, inside glabrous. Ovaries are dome shaped, short and with wide base. Ligules are present but difficult to discern.

278

Hoya recurvula subsp. bokorensis Kloppenburg & Yap

Published in Asklepios #108, 2010.

Outside surface of a flower enlarged about 8X. The sepal apices almost reach the corolla sinuses, corolla outer surface is glabrous, is deeply cut.

Outside surface of the calyx enlarged about 8X, Surface is finely granulate glabrous, sepals overlap at the base about 1/4, apex is obtuse are 0.22 cm. long and 0.20 cm. at the widest.

Inside view of a calyx enlarged about 10X. Surface is slick glabrous. Ovaries are sub-columnar, glabrous, 0.17 cm. tall and base pair 0.17 cm wide.

279

Hoya retusa Dalzell 1852

Published: In Journal of Botany 4 (1852) 294-295. “Contributions to the Botany of Western India” N. A. Dalzell (J. D. Hooker) (Kew Gardens Misc. Pub.). Natural Ord. Asclepiadeae. Pergularieae. § Hoyeae.

Pedicel, calyx with ovaries enlarged about 8X. Pedicel is terete, glabrous 1.2 cm. long and 0.07 cm. in diameter. Calyx base a little expanded, outside surface granulate glabrous. Ovaries bottle shaped, glabrous, 0.14 cm. tall and 0.08 cm. wide base pair.

Top view of the calyx and ovaries enlarged about 8X. Sepals narrowly triangular, ligules present, glabrous and slick inside, apices narrowly rounded, only slight overlap.

280

Hoya revoluta Wight ex Hooker 1883 Published: In Flora of British India 4 (1883) 55. Wight (J. D. Hooker).

Pedicel and calyx side view enlarged about 16X. Pedicels are of various lengths 1.0 - 2.0 cm. long x 0.07 cm. in diameter, curves forming a loose geotropic umbel, terete, glabrous, slightly larger at the peduncle end.

Outside view of the calyx cut from the pedicel greatly enlarged. Calyx is small and flat. Sepals do not reach the corolla sinuses. Glabrous both surfaces but outer surface is finely granulose, central portion thickened, broadly linear. Base - apex Center - apex Base width

0.11 cm. 0.15 cm. 0.07 cm.

Inside view of the calyx. Ligules are present but difficult to detect, as they are the same color and texture as the sepals. Base of the sepals are very slightly overlapped. In removing the pedicel and calyx the ovaries remain with the corolla.

281

Hoya rhodostele Ridley 1923 Described in Journal of the Royal Asiatic Society 1 (1923) 77. H. Ridley. "Sepals short, ovate blunt." The following photomicrographs were from a flower from herbarium sheet #1545 Hoya rhodostele Ridley, collected at 5000' elevation by H. S. Yates, Sumatra Forest Brostsagi, white latex. Listed as a Topotype.

Calyx. The pedicels are relatively thick and the calyx small. Pedicels 2.0 cm. long 0.06 cm. in diameter, glabrous Sepals 0.13 cm. long, apically rounded, with some overlap and large ligules present, widest 0.10 cm.; both surfaces glabrous.

282

Hoya rigida Kerr 1939 Described in Kew Bulletin (1939) 463. A. F. G. Kerr. "Calyx 5-parted high, glabrous, towards the outer base minutely papillose, inside minutely glandular; lobes linear-lanceolate, somewhat acute, 6 mm. long, bases 1.5 mm. wide."

Top view of the calyx. The sepals are ciliate, wide in the middle tapering to somewhat narrow rounded apices, and overlap about 1/2. There is a concave central cup around the ovaries. Ligules are present but difficult to detect. Calyx: 0.24 cm. long; widest 0.18 cm., punctate outside, glabrous inside, edges jagged, ¼ overlap, cupped near ovaries with a few cilia. Ligules present. Side view of a portion of the pedicel and the calyx with the ovaries enlarged about 16X. The pedicel is round and expands in a taper to the calyx base, and the surface has short sparse set hairs. The sepals lay flat, and are ciliate. The ovaries are unusually short and stubby with rounder apices. Pedicel: 2.3 cm. long; 0.11 cm. in diameter, strict, terete, hair cells hooked toward the apex. Ovaries: 0.18 cm. tall and base of pair 0.15 cm., short domed.

283

Hoya salmonea Kloppenburg, Mendoza & Ferreras 2013 Pub. In Jour. Of Nature Studies 12 (1) 22-24 2013. ISSN 2329-7336

Calyx: linear, glabrous entire, apex sub obtuse, unusually thin, 0.19 cm long and 0.10 cm widest, overlap 0.05 cm. Ovaries: broad domed, glabrous 0.12 cm tall and base pair 0.05 cm wide.

Ovaries apex greatly enlarged to show an apex with some modifications from the usual, here the end narrows, than apex has a center knob and side shelves.

284

Hoya salmonea subsp. pallida Kloppenburg, Mendoza & Ferreras 2013 ISSN 1055-4564

Calyx: enlarged ca. 32x, the sepals are linear with a few cilia, 0.13 cm long and 0.04 cm at the widest, calyx base is globular (bulbous). No ligules observed.

Ovaries: are stubby, glabrous 0.06 cm tall and base pair 0.06 cm wide, the apices are modifies. Ovary apices photo to the right showing apex modification here an extended rather flat ring like structure.

285

Hoya samarensis Kloppenburg & Siar 2012

Calyx: 0.13 x 0.085 cm glabrous, 1/3 overlap, small no ligules observed.

Sepals are triangular, here enlarged about 8X. Ovaries are short broad based, not measured.

286

Hoya salweenica Tsang & Li. 1974

Described in Acta Phytotaxanomica 12/1 (1974) 125. J. Tsiang & P. T. Li.

Side view of the calyx, and ovaries enlarged about 8X. Outer surface of calyx and pedicel sparsely pubescent. Ovaries columnar domed. Sepals inside glabrous and shiny surfaced, thickened in the middle, edges membranous, slightly cupped.

Top view of the calyx enlarged about 8X. Sepal bases overlap about ¼, ligules are present. Apices tapering to narrow rounded end, not sharply acute. Type description says no glands.

Corolla outside with calyx attached enlarged about 8X. Sepals with wide bases, slightly hirsute especially at the calyx base. Sepals do not barely reach the corolla sinuses. Corolla turns under at the sinuses. Outside glabrous inside thickly pubescent.

287

Hoya smithii Kloppenburg 2010

Calyx: Sepals 0.30 cm long. very pubescent outside, glabrous inside 0.60 cm in diameter. No ligules noticed. Calyx on the back of the flower enlarged about 8X. Sepal raised by a thin sewing needle long and narrow. Surface is actually hirsute. Apex just reaches the corolla sinus.

