Leaf Data from Hoya Type descriptions

Hoya acicularis Green & Kloppenburg 1992. semi-terete leaves (not linear-lanceolate……….. bifoliate at the node, that may be straight to curved and twisted; petiole curved. Hoya acuminata Bentham ex Hooker 1866. leaves opposite, oblong, acuminate, fleshy. Hoya acuta Haworth 1821. leaves ovate lanceolate acuminate. Hoya aeschynanthoides Schlechter. densely leaved; with leaves shortly petioled, broad ovate-elliptic, acute, pilose (felt like) on both sides, 1.5-2.3 cm. long, nearly 1-1.8 cm. wide, petiole hispid 0.3-0.5 cm. long. Hoya africana Decaisne 1844. leaves ovate or ovate-oblong acuminate, bases rounded or somewhat attenuate, glabrous on both sides, midrib above at the limb base glandular. Hoya alagensis (Shltr.) Kloppenburg 1990 leaves outspread ovate-lanceolate, with the base rounded, apex acute and beaked, 11 cm long by 4.5-5 cm. wide, both surfaces glabrous, not fleshy, veins on both sides 4-6 ascending, reticulate, costa (midrib) thick prominently raised below. Hoya alata K. Hill 1986. Leaves fleshy to coriaceous, ovate or rhomboid, acute, tapering to a narrowly chordate base, pale green, pink or pale bronze when growing in strong light. 3-7 cm long, 2.5-5.0 cm wide; margins and apex recurved; venation obscure, sub-parallel or somewhat palmate, petioles 3-8 mm long, 2-4 mm diam. Hoya alba Kosteletsky 1834. None Hoya albiflora Zippelius ex Blume 1848. leaves elliptic oblong cuspidate…….. Leaf 4.5 - 8 inches (long), 1 and 2/3 -2.5 inches wide, elliptic-oblong or oblong, with a constricted pointed apex, base somewhat rounded, texture very fleshy, with the lateral veins for the most part not visible, the a strong rounded prominent middle nerve on the back, above channeled and at the base warty. Hoya aldrichii Hemsley 1890. leaves with 5 nerves.

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Hoya alexicaca Moon ex Trimen 1895. glabrous; l. 1.5 - 3.5 in., ovate or oblong-ovate, usually cordate at base, with rounded lobes, very acute at apex, glabrous, thin, petiole long, slender. Hoya amboinensis Warburg 1907. leaves oblong and broad with a rounded base, acute apex 6 - 7 cm. long 2.5 - 5 cm. wide, above clear, at the base of the costa somewhat granulose, veins about 4 outspread, curving and curving and running together prior to reaching the margines , not at all distinct, reticulations barely thickened. Hoya amoena Bakhuizen 1950. leaves oblong-lanceolate, sometimes almost rhomboid or ovate with the base acute or wedge-shaped, rarely obtuse, acute or briefly acute acuminate, when dried thin but strongly leathery somewhat uninerved or distinctly trinerved at the base, lateral nerves and transverse nerves above fixed at the base which are barely different, glabrous 8 - 11 cm. long, 3.5 - 4.4 cm. wide, petiole thin, glabrous 8 - 15 mm. long. Hoya andalensis Kloppenburg 2005. Leaves opposite small, cupped below, ovate 5.0 x 2.2 cm. mostly uniform, medium flat green color above, below much lighter, most leaf pairs turned to one side of the stem. Hoya anncajanoae Kloppenburg & Siar 2008. Leaves: lanceolate- elliptic 5- 9 cm. x 1.5 – 2.8 cm. thick and fleshy, base subcuneate apex sub-acute. new growth with more pubescent than older foliage, juvenile lower surface with fine pubescence to finely papilate, edges rounded, upper surface with fewer but more uniform distribution. Mature foliage some hair cells below, upper with fine pustulate. Nerves obsolete but pinnate, 10 or so nerves on each side of midrib anastomosing. Hoya angustifolia Traill 1830. Leaves narrowly-lanceolate, Narrow leaved Hoya. Leaves membranaceous. Hoya angustisepala Schlechter 1830 Hoya angustisepala Burton 1987. leaves very leathery, glabrous, oblong nearly oblong-ovate, as long as 15 cm., 6 cm. wide, base broadly rotund or obscurely emarginate, apex acute nearly subacuminate, nerves on both sides 3-5, nerves thin; petiole thick nearly 2 cm. long, glabrous deeply grooved. Hoya anulata Schlechter 1905. loosely leaved; leaves flat outspread elliptic or oblong elliptic, ovate, sometimes obtuse acuminate, both sides glabrous, nerves barely conspicuous, of thick leathery texture, 6-8 cm. long, near the middle 4-4.5 cm. wide. Hoya apiculata Scheffler 1876.

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Leaves glabrous at the base almost cordate, somewhat orbicular, tips apiculate, one nerved, with gland inserted at the base. Hoya apoensis Kloppenburg & Siar 2010. Hoya archboldiana C. Norman 1937. Leaves thick, large, pinnately veined, up to around 13 x 7 cm., oblong, tips abruptly acute, when dry the base lightly cordate with the sinus closed. Hoya arnottiana Wight 1834. leaves fleshy, broadly ovate acuminate leaves fleshy, broadly ovate acuminate. Hoya attenuata Christophersen 1935. blades 5 x 1.6 - 6.5 x 1.9 - 8 x 2.8 cm. long and broad; petiole grooved, glabrous, 8 - 11 mm. long. Hoya aurantiaca Kloppenburg & Siar 2009. Foliage oblanceolate, apex shortly acute, base narrowing subcordate, surface dull green with occasional dull purplish staining. Nerves quintuplinerved of a lighter color distinct above to very distinct even raised, invisible below, first pair arise slightly above base, ending in looping secondary nerves below middle, second pair above base, looping with side reticulations most not reaching the apex. 7.5-13 cm. long and 3.0-5.0 cm. wide Petioles extremely variable, most thick, curved as large as stem or larger, some extremely long. Hoya australis R. Br. ex Trail 1830. Leaves on short petioles, ovate obovate or nearly orbicular, obtuse or shortly acuminate, rounded or rarely almost cordate at the base, thick and fleshy, 2 to 3 in. long or rarely more. Hoya australis subsp. nathaleae Kloppenburg & Siar 2009. Leaves ovate, cordate, short apiculate apex, 6.5 cm. long 3.6 cm. at the widest, dorsal surface stiff short pubescence, ventral side more dense indumentum. Peduncle densely pubescent, 4 triangular glands at the attachment point above, terete, curved 1 cm. long 0.10 cm. in diameter. Hoya balansae Constantine 1912. Leaves fleshy glabrous, grayish-green, at times with spots of dark green , oval rounded at the base, tip acuminate, nerves not very apparent 3; lateral nerves alternating, 2 to 4 on each side, meeting at the edge with the others; length 4.5 to 6.5 cm. by 2.5 to 3.5 cm.; petiole thick, to 5 mm. long. Hoya bandaensis Schlechter 1908. loosely leaved; leaves petiolate elliptic acute or briefly acuminate, both sides glabrous, with fleshy texture, 10-12 cm. long. in the middle 5-7 cm. wide, petiole fleshy about 1.5 cm. long.

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Hoya barbata Sprengel 1825. leaves ovate acute very glabrous.

Hoya barracki Horne ex Backer. None Hoya bella Hooker 1849. with copious foliage, leaves small ovate acute shortly petioled one nerved above intense green below pale, lateral. Hoya benguetensis Schlechter 1906. loosely leaved, with the leaves elliptical or ovate elliptical, acuminate, glabrous, with thick coriaceous texture, with 5 primary nerves very conspicuous, 6-10 cm. long in the middle or below 2.5 – 4 cm. wide. Hoya benvergarai Kloppenburg & Siar 2008. Blades 9-13 cm. long x 2-3 cm. at the widest. Petiole round 1.5-2.0 cm. long. Hoya betchei (Schlechter) Whistler 1908. loosely leaved, leaves outspread petiolate lanceolate-elliptic, with the base rounded, glabrous, texture leathery, 6.9-9.5 cm. long, in the middle 1.8-2.3 cm., petiole fleshy about 1 cm. long; cymes slenderly pedunculate shaped like umbels, of few flowers, peduncle round, glabrous, 5-6 cm. long. Hoya bhutanica Grierson & Long 1979. Leaves elliptic and not fleshy, corolla at least twice wider inside. Creeping shrub stem narrow 3-5 meters, rooting. Leaves elliptic (6.5) 9-15 x (2.75) 3-4.5 cm., apex acute base rounded, trinerved, lateral veins produced ½ above, slightly leathery, glabrous, margined entire. Hoya bicarinata A. Gray 1861. leaves glabrous somewhat fleshy flat obscurely penninerved oval obovate or ovate very briefly and abruptly acuminate bases rounded almost cordate, blade above petiole hairy glandular. Hoya bicknellii Kloppenburg 1999. Leaf: Opposite and whorled. Petiole terete, curved, 1.5 cm. long, nearly the same diameter of the stem from which it arises, glabrous, green. Blade broadly ovate to ovate elliptic, flat to sub curved with some undulations. 9.0—12 cm. long x 0.6—0.8 cm. wide in the center. Base cuneate, edges cupped upward slightly, pinnate anastomosing veining, visible above as is the midrib but obscure below. Apex shortly apiculate to slightly acute. Glossy green above duller below, glabrous.

Hoya bicolensis Kloppenburg & Siar 2009. Foliage ovate to lanceolate apiculate 7.8 – 12.5 cm. long and 2.0 – 4.0 cm. widest, triplinerved, prominent above obscure or less below. A basal pair plus midrib and a pair above, looping and anastomosing. Dull to dark green above paler below. 4

Petiole round curved to twisted 1.5-2.4 cm. long, larger in diameter than the stem 0.250.30 cm. in diameter. Hoya bicolor Kloppenburg 2003. Foliage opposite, blades ovate with a cordate base and extremely short acute apiculate apex, slightly furrowed at the midrib on the dorsal surface, otherwise horizontal, edges entire, pinnately at about a 40 degree angle, ca. 4 pairs on either side, looping in before reaching the margins and somewhat anastomosing. Lower surface along the midrib with scattered hair cells. Petiole long twister ca. 3 cm. long 0.4 cm. in diameter, slightly grooved near the blade otherwise terete; two small semicircular raised dark glands at attachment above. Internodes variable in length 5 cm.- 12.5 cm., puberulous, with nodes enlarged, leaves leaving circular scars. Hoya billardieri Decaisne 1844. leaves oval by no means or rarely bases emarginate or somewhat cordate apexes very shortly acuminate with both sides glabrous, limb above petiole glandular, with three 3 obtuse glands. Hoya bilobata Schlechter 1906. well leaved; leaves briefly petioled broadly elliptical or somewhat orbicular, obtuse, glabrous, leathery textured, 1.7-2.2 cm. long, 1.3-1.8 cm. long in the middle, petiole about 0.3 cm. long. Hoya blashernaezii Kloppenburg 1999. Leaf: petiole terete curved .7 - 1.2 cm. long 2.0 - 2.5 cm. At the widest, blade elongate elliptic- lanceolate with a slight curve inward apex tapering to a very acute apex, base cuneate or occasionally slightly rounded; venation prominently triplinerved; deep green and glossy above, dull and lighter green below, midrib obscure. Hoya blumeana Schlechter 1913. None Hoya bonii Costantin 1912. Leaves fleshy, opposite, round or heart shaped at the base end with the apex pointed, occasionally oval or even oblong and pointed at apex. Some hairs below, short and with intervals between the hairs visible, often seen with magnifying glass only; hairless and smooth above. Midrib very projecting below and hairy along its sides; secondary nerves 5 to 6 pairs, slanting at 45º for some distance towards the edge. Leaf 8 to 10 cm. long. by 3.8 to 7 cm. wide; petiole to 1 cm., 4mm. thick. Hoya bordenii Schlechter 1906. loosely leaved; with leaves outspread lanceolate-oblong or narrowly elliptical, acuminate, glabrous, texture leathery, 11-18 cm. long, 2.5-4.5 cm. wide at the middle or below the middle, with the petiole short, fleshy 1.5-2 cm. long. Hoya brevialata Kleijn & Donkelaar 1999.