Calyx outside surface enlarged about 8X. One sepal torn free. Very hirsute on this surface, lobes long triangular apices acute.

Inside view of the calyx enlarged about 8X. This surface is slightly cupped in the center under the ovaries (here deteriorated) This surface is glabrous but the hirsute cells show through the thin sepals.

288

Hoya samoensis Seemann 1866

Described in Flora Vitiensis (1866) 163. B. Seemann.

View of the pedicel and calyx on the flower enlarged about 8X. Pedicel is glabrous, terete, (here) translucent 0.7 cm. in diameter tapering larger as it approaches the calyx. Sepals extend only1/2 way to the corolla sinuses.

Inside view of the calyx and pedicel enlarged about 8X. Sepals are rather broad 0.16 cm. at the widest 0.16 cm long; 0.20 cm. long to the center. I did not observe any ligules. Ovaries are narrowly dome shaped 0.10 cm. tall and the base pair 0.08 cm. wide.

289

Hoya santiagoi Kloppenburg & Siar Described in Fraterna 24(4): 29 (-31) 2011. Calyx: linear to triangular 0.34 cm x 0.15 cm ciliate and pubescent more so than the pedicels, do not reach the corolla sinuses.

Hoya santiagoi subsp. mandozai Kloppenburg 2013 ISSN #2329-7336 Described in New Hoya Volume 1-3, 2013, rare-hoyas.com

Side view of the pedicel. calyx & ovaries enlarged about 8x. Pedicels: are very finely puberulous to glabrous, 3 cm long. terete, filamentous, curved at least 27 on this cluster. Calyx: pubescent outside, apex acute base broad 0.17 cm long x 0.10 cm at the base. No ligules observed.

Ovaries: columnar, glabrous, 0.17 cm long.

290

Hoya savaiiensis Kloppenburg 2010 Described in Fraterna 22 (4): 2009

View of the outside of the flower enlarged about 8X. The calyx lobes are very small and does not reach the corolla sinus. The pedicel is glabrous, terete, 2.4 cm. long 0.07 cm. in diameter.

Outside and inside view of the pedicel and calyx including the ovaries enlarged about 8X. The calyx is glabrous, apex to base 0.17 cm. long and 0.15 cm at the widest. Sepal apex is narrowly rounded, no ligules were observed. Ovaries narrowly domed, 0.17 cm. tall, glabrous.

291

Hoya scortechinii King & Gamble 1903 Described in Journal of the Royal Asiatic Society, Bengal Branch. 74/2 (1903) 567 (777). King & Gamble “Flora of The Malay Peninsula”. “Calyx small; lobes ovate, ciliate, membranous, glandular, recurved, .03 in. long.” The following photomicrographs are taken from flowers sent to me from Ted Green, Kaaawa, Hawaii, 8/2/94. This specimen does not conform to the Type description in that the inner lobes here are covering the anthers or nearly so. In addition it says of the pollinarium “attached by short caudicles (translator arms). Ours could not be considered short! It is difficult to say if we have a variant from the Type or a different species.

Top view of the calyx enlarged about 8X. Sepals are ovate with rounded apices, some slight ciliation; very thin and membranous. Calyx is flat rotate, does not reach the corolla sinuses, sepal edges ciliate, apex rounded 0.16 cm. long and at base 0.16 cm. wide with slight overlap up to 1/3; with indistinct ligules.

Pedicel, calyx and ovaries side view enlarged about 8X. The pedicel is short 0.90 cm. long and 0.08 cm. in diameter. Glabrous, yellow with raised red lenticels and striations Ovaries dome shaped 0.12 cm. tall, 0.09 cm. at the base of pair, glabrous.

View of the outside of the corolla with pedicel and calyx still attached enlarged about 8X.The corolla outside is glabrous crystalline, inside puberulous. A thickened raised collar in center.

292

Hoya siariae Kloppenburg 2002 Now a subsp. of Hoya blashernaezii Feb. 2014 Described in Fraterna 15/3:2002. 1-4.

Side view of a closed flower showing the calyx and pedicel enlarged about 8X. Pedicel is glabrous, 2.6 cm. long, terete, 0.10 cm. in diameter, rusty pink colored. Calyx is ciliate otherwise glabrous, inside and out, calyx base is rugose. The sepals do not reach the corolla sinuses.

Side view of the calyx and pedicel enlarged about 8X. The pedicel has a few scattered hairs, twisted and bent over. Calyx inside is waxy and glabrous. The two ovaries are short columnar 0.15 cm. tall and the base pair 0.10 cm. wide.

Calyx top view enlarged about 8X. Sepals are a little overlapped at the base, 0.20 cm. long and 0.17 cm. at the widest, edges are ciliate, apex narrowly rounded subacute. Small ligules are present at each sinus area

293

Hoya sigillatis T. Green

Described in Fraterna 17(3): 2 2004.

Pedicel calyx and ovaries enlarged about 8X. The pedicel and base to calyx have single-celled clear hair cells pointing apically. Calyx outside is granulose, inside shiny glabrous. Ovaries are 0.05 cm. tall and base pair are 0.10 cm. wide, glabrous, yellow.

294

Hoya sipitangensis Kloppenburg & Wiberg 2002 Described in Fraterna 15/3:2002, 1-6

Side view of the pedicel, calyx and ovaries. The sepals are very short with little overlap; ligules present. Sepals 0.08 cm. long. 0.26 cm. diameter. Sepal edged incised and an occasional cilia.

Pedicel and calyx with ovaries. Pedicels glabrous but granulose surface same for outside of calyx. Pedicels are curved, short, glabrous or extremely finely puberulent, terete. 0.80 cm. long, 0.09 cm. in diameter. Ovaries rather long and narrow.

295

Hoya stoneana Kloppenburg & Siar 2006

Described in Fraterna 19/4: 2006, 19-22.

Pedicel calyx and ovaries side view enlarged about 16X. Pedicel is lime green, widest at calyx, calyx lobes sticking straight out, glabrous, Ovaries glabrous, shiny surface, shot domed, 0.15 cm. tall and base pair 0.14 cm. wide.

Top view of the calyx and ovaries. Sepals are triangular with membranous edges, thickened in central region, outside tinged rust color, outer surface granulate, apex round- nearly acute, does not cone even half way to the corolla sinuses. 0.23 cm. long 0.12 cm. wide at the base, ligules present.

296

Hoya subquintuplinervis Miquel 1869 Described in Annales Museum Botanicum Lugduno-Batavum 4 (1869) 141. P. A. G. Miquel. "with sepals ovate obtuse and puberulous"

Top View of the calyx. Note: The calyx lobes are overlapped about 1/3 at the base. The edges are only slightly ciliate, Apex rounded, the ligules at the base of sinus of the lobes are very small and not noticeable here. Calyx: 0.23 cm., but variable length, shortest is only 0.117 cm. long. Widest 0.18 cm. Lobes a narrow triangle, ¼ overlap, ligules present.

Side View of the calyx. Note: the very short calyx lobes and stubby short ovaries, also the enlarging of the pedicel as it approaches the calyx. Pedicel: Straight to somewhat curved, glabrous, terete, fairly smooth with a few lenticels, it enlarges as it proceeds toward the calyx. 0.11 cm to 0.15 cm in diameter. Ovaries: 0.20 cm. high base 0.013 cm, domed columnar.

297

Hoya subquintuplinervis Miquel 1869 Published: In Annales Museum Botanicum Lugduno-Batavum 4 (1869) 141. View of the pedicel and calyx on the corolla enlarged about 16X. Pedicel is terete, glabrous, 1.0 cm. long 0.10 cm. in diameter. Calyx outside with short pubescence, ciliate. The sepal apex does not reach the corolla sinus.

Side view of the pedicel, calyx and ovaries enlarged about 16X. the base of the calyx is a little enlarged. Ovaries are domed with a fairly flat top. 0.15 cm. tall, glabrous, base of pair 0.13 cm. broad.

Outside surface of the calyx enlarged about 16X. This surface is rough, some surface hairs and ciliate, there is a slight overlap ao the sepals.

Top view of the calyx enlarged about 16X. The sepals inside are glabrous and shiny. Center to apex 0.35 cm.; base to apex 0.25 cm., 0.16 cm. widest.

298

Hoya surigaoensis Kloppenburg, Siar & Nyhuus 2010 Described in Asia Life Sciences 2010

Pedicel, calyx and ovaries enlarged about 16X. Pedicel is terete, straight, glabrous, lenticulate and slightly enlarged under the calyx. 1.7 cm. long and 0.09 cm. in diameter. Sepals are narrowly triangular, outside granulose inside glabrous and smooth, 0.14 cm. long 0.11 cm. widest at base, ligules present Ovaries are short domed shaped, glabrous, 0.10 cm. tall and base pair 0.10 cm. wide.

Bottom view of the flower with the calyx intact enlarged about 16X. The sepals reach only about 1/2 way to the corolla sinuses. Corolla surface glabrous. The corolla lobes are tightly curved under from the sinuses toward the apex leaving a channeled effect, with the apex turned under, Corona apexes are just slightly showing at the sinus areas. Sinus – sinus Sinus – center Sinus – apex Apex – center Widest

299

0.25 cm. 0.21 cm. 0.45 cm. 0.60 cm. 0.39 cm.

Hoya tannaensis T. Green & D. Kloppenburg

Calyx and ovaries on the pedicel enlarged as above, base slightly enlarged, surface is finely granulate, sepals are short triangular outer apex sub acute. 0.17 cm. long and base 0.12 cm wife, with a slight overlap.

Calyx top view enlarged ca. 10X. Ligules are present, inside surface is glabrous, slick. Ovaries are columnar, glabrous, 0.18 cm. tall and base of pair is 0.15 cm. wide, apex reddish color.

300

Hoya tauensis Kloppenburg 2009

Described in Asklepios 111: 14, 2011.

View of the corolla with the calyx and pedicel attached enlarged about 8X. Pedicel glabrous, terete, 2 cm. long x 0.45 cm. in diameter. Sepal apex nearly reaches corolla sinuses. Corolla outside glabrous, deeply cut.

Pedicel and calyx enlarged about 8X. Domed shaped pair of ovaries showing. Calyx is glabrous, thin, with striate vascular channels visible. Sepal sinuses have linear ligules present, 0.18 cm. long 0.20 from apex to center.

301

Hoya telosmoides Omlar 1996 Described in Nova 6(3)1996.

Pedicel, calyx with ovaries enlarged about 8X. Surface of calyx base glabrous, granulose and expanded from pedicel. Sepals membranous, short triangular, 0.10 cm. long 0.15 cm. at the widest, ligules present inside at the sinuses, do not reach the corolla sinuses. Ovaries: glabrous bottle shaped, 0.25 cm. tall and 0.16 cm. at the base of the pair.

302

Hoya tiatuilaensis Kloppenburg ISSN 2329-7336

Outside view of the flower enlarged about 8X. Pedicel is glabrous, terete, short, 1.05 cm long 0.05 cm in diameter, Calyx is very large, outside granulose, inside smooth, sepal apex reaches the corolla sinuses. Flower not yet opened.

Calyx: Left outside view, right inside view enlarged about 8X., glabrous. Sepals 0.13 cm long, 0.13 cm at the widest, near the base. One ligule seen.

303

Hoya tomataensis T. Green and Kloppenburg 2004

Described in Fraterna 17.2:2004, 1-5.

Outside view of the flower enlarged about 8X. The pedicels are glabrous 1.45 cm. long and 0.07 cm in diameter. Sepals reach about 3/4 the way to the corolla sinuses.

Pedicel, calyx and ovaries in side view enlarged about 16X. Pedicel near the calyx becomes granulose and slightly expanded. The sepals are narrowly triangular outside with a few cilia otherwise granulose. Inside shiny and smooth. Ovaries are glabrous, shortly domed 0.10 cm. tall and base pair are 0.11 cm. wide.

Top view of a calyx enlarged about 16X. Sepals are a little overlapped at the base, not much thickened centrally. There are multiple ligules at the sinuses. Apex - base Apex - center Widest

304

0.15 cm. 0.18 cm. 0.09 cm.

Hoya tsangii Burton ex Kloppenburg 1988

Described in The Hoyan 9 #4 (1988) 2:.i-ii. "calycis segmentis oblongis, obtusis, extus pubescentis et ciliatis." Burton " calycis segmenta ovata triangulata, glabra" Kloppenburg

Top view of calyx enlarged approx. 16X. Note: The sepals are broad and rounded, the whole structure is relatively small.

Calyx: 0.10 cm. long base 0.05 cm., edge membranaceous, undulant (possibly from drying). I did not observe any ligules, although they are usual present but sometimes difficult to detect.

305

Hoya tiatuilaensis Kloppenburg ISSN 2329-7336

Described in New Hoya Vol. 2-1, Feb. 2014, rare-hoyas.com. e-published.

Outside view of the flower enlarged about 8X. Pedicel is glabrous, terete, short, 1.05 cm. long 0.05 cm. in diameter, Calyx is very large, outside granulose, inside smooth, sepal apex reaches the corolla sinuses. Flower not yet opened.

Calyx: Left outside view, right inside view enlarged about 8X., glabrous. Sepals 0.13 cm. long, 0.13 cm at the widest, near the base. One ligule seen.

306

Hoya ubudensis Kloppenburg & Yap Published is Asklepios 108, 2010.

Calyx on the corolla enlarged about 8X. The sepals do not reach the corolla sinuses, extend about ½ way. The base is swollen and granulose puberulent.