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Leaves suborbicular to elliptic, light-green, yellowish to bright red when very exposed. Petioles 0.5--1 cm long. Hoya brittonii Kloppenburg, 1992. The leaves are noted to be thick and on drying are pinnately veined, 3-4 looping nerves on either side of a somewhat obscure midrib. They appear also to be reticulately nerved………… Leaf Blade: variable but generally obovate 8 cm. long, 3 cm. wide above the middle, thick, glabrous on both surfaces, with a cuneate base leading into the petiole; pinnately nerved, probably obsolete in the living state. Hoya brooksii Ridley 1925. leaves fleshy, elliptic, somewhat acute, bases cuneate, nerves invisible, midrib even obscure, 5 cm. long, 2 cm. wide, petioles 5 mm. long; peduncles somewhat robust 3 cm. long. Hoya browniana Koorders 1911. leaves opposite (but not always, rarely one aborts becoming pseudo-alternate), oblong, glabrous, fleshy, dried led-gray, with the base acute, with the apex acuteacuminate, distinctly triple-nerved, 14-20 cm. long and 6 - 7½ cm. wide. Petioles 2 - 2½ cm. long, thickened, glabrous. Hoya bruensis Miquel 1896. Leaves shortly petioled with rounded bases, slightly emarginate, above the petiole minutely callused, (a little) ovate-elliptic or (more or less) elliptic-oblong with the tip acute or obtuse-attenuate, fleshy, glabrous, below the midrib covered over, on both sides about 5 (from the very base) erect and outspread curved almost united. Hoya brunioniana Wight 1834. leaves fleshy, oval-oblong acuminate. Hoya bulusanensis Elmer 1938. See under Hoya panchoi Kloppenburg. . Leaves very thick, pale or yellowish green beneath, the upper surface paler green, normally opposite along the slender branches, more or less clustered toward the base or upon very short and thick seemingly abnormal branchlets, subdeciduous, glabrous but minutely punctate on both faces, the dry blades wrinkled on the upper but perfectly smooth on the lower side, flat, edges straight and entire, curing yellowish brown, of two distinct types, the orbicular to rotund shape and the elliptic to the obovately oblong type, the latter ones 2 cm wide above the middle and 4 cm in length, obtusely ro6nded at the apex, broadly cuneate at the base, the smaller kind 1.5 cm wide across the middle; petiole very short and very thick, curved and expanded toward and at the base, dull brown, rugose on my specimen, leaving large circular scars after falling; midrib scarcely evident or entirely obsolete in the smaller type of leaves, in the middle; petiole very short and very thick, curved and expanded toward and at the base, dull brown, rugose on my specimen, leaving large circular scars after falling; midrib scarcely evident or entirely obsolete in the smaller type of leaves, in the middle region longitudinally striate below especially so of the larger blades, lateral nerves none.

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Hoya buotii Kloppenburg 2002. The typical leaf is elliptic-lanceolate with a long acuminate apex, base narrowly rounded. Blade is 10 cm. long x 2.5 cm. wide near the base. The petiole is grooved above, 1 cm. long x 0.09 cm. in diameter. Nerves are pinnate 7 or so on each side, at from a 30º to nearly perpendicular angle to the midrib; anastomosing. Hoya burmanica Rolfe 1920. Leaves somewhat pendulous, shortly petiolate, narrowly ovate-lanceolate, acuminate, somewhat concave, slightly thick, glabrous, glaucous-green 5 to 8 cm. long, 1.3 to 2 cm. wide; petiole 5 mm. long. Hoya burtoniae Kloppenburg 1990. The leaves of this species are more regularly ovate-elliptic, rather than long and narrowly lanceolate. Hoya burtoniae contains a lot of anthocyanin pigmentation, which is visible throughout the plant, especially manifest on the lower leaf surfaces……… Leaves beautifully colored, opposite, far apart, thick or leathery, with revolute margins, narrowly oblong or oblanceolate, rigid, above covered with closely spaced velvety granules, beneath less pubescent and granular, 2.5 to 12 cm long and above middle 3 cm broad, bases attenuate, at apexes obtuse or somewhat acute; midrib not prominent, side nerves not at all prominent; petiole velvety 0.8 to 2.5 cm long. Hoya cagayanensis Burton 1987. Leaves elliptic, lightly leathery, penninerved, both sides glabrous, 6 - 8 cm. by 2.25 - 3 cm., petiole 1 com (cm.) plus or minus long. Hoya callistophylla Green. leaves opposite, elliptic to lanceolate with attenuate tips, cuneate bases, prominent, pinnate, dark green venation, substance heavy and hard, with a matte surface. Hoya calycina Schlechter 1908. Leaves spread or outspread, petiolate, broadly oval, apiculate, with base somewhat cordate, leathery, above sparsely puberulous, below shortly tomentose, petiole tomentose above slightly grooved. Hoya campanulata Blume 1826. leaves leathery almost veinless oblong, acuminate glabrous. Hoya camphorifolia Warburg 1904. Leaf blade glabrous on both sides, parchment like, ovate elliptic or elliptic, 6 to 9 cm. long, in the middle 2.5 to 3.5 cm. wide, apex acuminate acute often curled back, often convex on underside, bases narrowly rounded. Veins 2-3 on both surfaces ascending, not all reaching the apex, reticulate, distinct but not prominent. Peduncles long and filamentous, straight, glabrous, 8-9 cm. long, 0.1 cm. diam. Hoya cardiophylla Merrill 1920.

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. Leaves heart-shaped, apparently fleshy when fresh, rather flaccid when dry. 5 to 7 cm. long. 4.5 to 5 cm wide, pale when dry, glabrous or the younger ones slightly ciliate near the basal margins, the base broadly rounded and distinctly cordate, the sinus acute, the basal lobes broadly rounded, the apex shortly and sharply acuminate; petioles 5 to 7 mm long; lateral nerves about 4 on each side of the midrib, somewhat spreading, anastomosing, the reticulations lax and distinct on both surfaces. Hoya carnosa R. Brown 1802. Leaves oval-oblong fleshy, corolla hair. Hoya caudata J. D. Hooker 1883. Leaves 2-3 in. diam., sparsely papilose on both surfaces, midrib and arching nerves faint: petiole ¼ in., so thick as to appear globose. Hoya cavaleriei Leveille 1914. Leaves leathery, nerves conspicuous, middle nerve elevated. Hoya cebuensis Kloppenburg & Siar 2010. leaves are ovate lanceolate with an obtuse base versus linear-elliptic with a cuneate leaf base. Hoya cembra Kloppenburg 1990. leaves erect- spreading briefly petiolate, narrow elliptical beaked, glabrous, shinny, somewhat leathery, 8 - 10 cm. long, below the middle 2 - 3 cm. wide, petioles 0.5 cm. long. Hoya chinensis Traill 1830. None Hoya chinghungensis (Tsiang & P. T. Li) M. G. Gilbert, P. T. Li & W. D. Stevens. leaf blade broadly ovate, 1- 1.5 x 0.7 – 1.1 cm, glabrous, base rounded to truncate, apex acute to obtuse; lateral veins obsolete. Hoya chlorantha Rechinger 1908. Leaves leathery, when dry shiny green, nerves not prominent on either side, briefly petiolate, narrowly acuminate at both ends, penninerved. Hoya chlorantha var. tutuilensis Christophersen 1908. Leaves leathery, when dry shiny green, nerves not prominent on either side, briefly petiolate, narrowly acuminate at both ends, penninerved. Hoya chloroleuca Schlechter 1913. Leaves spreading or outspread, petiolate, lanceolate or elliptic- lanceolate, acute or sub acute, thinly coriaceous, both sides glabrous, petioles short, glabrous, above slightly grooved.

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Hoya chunii P. T. Li. 1984. None Hoya cinnamomifolia Hooker 1848. leaves fleshy-leathery, thick ovate or ovate oblong acuminate, 5 nerved, nerves above prominent, above at the petiole callused glandular; peduncle short. Hoya citrina Ridley 1922. Leaves thick, fleshy stiffly coriaceous ovate, base broad, round, subcordate; nerves three conspicuous when dry with about 4 pairs of lateral nerves from the central nerve broken up into reticulations large and lax. 4 to 4.5 in. long, 3 in wide; petiole very thick .75 in long. Hoya clandestina Blume 1848. not from type description (none) leaves fleshy elliptic-oblong acuminate with the base rounded, three nerved between the nerves obscure veined; Umbels with many flowered long peduncles’ the inside of the corolla silky lobes ovate acute revolute. Hoya clemensiorum Green. leaves opposite, thick and rigid, petiole 7 mm x 2 cm, curved, blade 10-20 cm wide x 20-50 cm long, with raised, prominent, pinnate venation, ovate-lanceolate with attenuate tip. Hoya cochinchinensis Loureiro 1817. leaves opposite, ovate. Hoya collettii Schlechter 1913. Leaves petiolate fleshy, linear-oblong or linear-lanceolate 3-4½ inches long, somewhat obtuse, base elongated, rounded, above shining, above paler midrib thick elevated; petioles thick 3-4 lines long. Hoya collina Schlechter 1913. Leaves erect spreading or widely outspread, shortly petioled, elliptic or ovateelliptic, obtuse, wedge shaped base, fleshy, both sides glabrous, petiole fleshy, above lightly furrowed (grooved). Hoya cominsii Hemsley 1890. Leaves fleshy, glabrous, wide ovate, with petiole thick, at least 5 inches long, obtuse or somewhat acute, palmately 5 to 7 nerved. Hoya compacta Burton Caulis puberulis, brunnei, albis pilis, nodis brevis 1 1/2 - 2 cm.. Folia 5 1/2 - 6 cm. longa X 3 cm. lata, obsolete plicata, inferiora exposita, superiora persipe occulta complanata, (basi cordatis, apices acuminatis), ocula nuda vel negligentius glaberrima