Outside view of the calyx enlarged about 8X. The sepals overlap ca. ½ this surface is granulose, puberulent and sepals are ciliate. Sepals are 0.15 cm. long and 0.15 cm. at the widest. The sepal shapes are intermediate between ovate and triangular.

Inside view of the calyx enlarged about 16X. Ligules are present, sepals widest just above the attachment area, inside glabrous slick. Ovaries are glabrous short domed, 0.13 cm. tall and base pair 0.12 cm. wide.

307

Hoya unruhiana Kloppenburg, Siar, Mendoza, Cajano & Carandang 2013

ISSN # 2329-7336

Calyx: side view enlarged ca. 25x. Sepals are thin, overlap slightly at base, no ligules, surface finely granulate, extend ½ or less to the corolla sinuses; 0.11 cm long and 0.08 at the widest.

Side view of a flower enlarged ca. 18x. Corolla outside glabrous and rotate when fully open. Sinus – sinus Sinus – center Sinus – apex Apex – center Widest

308

0.18 cm 0.14 cm 0.27 cm 0.37 cm 0.23 cm

Hoya valmayoriana Kloppenburg, Guevarra, & Carandang ISSN 1655-3179

Calyx: side view enlarged ca. 16x. Sepals are glabrous inside and out, sub-linear, overlap slightly at base, apices are obtuse, 0.22 cm long x 0.10 cm wide at base, ligules present. Ovaries: dome shaped, glabrous, 0.16 cm tall and base pair 0.11 cm wide.

309

Hoya vicencioana Kloppenburg, Siar, Cajano & Carandang 2013 ISSN 10055-4564

Calyx: side view enlarged ca. 25x. Sepals have a few hair cell on the outside surface, otherwise glabrous, inside glabrous slick, they reach the corolla sinuses, 0.14 cm long and 0.10 cm at the widest.

Ovaries: columnar with reduced apices, glabrous 0.09 cm tall and 0.08 cm wide at base of pair.

Outside view of a flower enlarged ca 18x to show the sepals beyond the corolla lobe sinuses. Corolla outside is glabrous. Corolla is reflexed, off yellow color.

310

Hoya vitellina Blume 1849 Described in Museum Botanicum Lugduno-Batavum 1 (1849) 45. C. L. Blume. Calyx not mentioned in the type description.

Top view of the calyx. Outside some hairs, edge ciliate, centrally thickened, glabrous inside. Sepals 0.15 cm. long ca. 0.13 cm widest. No ligules seen.

Pedicel, calyx and ovaries side view. Pedicel is straight, terete with scattered hairs curved and bent apically. 2.10 cm. long 0.11 cm. in diameter, enlarging slightly as it approaches the calyx. Ovaries domed, short, shot tips at the apex

311

Hoya walliniana Kloppenburg & Nyhuus 2003

Described in Fraterna 16/4:2003. 9-13.

Pedicel, calyx with ovaries enlarged about 16X. Pedicel: 0.7 cm. long, of variable lengths, to form a flat cluster, deep green lenticeled, rigid, rachis circular with fine bracts layered evenly, surface tan.

Outside view of a flower showing the small calyx; lobes of which reach only about 1/2 way to the corolla sinuses. Corolla is revolute, glabrous on this outer surface. Inner surface is puberulous with a large apical area glabrous. Enlarged about 16X.

Calyx and ovaries greatly enlarged. The sepals are small, centrally thickened with very short basal overlap, both surfaces are glabrous. Sepals 0.15 cm. long; from center 0.15 cm. Small dark ligules are present. Ovaries are glabrous, narrowly domed shaped with flaring apex. 0.15 cm. tall and base pair 0.06 cm. wide.

312

Top view of the calyx enlarged about 16X. The sepals are triangular shaped with a small basal overlap. Edges are irregular (slightly) and an occasional cilia, Small dark ligules are present.

313

Hoya wayetii Kloppenburg 1993 Described in Fraterna 1993/2:9-10. "The calyx segments are larger in this species than the former species, the apex rounded, the central width greater, the margins ciliate and with broad flat ligules and each overlap."

Calyx: lobes 0.17 cm long x 0.13 cm. at widest (at overlap), bright pale green, with ciliate margins, apex rounded, inside glabrous, outside granulose, broad flat ligule at sinus 0.03 wide x 0.02 long.

Pedicel: 1.0-2.0 cm. long 0.11 cm. diameter, curved variable in length, glabrous medium shiny green.

314

Hoya waymaniae Kloppenburg 1995 Described in Fraterna, International Hoya Association, 2 (1995) 8-9. R. D. Kloppenburg.

Calyx top view, very little overlap of sepals at their base, edges membranous. Ligules are present, Surface glabrous. Calyx: elliptic triangular, 0.8 cm. x 0.16 cm. at widest portion near the base, surface glabrous, outer punctate, ligules present.

Side view of the pedicel, calyx and ovaries. Pedicel is curved 2.0 cm. long with scattered hair cells and very finely puberulent, deep orange, 0.14 cm. in diameter. Ovaries: Domed 0.19 cm. tall and base pair 0.12 cm. wide.

315

Hoya whistleri Kloppenburg 2002 Described in Fraterna 15/1:2002,7-10. "calycis ovatis triangularis." Photomicrographs from flowers sent to me by Ted Green, Kaaawa, Hawaii 2/3/95. Collected by Art Whistler on Tau Island, Samoa. Flower large, lime green color.

Top view of the calyx. The sepals are somewhat irregular, ciliate with small dark ligules at the bases (sinuses). Apex is tapered obtuse with broad bases and slight overlap. Calyx cupped inwardly (apically), receptacle enlarged knobby and granulose, rough with some pinkish tones. Inner surface glabrous. Sepals broad and short, ciliate 0.19 cm. long and 0.18 broadest, overlapped at the base 0.06 cm.; small dark ligules are present.

Pedicel, calyx and ovaries side. Pedicel curved 2.5cm. long by 0.10 cm. in diameter, terete, glabrous with many longitudinal lenticels so surface is rough. Ovaries domed 0.17 cm. tall, base of pair 0.16 cm.

316

Hoya sp. 80-03 This species remains as of now 7/26/01 unnamed. It was thought to be Hoya chunii Li (Schlechter’s reticulata) See that file it is not that species. It was also suggested to be Hoya tjampeaensis Hochreuter (1936) although a large leafed species all the flower parts are much smaller in that species.

Pedicel, calyx and ovaries side view. Pedicel is curved, terete, glabrous. Calyx flat 0.50 cm. in diameter. Sepals finely granular outside, glabrous inside ciliate 0.25 cm. long, ligules present.

Calyx on outside of corolla. The sepals are very short with a thickened rugose base, The come no where near the corolla sinuses.

317

Hoya sp. laurifolia Miquel Described in Annales les Science Naturelles 9 #2 (1838) 272. Decaisne.

Pedicel, calyx and ovaries side view enlarged about 8X. The pedicel is 0.22 cm. long and 0.19 cm. in diameter, yellow, terete, glabrous enlarging slightly toward the calyx. Ovaries are tapered narrowing toward the apex, glabrous.

Top view of the corolla enlarged about 8X. The sepals are ciliate, glabrous inside, overlapped a little at the base. The ligules are plainly visible as darker spots and sepal sinuses. I did not take measurements. Since the pedicel diameter was measured it other measurements could be deduced.

Corolla outside with calyx attached enlarged about 8X. This corolla surface is glabrous, calyx is also glabrous, the sepals are ciliate and do not reach the corolla sinuses, corolla appears to be cut about half way. Again no measurements.

318

Hoya sp. as rumphii The following pictures are from a fresh flower supplied by Jerry Williams, Rainbow Gardens, Vista, California. The plant was originally from Ted Green, Kaaawa, Hawaii, labeled as Hoya rumphii. It is not that species, so what is it?

Top view of the calyx. The sepals are long and elliptic with ragged edges and ciliate; there is very little overlap at the base; edges are membranous and thickened internally. Small ligules are present. Sepals 0.17 cm. long. Inside streaked with red especially along the edges.

Side view of the pedicel, calyx and ovaries. The base of the calyx is expanded. 0.22 cm. in diameter with a rough surface. The ovary pair are somewhat bottle shaped. 0.19 cm. tall 0.12 cm. wide at the base 0.055 cm. wide at the apex.

319

Hoya sp. Bicknell 1994 Flower cluster sent by David Bicknell from Lilo-an, Cebu, Philippines. Collected by David January 1998 at Askoy, Cebu in forest, elevation 500 m. epiphytic. Flowers pale purple, 8 mm. Pedicels 2 ½ cm. pale green with purple flecks, glabrous. Calyx pale purple. Peduncle 13 mm. Cluster 28 flowers, geotropic. Section Acanthostemma (Bl.) Kloppenburg,

Pedicle, calyx and ovaries side view. Ovaries cone shaped 0.13 cm. tall base pair 0.07 cm. wide. Calyx: sepal apices acute 0.10 cm long and 0.08 cm. at base. Sepals outside granulose inside glabrous and shiny.

320

Hoya sp. BSI #1 Material sent via Ann Wayman, Photographed 14 February 1995. Compare this with Hoya cominsii Hemsley.

Top view of the calyx. Small overlap of the sepal bases, no ligules observed, a few cilia. 0.18 cm. long and widest 0.15 cm. Inside glabrous and waxy, rather triangular shaped with tapered rounded apices.

Side view of the pedicel, calyx and ovaries. Pedicel is dark red color. 2.2 cm. long, 0.10 cm in diameter, glabrous slightly larger at the calyx base. Ovaries columnar (odd shape here maybe slightly dried) 0.20 cm. tall and base pair 0.11 cm wide; glabrous.

321

Hoya sp. CAHUP 5965 It appears not to be Hoya bicolensis Klopp. & Siar although it is close.

Side view of the dried calyx enlarged about 16X. Here most sepals are reflexed on drying, sepals are very thin, membranous. 0.12 cm. long and 0.09 cm. at the widest. glabrous on both surfaces. Ovaries: 0.09 cm. tall and 0.07 cm wide at the base of the pair., glabrous, shortly domed.

322

Hoya sp. CAHUP 5967 Labeled Hoya incrassata. Most likely correct pollinium a little shorter.

Outside surface of the calyx enlarged about 32X. Pedicel widens as it Approaches the calyx, surface granulose, sepals short and overlapped about 1/3 at the base, apex obtuse.

Inside surface of the calyx enlarged about 32X. Sepals are broadly triangular, apex obtuse, here translucent, thin; 0.10 cm. long. Ovaries short domed 0.04 cm. tall and base pair 0.06 cm, glabrous.

323

Hoya sp. CAHUP 9136 Photos 14 November 2006 seems closest to Hoya crassicaulis Elmer ex Kloppenburg.

Back side of the calyx enlarged about 16X. Sepals are 0.12 cm. long and 0.07 cm. wide at the widest near base, thin, apex rounded.

324

Hoya sp. CAHUP 8359 (41559) Hoya imbricata

Outside surface of the calyx enlarged about X, sepals are large, apex acute, ciliate.

Inside surface of the calyx. Sepals are 0.35 cm. long and 0.18 cm at the widest, ligules are present at the base sinuses, overlap about 1/3. Ovaries are glabrous 0.23 cm. long and 0.15 cm. wide at the base pair.

Outside surface of a flower at the calyx area enlarged about 8X. Sepals reach a little over half way to the corolla sinuses. Corolla surface here glabrous.

325

Hoya sp. CAHUP 61933 Hoya mindorensis Schlechter

Calyx enlarged about 16X. Sepals are 0.16 cm. long and at the base 0.15 cm. wide with a slight overlap, thickened in the center and ciliate otherwise glabrous, outside granulose. Ovaries are columnar 0.13 cm. tall and base pair 0.12 cm wide, glabrous drying dark in color.

Pedicel and calyx on corolla. Sepals are not well differentiated.

326

Hoya sp. CAHUP 63827 23 Oct. 2006: Same as 81100 = quinquinervia Pedicel, calyx and ovaries side view enlarged about 16X. Calyx is broad triangular 0.07 cm. long 0.04 cm wide near base, overlap slightly, terete, outside granulose, ciliate. inside glabrous. I did not see any ligules. Ovaries columnar 0.07 cm. tall and base pair 0.05 cm. wide, glabrous, drying dark colored.

327

CAHUP #5297 Determined to be very close to 900307 (H. merrillii)

Outside of flower enlarged about 8X. Sepals do not reach the corolla sinuses, here dried and flared out from corolla surface which is glabrous.

Pedicel calyx and ovaries side view enlarged about 16X. Ovaries are narrow domed shape, 0.11 cm tall and 0.08 cm wide at abase pair, glabrous.

Top view of the calyx and ovaries enlarged about 16X. Sepals are glabrous inside and out, 0.13 cm long and 0.10 cm at the widest, overlap at the base about 1/4 . There are faint ligules, sepal apex narrowly rounded

328

CAHUP 18682 JP Determined to be Hoya cardiophylla same as DH 910301. Flower from Herbarium sheet for Identification.

Pedicel and calyx enlarged about 16X. The pedicel is 0.06 cm. in diameter, broken off so do not have the length. The surface is very finely crystalline puberulous. Calyx very small and ovate, 0.07 cm. long and 0.07 cm. at the widest with ca. ¼ overlap at the base, ligules present, outer surface as with the pedicel very finely puberulous and dense. inside glabrous and shiny, membranous. Ovaries, here dark 0.09 cm. tall and base pair 0.09 cm. wide, glabrous.