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sed reversis pilis minutis dispersis, ocula armata aspectabilibus utrinque conspersis. Petioli ± 1 cm. longi, supra non canaliculat. Hoya coriacea Blume 1825. leaves a little nerved oval acute or acuminate coriaceous, glabrous. Hoya costantinii P. T. Li 1984. Hoya reticulata Cost. in Lec., None Hoya cordata Li & Huang. caule terete cortice cineraceo-flavo 5 mm diametro. Folia opposita succulenta vel exsiccations crasse coriacea, ovate vel long ovate 5—9 cm longa 4.5—5 cm lata, apice acute vel obtuse basi cordata, supra glabra et subtus ad costam mediam puberula, nervis lateralibus utrinsecus 6— paribus utrinque leviter elevatis, petiolis robustis 5 mm longis pubescentibus, apice glandulis perpaucis 3—5-fascicularibus instructis. Hoya crassicaulis Elmer ex Kloppenburg 1995. leaves leathery elliptic or oblong-acute, shiny, 8-17 cm. more or less large, in the center nearly 5-8 cm. wide, base rounded or a little somewhat acuminate and recurved, margins somewhat revolute, midrib prominent below, veins on both sides about 4, spreading curving confluent to the margins. Hoya crassifolia Haworth ex Traill 1811. The leaves are obovate, very bluntly acuminate, upwards of four inches long, without veins, of a darkish green colour above, pale beneath. The petioles are thick and fleshy, of a purplish green colour, and much flattened above. Hoya crassior Hochreutiner 1935. Leaves opposite, midsize, petioles short, glabrous, more or less fleshy, channeled above and where united with the blade somewhat glandular; blade very glabrous, very thick and fleshy, the midrib it seems depressed on but sides, otherwise lanceolate, apex a little attenuate-acuminate, base acute, margines entire, secondary nerves inconspicuous, or in juvenile leaflet blade about 3, both sides arching and meeting and with reticulate nerves. Hoya crassipes Turczaninow 1848. leaves petiolate elliptic or obovate, the base attenuate, briefly acuminate, fleshy veinless glabrous. Hoya cumingiana Decaisne 1844. leaves ovate-cordate obtuse somewhat fleshy conspicuously veined papilose velvety with short petioles.

above

glabrous

below

Hoya curtisii King & Gamble 1908. Leaves sessile, very thick; orbicular, abruptly triangular-acuminate at apex, rounded or sometimes orbicular very slightly attenuate at the base; both surfaces

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scarious-puncticulate when dry; margin thickened; .4 to .7 in. long, .4 to .6 in broad; midrib when dry faint, other nerves invisible; petiole none. Hoya cystiantha Schlechter 1913. Leaves at bases generally acute, obverse oblong or lanceolate acuminate, leathery veined, 3-4 ½ inches long. Hoya dalrympleana F. Muell. 1861. = H. australis R. Br. ex Trail. None Hoya darwinii Loher 1910. Normal leaves elliptic oblong, 15-20 cm. long, 5-6 cm. wide, glabrous, leathery, with inconspicuous nerves, briefly petiolate; other leaves bullate, a globose pouch of 4-6 cavities, well closed. Hoya dasyantha Tsiang 1936. Leaves chartaceous, broad ovate, 4-7.5 cm. Long, 3-4.8 cm. wide, shortly acuminate, rounded at the base; nerves slightly elevated on both surfaces, lateral nerves 4-5 pairs, veinlets transverse, conspicuous; glands 4 to 6 in a group above the petiole; petiole 5 mm. long. Hoya davidcummingii Kloppenburg 1995. Leaf Blade: spreading lanceolate-elliptic, upper surface deep green even olivaceous, glabrous with fine granulation’s, edges rounded and slightly thickened, lighter green below: apex mostly acute turning down, base cuneate, overall thickened and coriaceous, veins not visible but pinnate, 5-7.5 cm. long, widest about the middle 2-2.5 cm. Hoya dennisii P. I. Forster et D. J. Liddle 1993. Leaves petiolate; lamina lanceolate-ovate, coriaceous, up to 72 mm long and 30 mm wide, discolorous, glabrous, venation obscure; upper surface dark green lower surface pale green; tip acuminate; base rounded to cordate petiole 5-7 mm long and c. 0.8 mm diameter, with scattered trichomes colleters 2, at lamina base. Hoya densifolia Turczaninow 1848. leaves somewhat sessile membranaceous densely imbricate elliptic, with short cordate sinus, slightly acute penninerved reticulate glabrous. Hoya deykeae Green 1990. Leaf blade: 5 - 12cm long x 5 - 8cm wide x 2 - 2.5mm thick, matte surface. Petiole 3 - 6 mm diameter x 10 -16 mm long, rough surfaced, tan. Hoya dickasoniana P. T. Li. leaves opposite even 3 or 4, elliptic or ovate, 9 – 19 mm. x 5 – 10 mm, base broad cuneate or rounded, apex obtuse, glabrous; petiole 1 mm. long.

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Hoya dictyoneura K. Schumann 1905. leaves briefly petiolate oblong or elliptic or somewhat orbicular acute with the base rounded five nerved both sides glabrous. Hoya dimorpha F. M. Bailey 1898. Leaves thick-coriaceous, oblong-acuminate, slightly cordate at the base; those on the thick portion of the stem exceeding 5 in. in length and 2 in. in breadth; those on the slender ends often cordate and under 2 in. long, glabrous, undersides pale, the gland on the face indicating the attachment of the petiole, beneath often a broad ciliate disk at times mote or less umbonate. Petiole stout, usually less then ½ in. long. Hoya diptera Seemann 1861. leaves ovate-elliptic, acuminate with the base ovate, fleshy, penninerved. Hoya dischorensis Schlechter 1913. Leaves spreading or outspread, shortly petioled, oblong, somewhat obtuse, acuminate, base obtuse, a little fleshy, both sides glabrous, petiole short, slightly grooved above. Hoya diversifolia Blume 1826. Leaves veinless oval acute orbicular fleshy glabrous. Hoya dodecatheiflora Fosberg 1940. leaves elliptic-lanceolate, thick, unequal, up to 8 cm. long, umbels axillary, many flowered, pedicles slender, 1 cm. long. Hoya dolichosparte Schlechter 1916. loosely leaved, glabrous with numerous wart like lenticels; leaves outspread, petiolate, broadly oval, obtuse, with 5 prominent primary nerves above, both sides glabrous 9-13.5 cm. long, almost 6.5-9 cm. wide at the middle, petiole fleshy 1.5-2 cm. long. Hoya edanoi C. M. Burton 1990. Leaves thick leathery, round or oblong, cordate or nearly (no such Latin word) peltate ? (shield shaped), with the tip round or mucronate 3-7 cm. long x 3-5 cm. wide. Hoya edenii King ex Hooker 1883. Leaves 3-4 by 1- 1 ¾ in., elliptic or obovate-lanceolate, obtusely caudateacuminate, base acute, thinly coriaceous, puberulous or glabrate beneath; petiole 1/12 in. pubescent. Hoya eitapensis Schlechter 1913. Leaves erect, spreading, elliptic, somewhat obtuse, with wedge shaped base, fleshy, glabrous with very short petioles, almost sessile. Hoya elegans Kosteletsky 1917.

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None Hoya elliptica Hooker f. 1883. leaves 1 ½ - 2 ½ in. elliptic obtuse et both ends thin in texture nerves very prominent, peduncle very short. Hoya elmeri Merrill 1929. leaves opposite, fleshy, when dry leathery, penninerved, somewhat elliptic, 11 to 15 cm long, 5 to 6.5 cm wide, when dry pale, both sides the same color, sparsely covered with small wort-like growths, both ends equally narrow, base acute, apex acute to briefly and obscurely acuminate, primary nerves on both sides of the costa 7, slender, distinct or sometimes somewhat indistinct, curved anastomosing, loosely reticulated; petiole thick, about 1 cm long; fleshy, glabrous 1.3 mm. long. Hoya elmeri Schlechter # 8896 (B). Not published Hoya enduensis Kiew 1989. Foliis glabris ovatis-cordatis apicibus acutis 2 ad 2.5 cm longis et 2 ad 2.5 cm latis a congeneribus. Hoya engleriana Hosseus 1907. leaves short or shortly petiolate, shortly lanceolate or narrowly ovate-lanceolate, margins curled inward leaves of the species accordingly at first appearing chiefly turned in the direction of the inflorescence, fleshy, white pubescent, nerves above inconspicuously marked, 1.5 cm long, 0.4 cm wide, narrow mobile. Hoya epedunculata Schlechter 1913. well leaved, round, glabrous. Leaves spreading or outspread, petiolate, elliptic, acuminate, somewhat leathery, both sides glabrous, with the peduncle glabrous, moderately thick slightly grooved above. Hoya erythrina Rintz 1978. Leaves coriaceous-fleshy, oblong with strongly corrugate margins and shallowly cordate basis; up to 15 cm long by 5 cm wide; 5-7 veined with a pair of veins parallel to the midrib glossy green or dull red above, red below. Hoya erythrostemma Kerr 1939. Leaves elliptic, base acute, apex mildly acuminate, up to 10 cm. long, 4 cm. wide, when dry papery, glabrous, midrib with the lateral and transverse nerves on both sides especially above, lateral nerves 5-6 pairs at 60° from the midrib joining the marginal nerves 3-6 mm. from the margins; petioles moderately robust, glabrous, 3-6 mm. long, above narrowly wedge shaped. Hoya esculenta Tsiang 1936.

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(H. liangii) have penninerved leaves; the leaves are of the same shape but much smaller. Hoya esquirolii Leveille 1912. None Hoya excavata Teijsmann & Binnendijk 1863. leaves somewhat fleshy, oval-oblong, base rounded, apex briefly apiculate…..Leaves opposite 0.15 long 0.09 wide, ovate-oblong, base rounded, or obtuse, apexes rounded, briefly aciculate, somewhat fleshy, above shinny, both sides with 6 nerves, with the petiole callous-glandulous. Hoya exilis Schlechter 1913. Leaves spreading or outspread, elliptic-lanceolate, obtuse, long-acuminate, texture for the genus thin, both sides glabrous, petiole glabrous, above slightly grooved. Hoya fetuana Kloppenburg 2003. Leaves short elliptic attenuate 5.3 - 5.5 cm. long, thin but possibly a little fleshy, apex subacute, base rounded 1.2 - 2 cm. at the widest near the middle; midrib narrow, visible below, pinnate netted nerves all the way to the apex. Petioles fine 0.5 cm. long, grooved above. Hoya filiformis Rechinger 1834. Leaves thin visibly reticulate veined, very glabrous bright……… Leaf texture thin, leaf exclusive of the petiole 6 cm long, 2 cm wide, 1 cm near the tapered point turns down. Petioles 8 mm long. Leaves are delicately feather nerved, nevertheless the nerves are distinct on the upper surface and visible on the lower surface, first of all from the median of the leaf bases extend outward in pairs almost to the edge of the leaf. …… Noteworthy through the long twisted apex of the leaf blade and the fine thin texture. Hoya finlaysonii Wight 1834. leaves oblong-lanceolate obtuse acuminate above (dried) reticulate, below somewhat veinless (5-7 inches long 1 ½ -2 wide) petioles short. Hoya fischeriana Warburg 1904. Leaves thick leathery broadly lanceolate 12 - 19 cm. long 1 - 7 cm. wide base and apex acute glabrous 5-nerved, nerves inside almost reaching the apexes, nerve reticulations distinct, petiole 2 - 3.5 cm. long 2 - 3 mm. thick; peduncles about 5 cm. long 2 cm. thick, upper half cylindrical 3 mm. thick. Hoya fitchii Kloppenburg & Siar 2008. Foliage is quintuplinerved, with the second pair of nerves nearly reaching the apex, outer connecting nerves anastomosing Nervation white or lighter green than the leaf blade color. Hoya flagellata Kerr 1940.