Outside surface of the flower enlarged about 16X. Sepals are small and do not come near the corolla sinuses. Corolla outer surface is finely granulate, glabrous.

329

CAHUP 41193 Determination: possible new species.

Side view of the calyx and ovaries enlarged about 8X. Calyx lobes do not reach the corolla sinuses are glabrous on both surfaces. Sepals are 0.10 cm long and 0.10 cm wide at the base, no ligules observed. Ovaries: short, glabrous 0.10 cm tall and base pair 0.05 cm wide.

Top view of a calyx with some sepals missing enlarged about 8X, they are thin, glabrous, triangular with broad base and rounded apex.

Closed flower outside surface with pedicel attached enlarged about 8X. Calyx lobes have turned up from the surface on drying. Outer surface of corolla is glabrous and granulose. Note the sepals do not reach the corolla sinuses and that the coronal lobe outer apex exceeds the sinus (lower right hand corner.

330

Hoya sp. CAHUP # 3935, Hernaez 2 March 1990

Pedicel, calyx and ovaries side view. Pedicel: terete, reddish color 2 cm. long, glabrous. Calyx: very small, edges ciliate, appears glabrous but very bumpy surface. Ligules present. Ovaries: narrow conical 0.06 cm. tall base pair 0.05 cm.

Outside of the flower with calyx attached. Sepals do not reach the corolla sinuses. Here corolla rolled up exposing the coronal scale bottoms.

331

Hoya sp. CAHUP #5271 Collected by Maximo Wayett at Benguet Village, Querino, Luzon, Philippines Photos 19 April 2006 Determination: near 5269 & 5293 Calyx enlarged about 16X. Sepals nearly reach the corolla sinuses, slight basal overlap; I did not see any ligules, edges entire, thin. 0.12 cm. long 0.10 cm. at the widest. Ovaries: short dome, glabrous, 0.07 cm. long and base pair 0.05 cm wide.

Flower outside showing a dark calyx and corolla lobes cupped up in drying. corolla outside glabrous, apex acute.

332

Hoya sp. CHAUP # 5293 & 5392 P. F. Hernaez 17 March 1990 Photos Roll 242 3/29/99

Calyx outside view. Sepals are centrally thick, granulose surface, broad triangular. Do not reach the corolla sinuses. 0.10 cm. long base 0.10 wide.

333

Hoya sp. IML 1056 This is a fuzzy yellow bordered flower collected 9/8/00 at Ted Green's, Kaaawa Hawaii.

Top view of the calyx. The edges are slightly irregular but not ciliate, there are well formed ligules at the sinus almost reaching the edge, apex is narrowly rounded. Calyx: outside slightly granular, inside glabrous with very fine crystalline surface, prominent small ligules at the sinuses of the sepals; small overlap of the sepals, edges entire, diameter 0.40 cm., sepals 0.13 cm. long base 0.11 cm. wide.

Side views of the pedicel and calyx. The pedicels are straight and glabrous; expanding only slightly as it approaches the calyx, that is also glabrous but granulose outside, it does not reach the corolla sinus. Pedicel: strict, terete, 2.4 cm. long., glabrous, yellow, 0.10 cm. in diameter, expanded slightly below the calyx, slightly granulose. Ovaries: paired, glabrous, dome shaped, 0.15 cm. tall; base of pair 0.11 cm.

334

Hoya sp. IPPS 8845 from Maggan Alm , Sweden 5/07 a cluster of 25 flowers in Kew solution.

Side view of a calyx enlarged about 16X. The surfaces are crystalline outside, ovaries very short.

Top view of a calyx enlarged about 16X. Sepals are broad based 0.12 cm. long and 0.10 cm. wide at the widest, slight basal overlap with ligules present, inner surface glabrous with a crystalline texture. Ovaries are short domed 0.09 cm. tall, glabrous, 0.08 cm. wide at base of pair.

Side view of flowers enlarged about 10X. outer corolla surfaces are glabrous, normally corolla is rotate, apices are acute, lobes broadest above the sinuses. Sepals are short of the corolla sinuses from a sub-expanded base.

335

Hoya sp. IPPS 8860 Umbels via Torill Nyhuus march 2007, data 3/28/07 Probably Hoya tomatoensis

Pedicel, calyx and ovaries enlarged about 16X. Pedicel is strict, terete, glabrous 2.5 cm. long, 0.06 cm. in diameter. Calyx: does not reach the corolla sinuses, edges are jagged with a few cilia, I did not find any ligules, overlap of sepals is ca. 1/8th. Sepals 0.13 cm. long , 0.09 at the widest.

Outside surface of the calyx enlarged about 8X, sepals are membranous, triangular with apices obtuse.

Inside surface of the calyx with one ovary missing, enlarged about 16X, sepals are not much thickened and with little basal overlap. Ovaries are short, domed, glabrous, 0.10 cm. tall and 0.08 cm. wide at pair base.

336

Hoya sp. IPPS 8870 Via Torill Nyhuus, March 2007. flowers in Kew solution, data 4/1/07 Probably Hoya tomatoensis

Pedicel, calyx and ovaries enlarged about 16X. Pedicel is strict, glabrous, 1,5 cm. long and 0.06 cm. in diameter. Calyx base bulbous, sepals glabrous, slight overlap at the base, triangular with obtuse apex. Ovaries, short domed, glabrous, 0.10 cm. tall and base pair 0.09 cm. wide.

Top view of the calyx enlarged about 16X. Sepals are triangular 0.12 cm. long and 0.10 cm. at the widest, glabrous with small ligules present at the sinuses. Apex does not reach the corolla sinuses.

337

Hoya sp. Matafoao Ridge Yellow Collected October 2003 2/3 way up the hog-back ridge to the West, where the trail turns South to the Peak. A butter clear yellow flower cluster dangling from a vine from there, leaves fairly large, pinnately nerved.

View of calyx attached to the corolla enlarged about 8X. The calyx outer surface is glabrous, granulose. Sepals are considerably thickened in the center, overlap a little at the base. The apex is narrowly rounded, 3/4 way the corolla sinuses.

Side view of the pedicel, calyx and ovaries enlarged about 8X. All parts are glabrous. Ovaries are short dome shaped 0.12 cm. tall and 0.12 cm. wide at the base pair.

Top view of the calyx enlarged about 8X. Lobes are triangular, netted with nerves, inside shiny-slick, glabrous. There are tongue like ligules present. Apex - base Apex - center Widest

0.11 cm. 0.27 cm. 0.15 cm.

Tongue- like ligule at each sinus of the calyx, here greatly enlarged.

338

Hoya sp. Matoata Cream 2003 SW end of Tutuila, American Samoa 50m. In bloom early October 2003 vine just above roadside in bloom with 26 large cream colored flowers on long rachis, bracteated.