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Leaves ovate or lanceolate, base rounded and minutely eared, with the apex acute, margine more or less revolute, 4.7 - 8.5 cm. long, 2 - 2.5 cm. wide, leathery, when dry pale brown, glabrous except that above it is sometimes provided with minute sparse hairs, midrib scarcely visible above, lateral nerves obscure on both sides; petioles thick, wrinkled, glabrous, 3-5 mm. long. Hoya flavescens Schlechter 1913. loosely leaved, round, glabrous. Leaves spreading or outspread, oblong or lanceolate-oblong acuminate, base somewhat cuneate rounded, fleshy, glabrous, petiole short, somewhat fleshy above lightly grooved. Hoya flavida P.I. Forster et D. J. Liddle 1993. . Leaves glabrous, petiolate; lamina elliptic-ovate, up to 140 mm long and 45 rum wide, discolorous, secondary veins 4 per side of midrib just visible on both surfaces, tertiary venation obscure; upper surface dark green; lower surface pale green; tip acute to short acuminate; base cuneate; petiole grooved along top, 10 - 12 mm long and c.1.5 mm diameter. Hoya flexuosa Spreng. 1825. leaves cordate oblong veined. Hoya forbesii King & Gamble 1903. Leaves coriaceous when dry; elliptic-oblong or obovate, shortly and bluntly acuminate at apex, rounded at bass; upper surface glabrous, shining; lower surface minutely papilose; both pale brown when dry; margins somewhat recurved; 4 to 7 in. long, 2.5 to 4 in. broad; midrib strong, much raised beneath, bearing on the upper surface a gland just above the petiole; main nerves about 7 pairs, irregular but usually at 40° with the midrib, more or less curved and branching; secondary nerves frequent, especially near the top between the main nerves; reticulations transverse, irregular, clearly visible only on the upper surfaces; petiole very thick, bent, articulate in the middle, about 5 in. long. Hoya formosana Yamazaki 1968. Leaves fleshy broadly ovate 7-9cm long 4-5cm wide apex abruptly attenuate base rounded when dry flat very glabrous base near the midrib sparsely rough below glabrous with nerves roughly punctate, primary nerves few 2-3 pairs conspicuously curved, petioles thick 2-3cm long above the beginning of the young shoot roughly punctate otherwise glabrous. Hoya fraterna Blume 1849. leaves leathery ovate-oblong elliptic acuminate base rounded somewhat veined above the petiole callous-glandulose. Hoya fungii Merrill 1934. leaves leathery, penninerved, elliptic to oblong, 8 to 14 cm. long, 4.5 to 8 cm. wide, when dry pale, opaque, both sides glabrous uniform color or below a little paler, apexes acute and shortly acuminate, base rounded, midrib thick, below strongly elevated,

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lateral nerves on both sides about 7, not at all evident, arched meeting at the margins, about 30° angle with the midrib, reticulations loose, obscure; petiole 1.5 to 3 cm. long, glabrous. Hoya fusca Wallich 1831. leaves oblong, acuminate. Leaves approximately opposite, very outspread, strongly thick and leathery, firm, flat, oblong narrowly acuminate ½ inch terminal, margins parallel convex, base obtuse, above purple veined, shinny on both sides of the petiole insertion a gland withered (at this place a solitary cilia) below flat, somewhat green, midrib very much elevated, various transverse nerves, capillaries near the margins anastomosing, 6 to 7 inches. Hoya fuscomarginata N. E. Brown 1910. Leaves thick fleshy; blade 17-22 cm. long, 7-8 cm. wide. ovate-lanceolate, sometimes slightly oblique, acute or gradually somewhat acuminate, obtuse or somewhat wedge-shaped, 3-5 nerves, glabrous, green, margins dark brown; petioles 2.5-4.5 cm. long, 7 mm. thick. Hoya gigantangensis Kloppenburg 1992. Leaf Blade: 11-16 cm. x 2.4- 2 cm., pinnate venation, both surfaces glabrous. Petiole: heavy, curved 1.3-1.6 cm. long, probably grooved above. Hoya gigas Schlechter 1913. Leaves erect spreading or outspread, oblong or elliptic oblong, acuminate, base rounded, glabrous above, below very shortly tomentose, petiolate, with the petiole minutely tomentose, above lightly grooved. Hoya gildingii Kloppenburg. linear lanceolate opposite leaves, lax or with the petioles upright along the stem. Hoya glabra Schlechter 1908. loosely leaved; leaves outspread ovate acuminate, base rounded, glabrous, fleshy texture, 14—16 cm long in the middle 7.5—9.5 cm wide, 3 primary nerves especially conspicuous, petiole thick 2.5—3.2 cm long. Hoya globiflora Ridley 1915. Leaves ovate subacute with rounded bases 6.5 cm. long, 3.5 cm. wide, nerves 3 pairs and nerrules few visible when dry, one pair from the base ascending, the others short horizontal soon broken up, petiole very thick, 6 mm. long. Hoya globosa LeConte 1912. None Hoya globulifera Blume 1849. leaves almost sessile and ovate elliptic acute with the base rounded fleshy flat lightly veined above toward the base large white spots below punctate.

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Hoya globulosa Hooker 1883. leaves 5-7 in. long-petioled elliptic or oblong cuspidate or acuminate base rounded midrib very stout, nerves distinct, peduncle long. Hoya golamcoana Kloppenburg 1991. The leaves are 5.3 cm. long and 2.7 cm. wide at the widest portion, enervis, although in dried specimens the midrib and five pairs of pinnate nerves are visible. It is a densely leaved hanging vine with almost sessile petioles, 0.3 cm. long. Hoya gonoloboides Regel 1883. Leaves membranaceous, penninerved, reticulately veined, cordate ovate, acuminate, petiolate, both sides with long ascending hairs, hairs below on the nerves. Petioles and pedicels also with long hispid hairs. Hoya gracilipes Schlechter 1905. leaves spreading outspread, lanceolate or elliptic-lanceolate acuminate, both sides glabrous, texture leathery, shortly petiolate, 7.5-10 cm. long, below the middle 2-3 cm. wide. petiole 0.4-0.7 cm. long. Hoya gracilis Schlechter 1908. loosely leaved, rooting; leaves shortly petioled spreading, lanceolate-elliptical acute or somewhat obtuse, base cuneate, glabrous, lucid, texture thick leathery, 5.5-7.5 cm long, in the middle almost 1.8-2.3 cm. wide. Petiole somewhat thicker 0.5-1 cm. Hoya grandiflora Blume ex Decaisne 1844. leaves oblong or elliptic-oblong base rounded apexes abruptly and shortly acuminate pubescent above somewhat soft and scattered below densely tomentose velvety reddish (When dried?). Hoya grandiflora Sprengel 1825. Hoya leaves cordate-ovate acute. Hoya graveolens Kerr 1939. Leaves oblong-elliptic or oblong oblanceolate, base broad wedge shaped or rounded, apex briefly acuminate, margins flat, 5 to 8.3 cm. long, 2 to 4.8 cm. wide, leathery, glabrous, midrib above sunken below almost flat, lateral nerves 4 to 5 pairs often ascending with an angle of 60° difficult to observe; petioles 4 to 6 mm. long, thick, above concave, glabrous. Hoya greenii Kloppenburg 1995. the veins of the leaves of this species are perpendicular to the midrib and finely reticulate, whereas those of Hoya darwinii Loher are at 45 degrees to the midrib, venation in neither is prominent………. Leaf blade: 12-15 cm. long x 3-4 cm. wide, narrowly elliptic, both surfaces glabrous, dull, very dark green, above slightly channeled along the midrib area, lighter green below. 10 +/- pairs of veins perpendicular to the midrib,

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somewhat obscure, with fine secondary reticulate venation with the base tapered acuminate, outer apex curved under acuminate, rigid. Hoya griffithii Hooker 1883. Leaves distant, 1-2 in. diam., midrib stout, nerves quite obscure in some, in others distinct and forming large areoles; petiole ½ - 1 in.. very thick. Hoya gymnanthera Wight 1834. The specimen of this plant which I have seen are leafless. Hoya hainanensis Merrill 1923. leaves when dry papery, ovate-elliptic, penninerved, opposite, 6 to 8 cm long, 2.5 to 4 cm broad, both ends somewhat equally narrow, apex acuminate, base acute, nerves about 4 on both sides slender, obscure, almost oblique, reticulations loosely arranged, inconspicuous; petiole 8 to 10 mm long. Hoya halconensis Kloppenburg 1990. Foliage 9-12.5 cm. x 4-6 cm. with most small, spreading, glabrous, base broadly rounded, rather rectangular tapering slightly to the mucronate apex. Midrib visible above and below, lateral nerves obscure. Hoya halophila Schlechter 1913. Leaves spreading or outspread, petiolate, broadly ovate or somewhat round-ovate, briefly acuminate, base very obtuse, both sides glabrous, lucid above, opaque below, fleshy. Hoya hasseltii Miquel 1849. leaves ovate acute or acuminate base somewhat cordate margines somewhat wavy glabris below white-punctate. Hoya hellwigiana Warburg 1907. leaves cuneate-lanceolate or oblong 9 to 15 cm long, apex and base acute thickly leathery, veins on both sides about 2 ascending almost entirely inconspicuous. Hoya heuschkeliana Kloppenburg 1989. Leaves without nerves, petiolate, ovate-elliptic obtuse, base wedge-shaped, fleshy, on the back puberulous both sides punctate (pustulous or granulate) 2 to 5 cm. long, in the middle 0.5 to 2 cm. wide, above convex. Petioles curved about 0.2 cm. long. Hoya heuschkeliana subsp. cajanoae Kloppenburg & Siar 2007. Foliage: opposite, ovate 2.5 – 4.8 x 1.9 -2.9 cm. glabrous, flat with edges slightly rolled under, lower surface finely circular granulate (pustulate). Petiole curved 0.8 cm. long , terete, glabrous with a grove above near leaf blade. Hoya hookeriana Wight 1834. leaves fleshy oblong-lanceolate 2 nerved (6 to 10 inches long 1 to 1 ½ wide).

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Hoya hypolasia Schlechter 1913. Leaves spreading or outspread, petiolate, lanceolate, acuminate, base somewhat cordate, leathery texture, glabrous above, lucid, shortly and densely puberulous below, petiole grooved above, very shortly and softly puberulous. Hoya imbricata Decaisne 1844. leaves aborted alternately unilaterally, imbricate orbicular above convex veinless below veined with reflexed margins. Hoya imperialis Lindley 1846. leaves leathery narrowly oblong, ciliate, glabrous, without veins, apiculate, recurved apex with the base rounded or slightly cordate, peduncles tomentose, leaves hanging and small. Hoya inconspicua Hemsley 1894. Leaves of primary flowering branches often reduced to filiform rootlike filaments; secondary leaf branchlets strongly abbreviated often erupt from the axils (?) bearing a single pair of leaves, one nearly reduced, briefly petiolate, thickly leathery or somewhat fleshy narrowly lanceolate. Hoya incrassata Warburg 1904. leaves thick and coriaceous, ovate to oblong, 9-14 cm. long, 5-6 cm. wide, bases rounded or a little subcordate, apexes acuminate, acute, glabrous and shiny above, costa beneath prominent, veins on both sides, 4-5 spreading, curving and meeting before reaching the margins. Hoya incurvula Schlechter 1916. loosely leaved; leaves outspread petiolate, ovate elliptic, obtuse, fleshy, glabrous, margins thickened below, 2.5-4 cm. long, above the middle 1.2-2 cm. wide, petiole fleshy, thick 3-5 mm. long. Hoya inflata (P. I. Forst., Liddle & I. M. Liddle) 2007. fleshy leaves. Hoya intermedia Schlechter 1913. not published Leaf blades: thin many in close clusters 6-10 cm. mostly 7.5 cm. long. glabrous, narrowly elliptic, thick base narrowing into petiole, apex acute long tapering to sub rounded. Waxy rounded leaf edge. Petiole 1-1.5 cm long, thin. Hoya intermedia A. C. Smith 1942. leaves; petioles thick (about 3 mm. In diameter) when dry very rugose 10 to 20 mm Long; blades fleshy oblong, 8 to 12 cm. long 2 to 4.5 cm. wide, base obtuse, apex somewhat acute, penninerved, midrib above lightly impressed below prominent, secondary nerves on both sides 5 to 7 ascending often sometimes immersed, both sides prominent, vein sometimes prominent on both sides.

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Hoya ischnopus Schlechter 1913. None. Hoya juannguoana Kloppenburg 2002. leaves, obovate to ovate-lanceolate, base cuneate, edges entire, with apiculate acute apex. Blades usually somewhat cupped, waxy medium green upper surface, lighter beneath. Pinnate venation 3 veins on either side of midrib not readily visible (in contrast to those of H. finlaysonii Wight) 4 ½” – 6” long 2- 2 ½” widest near the middle or beyond. Nodal length variable, stems corky, tan colored, new growth rose bronze, the whole plant glabrous. Hoya kanyakumariana Henry & Swaminathan 1978. Leaves 1.5-3.3 x 1.5-2.2 cm, obovate to obcordate, entire, fleshy, glabrous, rounded or emarginate at apex, tapering at base; mid and lateral nerves obscure; petioles 2-6 mm long, thick. Hoya kastbergii Kloppenburg 2003. Leaves: nearly round 6 cm. long x 4.5 cm. wide, with variation, nerves obscure, glabrous, medium green.

Hoya kenejiana Schlechter 1913. loosely leaved, round, glabrous. Leaves spreading or outspread, petioled, narrowly oblong, somewhat obtuse, base rounded, fleshy, both sides glabrous, with a short petiole, lightly grooved above. Hoya kentiana Burton 1991. Folia 3-12 cm. longs X 0.8-1.5 cm. late, utrinque acuti; petioli ½ -1 cm longi. Hoya kerrii Craib 1911. Leaves opposite, obovate or orbicular, apex bilobed lobes rounded, base broadly wedge-shaped or somewhat rounded 4 to 12 cm. long, 5 to 9.5 cm wide, thickly fleshy, both surfaces glabrous, margins turner under; petiole 5 to 15 mm. long, 5 mm. thick. Hoya keysii Bailey 1884. Leaves ovate-cordate 1 to l 1/2 inches long, 3/4 to 1 inch wide, thick, fleshy, prominently penninerved; conspicuously anastomosing within the margin. Hoya kingdonwardii P. T. Li 1994. leaves opposite, succulent, elliptic, 1 – 1.7 cm x 0.5 – 1 cm, both ends acute, glabrous; petiole 1 mm. long. Hoya kloppenburgii Green 2001. opposite, lanceolate leaves, matte, dark green upper surface with occasional flecking, erose, heavy, thickened edge to blade, underside dull purple with 2-4 mm green edge, 10 – 21 cm long and 4 - 5 cm wide, apex acute base cuneate. Pinnately veined, with secondary veins at 45 60 degrees to midvein, nearly obscure. Petiole curved, 0.7 cm long and 0.4 cm in diameter.

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Hoya klossii Moore 1916. Leaves with short petioles, ovate, fairly long cordate-acuminate, apex obtuse, somewhat rounded, margines recurved, leathery, glabrous, surface skin shiny and (especially the upper surface) with prominent reticulate nerves, 4 to 6 by 1.5 to 2.5 cm. Hoya krohniana Kloppenburg 2009. Small cordate deep green opposite foliage.

Hoya kuhlii Koorders 1912. Leaves lanceolate acuminate, base narrow: Hoya kwangsiensis Tsiang et P. T. Li 1974. leaves long lanceolate up to 17 cm long and flowers much larger 3 cm in diameter. Leaves fleshy and leathery long lanceolate or lanceolate-oblong 8 to 17 cm long and 2 to 4 cm wide, margines slightly recurved, midrib on both sides nearly free of nerves and at other times not at all obscure, petioles strong grooved above 1 to 2 cm long apex with small blackish gland. Hoya lactea S. Moore 1914. leaves petiolate oblong-ovate with the apex rounded, sometimes itself briefly acute and cuspidate with a rounded base very slightly cordate thick leathery both sides glabrous. Hoya lacuna Wight ex Don 1837. foliis coriacco-membrannaccis cordatis ovatisve acuminatis supra pubescentibus subtis velutinis. Hoya lacunosa Blume 1826. leaves veinless ovate acuminate glabrous laciness above beneath convex. Hoya lambii Green 2000. leaves opposite, elliptic with apiculate tips and prominent palmate venation, substance papery, with a matte surface…….. Leaf blade: Elliptic, 10-15cm long x 610cm wide x 1mm thick, matte surface, light green. Leaf venation: Palmate Petiole: 45mm diameter x 10 -15mm long, rough surfaced, tan and green. Hoya lamingtoniae Bailey 1898. Leaves lanceolate-acuminate, 2 to 3 in. long, scarcely exceeding 1 in. wide, of rather pale-green, the gland on the face indicating the position of the attachment of the petiole somewhat raised; lateral nerves few, very oblique, triplinerved at the base, glabrous on both sides, the underside more or less speckled with purple. Petiole 3 or 4 lines long. Hoya lanceolata Wallich ex Don 1825. leaves lanceolate acuminate glabrous.

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Hoya lanceolata Moore 1916. Leaves shortly petioled, ovate-oblong, above ending in a tale or attenuate, apex obtuse, base obtuse briefly rounded, decidedly nerved, fleshy-leathery, both surfaces glabrous, 5.5 to 7.5 by 1.3 to 2.9 cm. peduncles shorter than the leaves. Hoya lancilimba Merrill 1932. leaves lanceolate to oblong-lanceolate, flashy, leathery 11 to 14 cm. long 2.5 to 3.5 cm. wide, both ends somewhat equally narrow, base acute, apex acute acuminate, in the dry state pale, wrinkled, with few primary nerves on either side, spreading, obscure arching and anastomosing; petioles thick, 2 to 3 cm. long. Hoya lantsangensis Tsiang et P. T. Li 1974. (This is not a hoya) Leaves erect, flat, thick or somewhat leathery all together glabrous, blades somewhat triangular base wedge shaped and apex obcordate rarely truncate 2 to 3 cm long and 1.5 to 9.5 cm wide, lateral nerves obscure, when dry wrinkled on both surfaces, petiole glabrous about 2 mm long, apex with a very small gland. Hoya lasiantha Blume ex Korthals 1849. Leaves ovate briefly cuspidate with obscure veins. Hoya latifolia G. Don 1838. leaves broad-ovate, acuminated, obtuse or sub-cordate at the base, 3-nerved, fleshy, glabrous. Hoya lasigynostegia P. T. Li 1984. Leaves fleshy when dry, thick paper like, ovate lanceolate 4.5-7 cm. long 1-2.7 cm. wide, apex caudate (tail like) base rounded, above both sides flattened with lateral nerves 4-5 pairs curved ascending anastomosing far from the margines, petiole fleshy 2-4 mm long puberulous becoming glabrous, apex provided with a small brownish gland. Hoya laurifolia Decaisne 1838. leaves lanceolate oblong lanceolate, above petiole glandular margins somewhat reflexed undulate leathery very glabrous. Hoya laurifolia Miquel 1838. leaves lanceolate oblong lanceolate, above petiole glandular margins somewhat reflexed undulate leathery very glabrous. Hoya laurifoliopsis Hochreutiner 1936. Leaves not very fleshy, but nevertheless more or less coriaceous, opposite; petioles glabrous, lenticellate, more or less laterally compressed; leaves very glabrous, elliptic, margins entire and a very little revolute, apexes long acuminate, acute, bases long attenuate and toward ( the margins revolute acuminate. At the leaf base where the petiole meets it, probably glandular. Dried midrib and secondary nerves (about 5 on both sides) more or less conspicuous.

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Hoya lazaroii Kloppenburg & Siar 2007. Foliage: Glabrous, narrowly elliptic, apex acute, base cuneate to sub-rounded, enervis length 11.0 cm. and 2.8 cm. wide. Hoya leembruggeniana Koorders & Schumacher 1911. Leaves elliptic-oblong, obovate-oblong or rhomboid-oblong, fleshy, glabrous, base wedge shaped, apex acute or somewhat acuminate, base one nerved, secondary nerves very obscure, 4 to 6 ½ cm wide. Petiole thickened, ½ to 1 cm long, glabrous. Hoya leucantha S. Moore 1916. Leaves among the smallest, shortly petioled, lanceolate, acuminate, base obtuse slightly rounded margines recurved, leathery, both sides glabrous, 3.5 to 4.5 cm. long, near the base 8 to 10 mm. broad. Hoya leucorhoda Schlechter 1913. Leaves spreading or outspread, petiolate, ovate acuminate, base rounded, texture leathery, both sides glabrous. Hoya leytensis Elmer ex Burton. Leaves coriaceous and subglaucous, more or less persistent, glabrous, opposite or in subwhorles at the ends of abortive branchlets, quite variable in size and shape, entire, narrowly oblong to rotund, most of them are from 2-3.5 cm. long and from 6-10 mm. wide across the middle or widest portion, the rotund ones measure 10 by 12 mm., the normal or the oblong blades equally tapering toward both ends, obtuse to subacute or even rounded for the smaller rotund to elliptic blades which are in the minority, flat, grayish to brown on my dry specimen; midrib quite fine or obscure, plainer on the lower surface, lateral nerves none, only the upper side marked with faint transverse lines; petioles 3-5 mm. long or much shorter, dark brown, crinkle rugose when dry, leaving orbicular flat scares upon falling Hoya liangii Tsiang 1936. Leaves fleshy obovate or obovate-oblong 4.5 to 7.5 cm. long 3 to 4.5 cm. wide, apex rounded or obtuse, base somewhat cordate or rounded, above shiny green below dull when dry pale yellow, lateral nerves 4 in pairs, veins faint, above the petiole 2 to 5 glands, petiole thick 1.5 cm. long. Hoya lii Burton 1991. Hoya limoniaca S. Moore 1921. leaves far apart, moderately thick; leaves 7 to 8 by 3 to 3.5 cm., petiolate (petiole all 15 mm. long) ovate, attenuate towards the apex the apex itself obtuse, base rounded-obtuse, 5 nerved, moderately thick, leathery. Hoya linavergarae Kloppenburg & Siar 2006.