Outside view of the flower pedicel and calyx enlarged about 16X. The sepal apices reach 2/3 the way to the corolla, sinuses are broadly triangular. Outside finely granulose.

Side view of the pedicel, calyx and ovaries enlarged about 8X. Apex - base Apex - center Widest Ovaries base.

0.20 cm. 0.30 cm. 0.17 cm.

0.15 cm. tall and 0.13 cm wide at the

Top inside view of the calyx and ovaries. There is a slight overlap at the base and there are linear ligules in a pattern 2-2-1-1-1. Enlarged about 16X.

339

Hoya sp. Maximo Wayet Collected in the Diffin Area near Baguio, Luzon, Philippines. 1990.

Calyx side view. Pedicel is glabrous, terete . Sepals are 0.15 cm long and 0.13 cm. at the widest, granulose wrinkled outside, inside glabrous and shiny, a few cilia.

Calyx top view. Some sinuses have ligules but I did not find them at all places. Overlap of sepals is about ¼ - 1/3. Color rich yellow.

340

Hoya sp. Monina #1 Data 17 Dec. 2006, Fresno CA. Flower cluster and leaves sent via Monina Siar. Pollinarium indicates H. bilobata.

Pedicel and calyx enlarged about 18X. Pedicel: glabrous, terete, curved, 0.9 cm, long, longitudinally very fine sulcate. Calyx: membranaceous, entire, outside crystalline granulate, inside slick, glabrous. Sepals 0.10 cm. long and widest 0.05 cm., apex obtuse, ovate shape. Ovaries: cone shaped, glabrous, 0.08 cm. tall and 0.05 wide at base pair.

Outside view of a flower enlarged about 18X. surface glabrous finely granulate, sepals almost reach the corolla sinuses, pedicel enlarged near the calyx, corolla revolute. Sinus – sinus Sinus – center Sinus – apex Apex – center Widest

341

0.15 cm. 0.15 cm. 0.21 cm. 0.30 cm. 0.17 cm.

Hoya sp. Monina #2 Data & photos 18 Dec. 2006 3 flowers and 3 leaves from Dr. Monina Siar. Pollinarium indicates H. bilobata.

Side view of the pedicel, calyx and ovaries enlarged about 16X. Sepals with ragged edges, 0.07 cm. long and 0.06 cm at the widest, ligules are present, length of pedicels as with #1. all surfaces glabrous. Ovaries: here more done shaped, not cone shaped, 0.07 cm. tall and base pair 0.05 cm wide, glabrous.

342

Hoya sp. MT. 13 via May Tolentino, Manila. Philippines. 5 flowers in Solution from Torill Nyhuus 3/20/07

Side view of a calyx enlarged about 16X. Base of calyx bulbous. sepals ciliate, outer surface rough but glabrous.

Bottom view of a calyx, pedicel cut off enlarged about 8X. Sepals are centrally thickened, edges ciliate, semi linear to triangular. Sepals 0.30 cm. long and 0.19 cm. at the widest above the sinuses, overlapped only slightly at the base.

Top view of the calyx Sepals inside glabrous and slick, shiny, ligules 1-1-1-1-2 at sinuses. Apices narrowing obtuse. Ovaries columnar, glabrous, 0.13 cm. tall and base pair 0.10 cm.

343

Hoya sp. Nagtabon

Side view of the pedicel, calyx and ovaries enlarged about 8X. Pedicel: Strict, terete, 0.15 cm. long with globose cluster, fine dense short pubescence. Calyx: sepals short, triangular, do not reach the corolla sinuses, punctate. Ovaries: glabrous 0.12 cm. tall and base pair 0.11 cm. wide.

Top view inside the calyx.. Sepals 0.13 cm. long and 0.11 cm. at the widest, short ciliate, apex rounded, overlap about ¼ and ligules are present (very obscure, same color as the sepals.

Outside view of a flower with pedicel and calyx in place. Notice the pedicel surface.

344

Hoya sp. NS-0009 Collected by Nathalie Evelinadotter on Mindoro. Philippines Photographed 23 Feb. 2005

Calyx side view, outer surface with scattered hair cells (white) and granulate. The sepals reach less than ½ way to the corolla sinuses. Ovaries: bottle shaped, 0.30 cm. tall and the base pair 0.16 cm. wide, glabrous.

Top view of the calyx, edges are ciliate, inside surface glabrous and shiny. Sepals 0.30 cm from center to apex and 0.16 cm. long 0.18 cm at the widest, ligules are present at the slight basal overlap. Triangular in shape with apex rounded

345

Hoya sp. NH #1 Flowers from Ann Wayman, Photographed 30 March 1995. Role 193-194:

Outside of the corolla with pedicel and calyx. Pedicels glabrous, terete, with occasional bumps and a hair cell or two. Calyx small, sepals short and do not reach the corolla sinuses, 0.09 cm. long base 0.10 cm., triangular, glabrous both surfaces.

Sepals are triangular and small arising from a relatively large pedicel.

346

Hoya sp. NS05-206

Side view of pedicle, calyx and ovaries enlarged about 16X. The base of the calyx is rounded and surface granulate, glabrous as are the outer surfaces of the calyx. Ovaries: short domed 0.10 cm. tall and 0.10 cm. wide at base of the pair, glabrous.

Back of flower enlarged about 8X. The sepals reach or slightly exceed the corolla sinus, are triangular linear, with no overlap at the base, 0.15 cm. long , 0.10 cm. at base the widest part. The granular surface of the sepals is somewhat linear and reach the edges, which gives the appearance of cilia.

347

Hoya sp. NS05-213 Hoya halconensis Kloppenburg Flowers preserved in vial. This is the same species as NS05 225.

Outside surface of the calyx enlarged about 16X. Sepals are thickened triangular-linear, ciliate with a few cilia on the immediate surface, apex narrowly rounded, little base overlap.

Inside view of the calyx, surface is shiny glabrous, a little overlap at the base. Sepals: 0.30 cm. long, 0.19 cm. wide near the base, very small ligules present. Ovaries: columnar 0.24 cm. tall and base of pair 0.14 cm. wide.

Outside surface of the flower at the calyx area and corolla sinus enlarged about 8X. The sepals do not extend to the corolla sinus. Corolla is fleshy-thick. and the coronal outer lobe projects beyond the sinus. Sinus – sinus Sinus – center Sinus – apex Apex – center Widest

348

0.50 cm. 0.40 cm. 0.95 cm. 1.20 cm. 0.60 cm.

Hoya sp. NS05-231 Mt. Halcon, Philippines via Torill Nyhuus march 2007, 4 flowers in solution

Side view of the calyx enlarged about 8X. Pedicel enlarges a little below the bulbous base of the calyx. Calyx lobes outspread. edges ciliate, outside a few hair sells and center thickened apex obtuse.