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The growth of this plant is similar to Hoya cembra Kloppenburg, Hoya odorata Schlechter and Hoya paziae Kloppenburg except here the leaves are more undulant and rigid reminding me of a holly leaf. Foliage opposite, glabrous blade 4-5 x 1.1-2.7 cm. Hoya lindaueana Koorders 1911. Leaves opposite, elliptic, when dried lead gray, bases acute, apexes shortly-somewhat-minutely-acuminate, 3-plinerved (or indistinctly 5-plinerved) 12-21 cm long and 5-10 cm wide, glabrous. Petiole 2-2.5 cm long, glabrous. Hoya linearis Wallich 1825. leaves lanceolate obtuse fleshy both sides stem-like filiform densely pilose. Three times smaller than above (H. lanceolata) and more slender. Hoya lipoensis P. T. Li 1985. Leaves long elliptic, 9 to 15 cm long by 3 to 5 cm wide, apex caudate, cauda 1.5 cm long curved, base wedge-shaped asymmetrical, lateral nerves parallel 10 to 12 pairs, prominent above, obscure below, anastomosing near the margins. Petiole 1 cm long. Hoya litoralis Schlechter 1905. loosely leaved, with aerial roots; leaves outspread , spreading, elliptic-lanceolate, acuminate, with the base generally narrowing to the petiole, both surfaces glabrous, texture thickly leathery, 4.5-7 cm. long, near the middle 1.3-2.2 cm. wide, petiole about 0.7 cm long. Hoya lobii Hooker 1883. leaves 2-3 in. subsessile elliptic or ovate acute or ovate acuminate very fleshy, nerves 6-8 pairs arched spreading…………. Leaves 1-2 in. diam., base narrowly cordate; midrib rather slender. Hoya loheri Kloppenburg 1991. Leaf blade: 2.6-4.4 cm. long, subovate, enervis, glabrous 1.2 cm. at the widest. Petiole: c. 0.5 cm. long 0.15 cm. in diameter, grooved above, bare, rugose. Hoya longifolia Miquel 1857. leaves at the base somewhat wedge-shaped otherwise obtuse oblong lanceolate, acuminate, fleshy glabrous 4 to 2 inches long, peduncles somewhat germinal multiflowered. Hoya loyceandrewsiana Green 1994. loosely leafed with flat to conduplicate leaf blades, 10-20 cm wide, 15-25 cm long; leaf petiole 1 cm thick x 1x8 cm long with a thick callus on the dorsal side at the blade attachment; leaf blade thick, 2 to 5 mm, ovate to widely cordate with acute tip and obtuse to cordate base; palmately veined with 3 major veins, 2 secondary palmate veins and network of tertiary veins. Hoya lucyae Kloppenburg & Siar 2006.

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Leaves narrowly elliptic to oblanceolate, margins entire not revolute, somewhat concave above, Enervis, base cuneate, widest toward the outer apex that is acute, fine scattered hair cells especially on new growth on blade, stems, and petioles, all surfaces. Blades 6-10 cm. long and 1.8 – 2.0 at the widest. Glossy deep waxy green above, lighter and more dull on the under surface. Petioles: straight or curved, terete, 0.25 cm. in diameter, 1.5 – 2 cm. long, with small gland at blade juncture or none, matt finish.

Hoya lutea Decaisne 1844. leaves elliptic, base and apex round or at apex acuminated, briefly terminated, fleshy, lightly veined and cupped below, glabrous as in the preceding (ariadna) but a little smaller and rounder, pale green. Hoya luzonica Schlechter 1904. leaves far apart; with the leaves outspread oblong obtuse or acute; both surfaces glabrous, leathery-fleshy about 9-13 cm. long, in the middle almost 5-6 cm. wide, petiole somewhat fleshy, glabrous 1.5-2 cm. long. Hoya lyi Leveille 1907. opposite leaves, leathery on both sides corrugated and pubescent, broad green, margins entire, margins revolute, ovate-lanceolate, one nerved, at the apex and base obtuse shortly petioled. Hoya macgillivrayi Bailey 1914. Leaves subcoriaceous, about 4 in. long, 2 in. broad across the blunt cordate base, from which it tapers to an acuminate apex; margins recurved; nervation seen under a lens quite dense, the nerves alone prominent and only a few of these conspicuous. Petioles under 1 in. long, minutely glandular at its junction with the lamina. Hoya macrophylla Blume 1826. leaves trinerved ovate or ovate-oblong acuminate fleshy. Hoya macrophylla Wight 1834. leaves broad ovate acuminate base obtuse somewhat cordate 3 nerved fleshy glabrous (7 to 10 inches long 4 to 10 wide). Hoya magniflora P. T. Li = Hoya grandiflora Blume Not a hoya Hoya maingayi Hooker 1883. Leaves very thick, fleshy, coriaceous when dry; elliptic-lanceolate or oblanceolate, shortly acuminate at apex, long acuminately narrowed at base; both surfaces glabrous, dull when dry; margins recurved; 7 to 9 in. long, 2.5 to 3.5 in. broad; midrib slender but prominent when dry, with a gland or enlargement at the base above the petiole; main nerves about 5 pairs, prominent when dry, especially above, at about 60° to 70° with the midrib, nearly straight at first, then branching and forming a rough

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intramarginal nerve; secondary nerves few; reticulations irregular, distant; petiole thick, .25 to .5 in. long. Hoya manipurensis Deb 1955. Leaves simple, opposite, decussate, thick, coriaceous, puberulous, exstipulate, petiolate, obovate, truncate, pale gray in colour; veins obscure 3-5, at first parallel then diverging, apex slightly depressed and mucronate; base acute attenuated; margin entire; petioles pubescent, 2-4 mm. long, 2-2.5 cm broad at the broadest region. Hoya marginata Schlechter 1905. loosely leafed, rooting; leaves spreading, elliptic-ovate, acuminate, both sides glabrous, thick leathery, 5-7 nerved, with nerves almost fan shaped, ornate, 13-17cm long, in the middle 5.5 - 10 cm broad, petiole thick, up to 4cm long. Hoya matavanuensis Kloppenburg & Siar 2009 Leaves glabrous, elliptic, attenuate apex 2.5 -5 cm. long, 1 - 1.5 cm. wide; nerves obscurely pinnate, finely netted, whole plant here fine and delicate with relatively long peduncles & large flowers. Petioles glabrous, 0.5 -0.7 cm. long ca 0.1 cm. in diameter, grooved above. Hoya maxima (Karsten) Koorders 1920. leaf-base is very united fused folded together or ± fragile and sometimes, profoundly (heart-shaped). Hoya maxima Teijsmann & Binnendijk 1863. leaves thick fleshy, elliptic-oblong, shortly acuminate, bases rounded, both sides glabrous, margins reflexed, petiole thick. Hoya mcgregorii Schlechter 1906. Leaves widely spread, lanceolate-elliptic, acuminate, glabrous coriaceous; 7-11 cm. long 2.5 -3.5 cm. wide in the middle, triplinerved. Petiole fleshy, almost terete, glabrous, 1-1.5 cm. long. Hoya megalantha Turrill 1915. Leaves elliptic-ovate, apexes attenuate acuminate, bases rounded, up to 6.5 cm. long and 3 cm. wide, entirely, glabrous, somewhat leathery, not very fleshy, midrib impressed above, prominent below, about 5 lateral nerves on both sides, somewhat prominent on both surfaces, as are the transverse reticulations; petiole about 1 cm. long, round glabrous. Hoya megalaster Warburg 1901. None Hoya mekongensis M. G. Gilbert & P. T. Li. leaf blade oblong-oblanceolate, 10-133 x 4-5 cm, minutely hairy, base rounded, apex acuminate to caudate-acuminate; lateral veins ca. 7 pairs, obscure.

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Hoya meliflua Blanco ex Merrill 1837. Opposite leaved, oblong, round, somewhat fleshy, hairless without veins with a gland at the base. Hoya memoria Kloppenburg sp. nova 2004. leaves shortly petioled, elliptic-lanceolate, base cuneate glabrous.

Hoya membranifolia Costantin 1912. Leaves lanceolees, attenuated in corner or a little rounded toward the basis, acuminate, very sharp to the top, very thin, translucent on the edges, glabrous; prominent median rib underneath; nerves secondary 5 pairs, inserted perpendicularly from the median rib but immediately arching themselves toward the top and coming 1 mm. from the sides of the leaf; limb 7-12 cm. long on 14-32 mm. glabrous lateral, stalked Hoya mengtzeensis Tsiang & Li 1974. . Foliage thinner than general, lanceolate, apexes acuminate, base attenuate and running down into the petiole 6-11 cm long, 1.5 -2 cm broad, midrib impressed above, nerves and veinlets when dried rugose, petioles 1-2 cm long provided with 2-3 small bunched glands at apex. Hoya meredithii Green 1989. loosely, appositely leaved (sometimes lacking the twin of a pair); leaf open, with undulate margins, ovate with acuminate tip, obtuse base, subauriculate, with one large gland, hard and rigid blade; 7.5 to 18 cm. wide by 12 to 30 cm. long; petiole hard and thick, 8 mm. in diameter by 2.5 - 3 cm long, twisted; leaf venation pinnate, with 8 - 12 secondary veins ending in reticulations, showy, all veins dark green against a light blade. Hoya merrillii Schlechter 1904. leaves far apart, fleshy; leaves outspread, ovate acuminate, fleshy, both sides glabrous, with 5 prominent nerves when dry, 6-9 cm. long, below the middle 5-6 cm. wide; petiole short fleshy, 1.5-2 cm. long. Hoya micrantha Hooker f. 1883. leaves ovate or elliptic lanceolate acute thick and fleshy, margins flat. Hoya microphylla Schlechter 1913. loosely leafed, round, sparsely puberulous, erect, spreading, ovate, somewhat obtuse, margins recurved, reticulated nerves, leathery, on both sides glabrous; petiole puberulous above lightly furrowed. Hoya microstemma Schlechter 1913. Leaves spreading or outspread, shortly petioled, elliptic, obtuse, wedge shaped base, fleshy, both sides glabrous. Hoya minahassae Schlechter 1916.

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, loosely leafed; leaves petiolate, spreading, elliptic or lanceolate-elliptic, obtuse or almost obtuse, bases wedge-shaped fleshy, both aides glabrous, 4-7 cm long, in the middle almost 1.5 to 2.5 cm wide, petiole thick 7- 10 mm long. Hoya mindanaensis Elmer 1938. Leaves opposite, persistent, scattered, mainly descending, thickly coriaceous, yellowish green beneath, tips recurred, otherwise nearly flat or only the entire margins slightly imbricate, also glabrous and smooth, curing subolivaceus on both sides, oblong or more often ovately oblong, base broadly rounded and sometimes obscurely emarginate, the upper portion of the laminae gradually tapering into the acute to subacuminate apex, the average blades 15 cm long by 6 cm wide across the middle or a trifle below it, persistent; midrib thick and pronounced beneath clear into the apex and olivaceus in color, on the upper face deeply and narrowly grooved, totally glabrous on both surfaces; nerves faint, 3 to 5 on each side of the midrib, slightly ascending and straight, forked from above the middle and reticulately united, sometimes with secondary nerves in between, reticulations coarse and only visible from beneath; petioles up to 2 cm in length, blackish brown on my dried specimen, also thick, rugose, glabrous, calculate along the upper side, persistent, leaving a triangularly shaped scar after falling.

Hoya mindorensis Schlechter 1906. loosely leaved, with the leaves outspread or spreading, oblanceolate-elliptic, shortly acuminate, glabrous, leathery textured 9-12 cm. long, above the middle 3.5-4.5 cm. wide, with the petiole thick, somewhat terete (circular), 2-2.5 cm. long. Hoya mindorensis subsp. superba Kloppenburg 2005. None. Hoya minima Costantin 1912. Leave glabrous, opposite, lanceolate, very small, thin, attenuate at the base, acuminate at the apex, wrinkled, reticulate in dry state; nerves not apparent except when light shines through; secondary nerves not visible, very oblique, towards the top, 4 to 5.5 cm long by 7 to 10 mm wide; petiole 5 to 5 mm, glabrous. Hoya mitrata Kerr 1940. Leaves oblanceolate or elliptic, base rounded and lightly eared, apexes briefly acuminate, margins flat, 6.5 to l2 cm long, 4.7 to 6 cm wide, leathery, dried gray, glabrous, above shining, beneath opaque, costa above concave, below prominent, lateral nerves 5-6 pairs towards margins very arching and 5-6 mm from margins anastomosing with network veining prominent on both sides; petioles 6-8 mm long, above concave, glabrous. Hoya monetteae Green 2004. Allied to Hoya diversifolia Blume but differing in that the foliage larger 20 cm. x 10 cm versus 5-10 cm. x 3.75-7.25cm.and flowers larger 2cm in diameter verses 1.3cm, and with clear sap versus milky.