Top view of the calyx and ovaries enlarged about 16X

Ovaries: columnar, apex nippled, glabrous. 0.25 cm. tall and base pair 0.16 cm. wide. Sepals: broadly triangular with apex obtuse, overlap ca. 1/3 0.23 cm. long and 0.23 at the widest. Inside slick, glabrous. Ligules present.

349

Hoya sp. PNG #4 Hoya lamingtoniae Bailey 1898 From flowers collected at Edward Gilding's, Pearl City, Hawaii 9/8/00 Top view of the calyx, for the most part the sepal edges are entire and both surfaces are glabrous. There is quite a bit of basal overlap, ligules are present and easily seen. Calyx: edges of sepals entire, 1/4 overlap, clear basal ligules almost to sinus. Diameter 0.23 cm., sepal length variable 0.100.20 cm. widest 0.13 cm.

Pedicel and pedicel with calyx. The pedicel (visible at left) had some outgrowths I have never observed before growing from the pedicel, The pedicel is glabrous Pedicels: strict, 2.5 cm. long, terete, glabrous, red color, 0.09 cm. in diameter. Ovaries: domed 0.18 cm. tall base width of pair 0.14 cm., glabrous and smooth.

350

Hoya sp. PNH 11859 Species found by G. E. Edano 12 March 1950 at Antilao River, Ormoc. Leyte, Philippines. Field No. 2018, along a ridge in forest, elevation 500 meters. Flower green, fruit green.

Pedicel, calyx and ovaries. Pedicel: ca. 1.3 – 2.0 cm. long , terete, filiform finely hirsute.

Pedicel and calyx, to show the dense hirsute surfaces. Sepals are in addition ciliate; inside glabrous. 0.1 cm. long base 0.05 cm.

351

Hoya foxii Kloppenburg Hoya sp. PNH 5023 Described in New Hoya Vol. 2-1, e published, rare-hoyas.com This species was labeled Hoya gracilis Schlechter ?, collected by Robert B. Fox at Dibutunan, Baler, Quezon Province, Luzon, Philippines march 1948. Determination made by E. D. Merrill. This species is not a Section Acanthostemma (Bl.) Kloppenburg species. It is not Hoya gracilis Schlechter. The notation on the herbarium sheet says "The leaves of this plant are crushed in a coconut shell of water, removed, and then the leaves are rubbed over the entire body of a person suffering from fever”. Photomicrographs from the herbarium sheet: Pedicel, calyx and ovaries. Pedicel is long and curved, terete, glabrous, filiform 1.7 cm. long and 0.035 cm. in diameter. Sepals have about 1/4 overlap at their base, edges membranous, apex rounded 0.12 cm. long and 0.08 cm. wide. Ovaries are bottle shaped 0.10 cm. tall base pair 0.08 cm. wide.

Calyx and ovaries top view.

352

Hoya sp. PNH 5733 Collected by Barbon, Garcia & Sagcal at Samar Brg. Inuntan, Basey, 29 Jan 1992 studied 5 Nov. 1993. Scales like H. pentaphlebia but leaf very small for that species.

Side view of a pedicel and calyx. enlarged about 16X. Pedicel is 4 + cm. long x 0.06 cm. in diameter, glabrous, terete enlarging toward calyx. 15 + flowers Calyx; small outside granulate inside slick, smooth, 0.09 cm. long x 0.08 at the base, edges with a few cilia. Ovaries cone shaped, glabrous 0.07 cm. tall and base pair 0.08 cm. wide.

Outside view of the corolla enlarged about 8X, surface is glabrous, lobe apices acute. Sepals very far from reaching the corolla sinuses.

353

Hoya sp. Western Samoa Double Corolla Cutting was collected by me in Western Samoa, Lake Lanatoo area in 1988, sent to Michael Myashiro who bloomed it and sent me flowers. It was photographed in 5/21/90. Michael could not remember the plant, but I assume it may still be in existence. Photomicrographs follow:

Top view of the calyx. (picture blurred). Calyx 0.50 cm in diameter. Sepals 0.17 cm. long 0.16 cm. wide. Ligules present in pattern 2-2-1-1-1,

Side view of the pedicel, calyx and ovaries enlarged. Pedicel is terete, glabrous, filiform, creamy yellow color 1.6 cm. long 0.10 cm in diameter. Calyx outside granulose with a few scattered hair cells.

354

Hoya sp. Sulit # 10147 M. D. Sulit (PNH) 10147 15 April 1949 Mt. Katanglad, Bukidnion Prov. Mindoro. Flower creamy 1800 m. forest edge (roll 144 drawing 136). I had determined this to be Hoya cardiophylla but here the corolla is densely pubescent inside which negates it being that species.

I failed to make measurements.

Pedicel, calyx and ovaries.

355

Hoya sp. T. G. revoluta Flowers sent by Ted Green, Kaaawa, Hawaii as H. revoluta Wight 21 June 1993. I do not believe it is that species since the cluster is too tight and pedicels to regular. Photomicrographs of this species follow: 16 flowers/ cluster.

Top view of the calyx. Sepals do not overlap, edge with a few cilia, edges membranaceous. 0.14 cm. long and 0.10 cm at base. Ligules present.

Side view of the pedicel, calyx and ovaries. Pedicel is terete, 0.10-1.40 cm. long, glabrous with blotched and linear red markings, otherwise bufforange, enlarging toward the calyx base. Ovaries: bottle shaped, 0.15 cm. tall and base pair 0.11 cm.

356

Hoya sp. USDA 354238 Photomicrographs follow; Flowers supplied by Ann Wayman, Central Point, Oregon.

Side view (left) and top view (right) of the calyx. The calyx with same outside surface but inside smooth. Sepals with 1/3+ overlap (0.12 cm.) length 0.27 cm. an occasional cilia.

The pedicel is straight 1.7 cm. long 0.11 cm. in diameter, very fine glistening texture and very densely punctate (protopubescent). Ovaries are domed 0.20 cm. tall and base pair 0.15 cm. glabrous. They are well short of the corolla sinuses.

357

Hoya sp. camphorifolia similis This plant bloomed in October 2000 at the end of the season. It has small flowers that stay open only 1-2 days or so. This species is evidently not H. camphorifolia Warburg, as for one thing the outer coronal lobes here are acute (not obtuse). In addition the venation on the foliage is different and the veins are not distinct, also the color on the top side is glossy green and deeper color green than the dull lower surface. Photos taken 10/20/00 from flowers at Fresno, CA.

Top view of the calyx. Diameter is 0.15 cm; sepals are 0.11 cm. long, widest 0.10 cm., outside granulate, inside glabrous smooth. Ovaries are tapered domed 0.10 cm. tall and the base of the pair is 0.08 cm. wide.

358