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Hoya montana Schlechter 1913. Leaves erect-spreading or outspread, shortly petiolate, oblong-elliptic, shortly and somewhat obtusely acuminate, bases obtuse, somewhat fleshy, both sides glabrous, petioles very short, above lightly grooved. Hoya motoskei Teijsmann & Binnendijk 1836. (not from Type description). leaves fleshy, ovate-rounded, acute, base slightly cordate, margins reflexed, glabrous above, pubescent below, with long petioles. Hoya mucronulata Warburg 1901. (not from Type description). leaves leathery oblong 6 to 12 cm, long, 3 to 5 cm. wide rounded on both ends, apex sometimes briefly, obliquely mucronulate, glabrous on both sides, above on midrib minutely glandular, veins on both sides 7 to oblique long, meeting and joining at the margins. Hoya multiflora Blume 1823. leaves somewhat leathery veined wedge-shaped oblong or oblong-lanceolate glabrous below paler, stiff hairs at base. Hoya myanmarica P. T. Li 1995. leaves opposite, succulent, oblong 4.1 – 16 cm. x 1 – 3.3 cm. base cuneate or rounded, apex acuminate; petioles 2 – 3 mm. long, apex with 2 or 3 glands, umbel terminal, 6-8 cm. long. Hoya myrmecopa Kleijn & van Donkelaar 1999. Leaves 2.5--7.5 cm long, ovate to elliptic, acute to acuminate, venation obscure. Petioles 0.5--1.5 cm long. Hoya nabawanensis Kloppenburg & Wiberg 2002. Opposite leaved, fleshy, petioles curved to slightly bent, terete, 5-6 times shorter than the leaf blade, 1 cm. long, glabrous. Blades ovate to ovate- lanceolate apex acute to somewhat apiculate, flat to slightly curved down at the apex, deep green, undersides slightly lighter in color; glabrous 6 - 10 cm. long by 2.5 - 4 cm. wide, pinnately veined about 4 on each side of the midrib at 45 degrees to it, midrib slightly raised on the upper side, edges entire, texture thick and leathery, sap milky white. Hoya naumanii Schlechter 1908. loosely leaved; leaves petiolate spreading outspread, broadly elliptic, widely apiculate, with the base somewhat cordate-rounded both sides glabrous, about 10 cm. long, near the middle about 7.5 cm. wide, petiole somewhat thickened, about 2 cm. long. Hoya neoebudica Guillaumin 1937. leaves 3 to 4 together, lanceolate (7 to 9 cm. by 3-3.5 cm.), base wedge-shaped, apexes acute acuminate, glabrous, fleshy, 5 ply nerves, petiole 1 to 1.5 cm. long.

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Hoya neocaledonica Schlechter 1906. loosely leaved, with the leaves erect-spreading very outspread, ovate or ovateoblong acuminate, both sides glabrous, texture fleshy, 3 primary nerves very conspicuous above on the herbarium specimen (dried), petiolate, blade 5-9 cm long, in the middle and below the middle 3-5 cm. wide, petiole fleshy, round 1-2 cm. long. Hoya neoguineensis Engler 1886. leaves supported by lightly channeled petioles 3 to 4 times shorter than the leaves, oblong-ovate with obtuse bases, almost truncate, distinguished by long acute accumulation, having about 6 thin lateral nerves on each side below (on dried leaves only) scarcely prominent. Hoya nervosa Tsiang et P. T. Li 1974. Leaves large ovate or ovate-oblong 8.5 to 17 cm long 4 to 6.3 cm wide, apexes acute or shortly acuminate and bases rounded, costa nerves and veins very conspicuous; lateral nerves 5 to 6 pairs arching-ascending anastomosing next to margins veins reticulate, petiole strong 1 to 2 cm long with a few glands at apex, 2 to 3 together. Hoya nicholsoniae Mueller 1866. leaves ovate acuminate, somewhat fleshy, flat, 5-nerved, reticulate-veined; petiole often longer. Hoya nicobarica R. Brown ex Traill 1827. (I do not have original text) leaves ovate-lanceolate. Hoya nummularia Decaisne ex Hooker 1883. leaves very small orbicular shining. Hoya nummularioides Costantin 1912. . Leaves fleshy, small, rounded or oval, but with the apex obtuse and apiculate, very pubescent on both surfaces; nerves not visible; blades 9-16 mm., by 12-25 mm. long; petiole pubescent, 2-5 mm. long. Hoya nuuuliensis Kloppenburg & Siar 2009. Leaves, glabrous, thick wrinkled, elliptic, trinerved indistinct 6.8 cm. long , ca. 2 cm. wide, apex acute, base cuneate. Petiole glabrous, thick 1 cm. long (cannot see a groove above). Hoya nyhuusiae Kloppenburg 2003. None. Hoya obcordata Hooker 1883. leaves obcordate or obreniform, …Leaves ½ in. long, sometimes broader than long, lobes rounded, sinus acute, base rounded or subacute, rather thin, opaque, nerves obscurely reticulate; petiole 1/16- 1/15 in.

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Hoya oblanceolata Hooker 1883. leaves 4-6 in. narrowly oblanceolate acuminate very fleshy. Hoya oblongacutifolia Costantin 1912. Leaves glabrous, ovate-oblong, or lanceolate, pointed at base, acute or acuminate at apex, as a rule not very fleshy but nevertheless sometimes thick; secondary nerves 4-6 pairs, prominent, at 80° arching and running together as 1 nerve at margins. Leaves 5-8 cm by 2.5-3 cm; petiole 4-8 mm long. Hoya obovata Decaisne 1844. leaves obovate apexes rounded or rarely emarginate, bases thick attenuate, towards the petioles a callused gland arising from the limb, margins reflexed, both sides glabrous. Hoya obreniformis King 1901. leaves obcordate or obreniform, sometimes broader than long, lobes rounded, sinus acute, base rounded or subacute, rather thin, opaque, nerves obscurely reticulate, faintly visible above not evident beneath, bright-green above, paler beneath; length 1.25—2.5 cm., width 2.25—2.5 cm. Hoya obscura Elmer ex Merrill ex Burton 1986. Folia oppositae, glabra, laevis, stature variabilis, plane elliptica ad obovata-oblonga, in folia longiores, folia minores subelliptica 2 X 3 cm., aliae usque 10 cm. longa et fere 4 cm. late ad medium vel parve supra, gradatim contracta ad acuta-subacuainata apex, folia minores late obtuse ad basim, folia magnae cuneatae, interdum graciliter cuneatae omnes gradatum inter; costa non promines, laterales nervi 3-5 utrinque costa, admodum obliquus praecipue nervi basim, obscure furcati et plus minusve anastomosans ad margine; petioli 5-10 mm longi vel etiam usque 1.5 cm long), glabri, supra canneculeti. Hoya obscurinervia Merrill 1923. leaves opposite, fleshy, when dry leathery, pale, elliptic to oblong-elliptic, somewhat thin acuminate, bases for the most part acute, obscurely 3 or 5 plinerved, about 10 cm long, 4 to 4.5 cm wide, nerves thin, reticulations irregular; petiole thick, 2 to 2.5 cm long. Hoya obtusifolia Wight 1834. leaves fleshy oval-oblong bases of the apex obtuse cuspidate, above shining below transversely obscurely veined (dried). Hoya occlusa Ridley 1912. Leaves ovate acuminate in pairs, base obtuse glabrous above shining greenish when dry, below brownish, coriaceous, nerves 5 pairs rising at an acute angle, slender, elevated beneath, 4 inches long 2 inches wide, petiole 1 inch long. Hoya odetteae Kloppenburg 1998.

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Leaf Blade: l x 2-7 cm. Lanceolate. Matte midrib prominent, secondary veins obscure. Petiole: .25-1 cm. long. Hoya odorata Schlechter 1906. well leaved; leaves erect-spreading, short petioled, elliptic or lanceolate-elliptic, acuminate, glabrous, lucid, texture thinly leathery 3.5-5 cm. long, below the middle 1.32.2 cm. wide. Hoya oleoides Schlechter 1913. Leaves erect spreading or outspread, petiolate, elliptic obtusely acuminate, leathery, both sides glabrous, petiole short, somewhat thick, above lightly grooved.. Hoya oligantha Schlechter 1913. Leaves erect-spreading or outspread, shortly petioled, narrowly lanceolate, minutely apiculate or somewhat acute, base rounded, leathery texture, both sides glabrous, petiole lightly grooved above, glabrous, very short. Hoya oligotricha K. Hill 1986. Leaves fleshy, broadly pinnately veined, elliptical, ovate or orbicular, 4-12 cm long, 3.0-8:0 cm wide; petioles 1.0-3.5 cm long, 3-7 mm diam. Hoya onychoides P. I. Forster & Liddle 1995. Leaves petiolate; lamina lanceolate-ovate, up to 120 mm long and 55 mm wide, succulent, discolorous glabrous, with venation obscure, upper surface dark green; lower surface pale green; tip acute to shortly acuminate; base cordate; petiole 18-22 mm long, 4-5 mm diameter, grooved on upper surface; collators 3 or 4 at lamina base, often coalesced. Hoya opposita G. Don 1837. opposite leaves, the narrower petioles, which are convex beneath, and concave above. Hoya orbiculata Wallich ex Wight 1834. leaves fleshy orbicular glabrous. Hoya oreogena Kerr 1939. Leaves elliptic, base wedge-shaped or narrow slightly ear-shaped, apex somewhat obtuse margines plus or minus reflexed, 5 to 7 cm. long, 1.8 to 2.9 cm. wide, leathery, glabrous above, below with moderate copious short tawny pilose, when dry midrib obscure above bellow slightly prominent at the base narrowly winged. Lateral nerves obscure on both sides; petiole thick, warty, pilose, 5 to 8 mm. long. Inflorescence lateral, umbel-like, many-flowered: peduncles moderately robust, pilose 5 to 11 mm long. Hoya oreostemma Schlechter 1913.

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Leaves spreading or outspread, petiolate. lanceolate elliptic or elliptic, acuminate, base rounded, leathery texture, both sides glabrous, lucid above (shiny), below opaque, petiole glabrous, above slightly grooved. Hoya orientalis P. T. Li =Hoya reticulata Merrill 1920. Leaves fleshy when fresh, coriaceous when dry, ovate to oblong-ovate, shining, 8 to 12 cm long, 4 to 7 cm wide, the base broadly rounded, the apex slenderly and sharply acuminate, the epidermis on both surfaces coarsely reticulate-wrinkled when dry; petioles 1 to 2.5 cm long; lateral nerves about 4 on each side of the midrib, somewhat spreading. Hoya ottolanderi Koorders 1911. Leaves opposite very thick, 15 to 28 cm. long and 4 to 7.5 cm wide, glabrous distinct 5 nerved below above about 3 plinerved, lanceolate base acute, apex acuteacuminate. Petiole 1.5 to 3.5 cm long. Hoya ovalifolia Wight & Arnott 1834. leaves fleshy ovate, acuminate at both ends, peduncles shorter than the leaves. Hoya oxycoccoides S. Moore 1916. Leaves bright green and faintly shining above when dry, opaque and pale below; petioles 3-5 mm. long, puberulous. Hoya pachyclada Kerr 1939. Leaves obovate, bases cuneate or somewhat rounded apexes very shortly and obtusely acuminate 7.8 to 10.5 cm. long, 5.2 to 6 cm. wide, thickly leathery, both surfaces having a few sprinkled short hairs, dried costa and nerves on both sides obscure but seen with transmitted light, lateral nerves 2 pairs and basal (or almost at the base) arising acutely, arching and ascending almost reaching apex; petioles thick, above concave, minutely puberulous, 6-12 mm. Hoya pachyphylla K. Lauterbach & K. Schumann 1901. leaves with short petioles elliptic, obtuse and apiculate with base rounded fleshy shiny both sides very glabrous. Hoya pachypus S. Moore. leaves petiolate broadly ovate-oblong sometimes almost square both ends rounded apex often mucronate thick leathery surface on both sides glabrous. Hoya padangensis Schlechter 1916. leaves spreading and widely spaced, narrowly elliptic-lanceolate, acute or acuminate, 7 to 10 cm long, in the middle almost 1.3 to 2 cm wide, fleshy, glabrous, petiole fleshy about 5 mm, long. Hoya palawanica Kloppenburg 1990. This species has a very distinctive leaf with venation similar to Hoya finlaysonii Wight with looping inner pairs anastomosing…. Leaves spreading or outspread, petiolate

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lanceolate acuminate, texture leathery, with both surfaces glabrous 13.2-14.5 cm. long, in the middle 3.8-4.2 cm. wide, deciduous with 5 nerves, inner ones nearly reaching the apex with distinct reticulations. Petiole about 1.8 cm. long, 0.6 cm. in diameter, glabrous, grooved above. (from Schlechter). Hoya pallida Lindley 1826. leaves ovate-lanceolate acuminate fleshy veined. Hoya pallilimba Kleijn & van Donkelaar 1999. Leaves 3--8 cm long, ovate, acute, without obvious venation, margins on basal half of the leaves thin, pale green. Hoya panayensis Kloppenburg & Siar sp. Nova. Hoya panchoi Kloppenburg 1988. Leaves very thick, glabrous, with both surfaces minutely punctate, subdeciduous, edges straight and entire; of two distinct types, some round 1.5 cm. in diameter, others obovately oblong 2 cm. wide above the middle and 4 cm. in length, obtusely rounded at the apex, with a broad cuneate base. Petiole very short, thick, curved and expanded toward the base, leaving circular scars after falling. Hoya pandurata Tsiang 1939. Leaves erect spreading, a little fleshy, glabrous pandurate or narrowly oblong, margins involute, apex acuminate base rounded and in China very slenderly 2-eared 4 to 7 cm. long and 1 cm. wide (Wong's #74427, 10 cm. by 2 cm ), nerves on both sides not conspicuous, petiole glabrous about 4 mm. long apex glandular 2 to 4 fascicled. Hoya paniculata Sprengel 1825. Hoya leaves somewhat cordate  ovate acuminate very glabrous. Hoya papillantha K. Schumann 1898. leaves petiolate, petioles thick round, leaves oblong acute bases somewhat rounded fleshy both sides very glabrous; umbels outside the axils on short peduncles. Hoya papuana Schlechter 1913.

loosely leafed; leaves lanceolate-elliptic, long acuminate, both sides glabrous, texture thinly leathery, 6 to 8 cm long, below the middle 1 to 2.3 cm wide, petiole short, glabrous, 0.5 - 0.7 cm long; rachis very short, about same length as petiole, cylindric. Hoya parasitica Wallich 1830. Leaves ovate-lanceolate, fleshy, drooping…… Leaves, opposite, petioled, retrofracted, ovate-lanceolate, acute, of a firm fleshy texture, and smooth on both sides, veins scarcely conspicuous above, and invisible underneath; from two to four inches long, and about one broad. Petiole short, round, ash coloured, in fact, more like a part of a branch than a petiole.

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Hoya parviflora Wight 1834. leaves fleshy lanceolate base and apex attenuate. Hoya parvifolia Schlechter 1908. loosely leaved, having aerial roots; with the leaves shortly petiolate oblong or oblong-elliptic, very obtuse, both surfaces densely and shortly hispid, 1.2-1.7 cm. long, 0.8-1 cm. wide, petiole hispid about 0.3 cm. long. Hoya patella Schlechter 1913. loosely leaved, round, minutely and sparsely pubescent. Leaves spreading or outspread, briefly petiolate, oblong-elliptic, somewhat obtusely acuminate, below pubescent, petiole pubescent, above slightly grooved. Hoya pauciflora Wight 1848. leaves approximated, fleshy, glabrous, narrow lanceolate, blunt pointed; flowers few, generally paired, from a short thick peduncle. Hoya paxtonii Nichols 1885. Closely resembles the H. belle in habit and appearance, but has less fleshy, more acuminate and lighter green leaves. Hoya paziae Kloppenburg 1990. Foliage: held flat (surface to stem) at 90°-100° on twisted ½ turn petioles which are grooved above, also pale green 1 cm. long, 0.18 cm-0.20 cm. in diameter. a divided leaf gland at the base of the leaf tan in color, Midrib a little sunken an the upper surface, blade slightly concave on top edges slightly undulant, apex attenuate, narrowly acute and slightly turned under. Blade 6-7 cm. long and 2.5-3 cm. at the widest below the middle. Pinnately veined, 5 pairs not prominent, surface glabrous, waxy, not punctate, very smooth above, granular below, midrib extending below, channeled above. New growth pink-bronze. Hoya pedunculata Schlechter 1913. In the leaf form it is midway between Hoya halophila Schlechter and Hoya litoralis Schlechter. Hoya peekelii Markgraf 1927. . Leaves fleshy, elliptic, 12 cm. long, 7 cm. wide, shinny, apex briefly acuminate, netted veins sparsely hairy, base near the petiole 3 to 4 cm. long shortly connate. Hoya pentaphlebia Merrill 1918. Leaves thickly coriaceous, pale and shining when dry, broadly ovate to elliptic, 8 to 16 cm long, 6 to 9 cm wide, the apex broadly and rather prominently acuminate, base rounded, prominently 5-nerved, the interior pair of nerves reaching the apex, the reticulations very lax, distinct on both surfaces, the margins somewhat recurved; petioles stout, 2 to 5 cm long.

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Hoya pendula Wight & Arnott 1834. leaves fleshy glabrous from oblong-oval acute to broad ovate acuminate, margins revolute. Hoya perakensis Ridley 1910. Leaves ovate acute coriaceous glabrous base broad truncate rounded 4 ½ inch long 3 inches wide, nerves from the base 5, with few arched secondary nerves, drying brown with recurved edges, petiole thick ¼ inch long. Hoya phyllura O. Schwartz 1931. Leaves opposite, blade oblong-elliptic or very slightly ovate-oblong, bases abruptly narrowed, apex long and acute caudate-acuminate, glabrous in the dry state coriaceous shining, nerves very prominent, nervation pinnate veins 3 to 4 on each side disappearing very near apex, curving and anastomosing, nerves narrowly reticulate. Leaf margins entire and very lightly revolute, 8 to 16 cm long, 3.2 to 5.2 cm wide, acumen 1.5 to 2 cm long, in the living state lighter green beneath than above. Petioles robust 1 to 1.7 cm long. Lateral inflorescence umbellate, supported by robust short peduncles 2.3 to 3 cm 2.3 to 3 cm long. Hoya picta Siebold 1848. leaves ovate-lanceolate, apex attenuate acute, fleshy, arched in the middle, yellowish and when young purplish or azure-rose and margines green variegate. Hoya picta Blume 1849. leaves (small) slightly round to ovate, sharply pointed; both sides purple-stained and glandular; below convex. Hoya picta Miquel 1856. Leaves on short petioles, broadly ovate mucronate-acute, both sides purple-dyed and glandular, in the dried state almost three nerved, beneath convex, 4-5 lines long. Hoya piestolepis Schlechter 1913. Leaves spreading or outspread, petiolate oblong or elliptic, acuminate, base briefly cordate, texture leathery, both sides glabrous. Hoya pilosa Seemann 1856. the leaves more hairy below. Hoya polyneura Hooker 1883. leaves 3-4 in. subsessile rhombic-ovate or lanceolate acuminate very fleshy, nerves very many oblique straight parallel. Hoya pimenteliana Kloppenburg 1999. foliage is opposite, linearly lanceolate with an acute apex and cuneate to slightly rounded base, where a distinct gland is present. The leaves are very distinctive, of a deep dull (flat color) iridescent green, due to an extremely fine velvety upper surface. Venation

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is somewhat obscure but pinnate with veins at 45 ° to the midrib (noted when dry). Leaf edges are entire but with slight undulations, the edges being slightly turned under. The foliage is profuse……… Leaf: 6-10 cm long; 2.5-3 cm wide near the middle, apex acute to slightly apiculate, base cuneate to slightly rounded, gland at base, pinnate venation obscure on either surface, Upper surface very finely velvety and deep green, lower surface lighter green, dull with numerous lighter punctations. Leaves rather thick, but not succulent, somewhat leathery. Petiole: short, thick, glabrous, 0.08 cm. long, curved, terete, 0.05 cm. in diameter, not grooved. Hoya plicata King and Gamble 1908. Leaves fleshy coriaceous when dry; ovate-lanceolate, acuminate at apex, acute at base; smooth and glabrous on both surfaces; margins slightly recurved; 3 in. long, 1 in. broad; midrib very faint and then only when dry; remaining nerves invisible; petiole stout, .25 in. long, flattened. Hoya polystachya Blume 1849. leaves broadly-ovate or oblong-ovate, acuminate, with base rounded or somewhat cordate; fleshy, thick 3-5 nerves, margins a little recurved. Hoya poolei White & Francis 1928. Leaf blades fleshy, lanceolate or elliptical, acuminate, venation indistinct in dried specimens, 6-9 cm. long, about thrice as long as broad. Hoya pottsii Traill 1830. The leaves are cordate, sharply, acuminate, having occasionally a rusty stairs partly spread over them, beneath of a pale whitish green without veins, above light yellowish green, with three distinct veins of ratter lighter colour than the leaf, from which smaller veins occasionally branch off. The petioles are not particularly thick, and less than half an inch long; they are of the same colour as the stems. Hoya praetorii Miquel 1857. leaves oblong or ovate-oblong or somewhat lanceolate acuminate, midrib on both sides with 5 outspread ascending veins prominent beneath, above somewhat grooved,6 to 3 inches long. Hoya pruinosa Miquel 1856. Leaves (small) somewhat round above convex granular- punctate, beneath flat and glaucous-pruinose. Hoya pseudolittoralis Norman 1937. Leaves somewhat fleshy in those collected, red-brown ovate acute without nerves, 7 times 3 to 9 by 3 to 4.5 cm, with petiole very short and narrow. Hoya pseudomaxima Koorders 1918. foliage is profoundly cordate and from Hoya maxima Koorders leaves above glabrous and at the apex not at all or rarely emarginate……………………. Leaves

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round-circular, at the base to 3 cm deeply-heart-shaped, on the top entirely glabrous (not only glabrous, but also without Cutiular-protruberances and polished, undersides glabrous and also without Cutiular-protruberances, to 8 cm long and 9 cm broad. Hoya pseudovalifolia Costantin 1912. Leaves ovate-acuminate, fleshy; 4.2 to 6 cm long and 2.4 to 3 cm wide; petioles up to 3 mm long. Hoya pubens Costantin 1912. Leaves not Isis fleshy as the preceding species (parasitica) oval-oblong, rounded or often pointed at base shortly apiculate at the apex; veins 2 lateral, in addition to pairs departing at the base and joining the others, reappearing near apex, very strong and prominent on both sides. Three rib-net-work very apparent; leaf 9.5 to 13 cm long by 3.6 to 6.8 cm. ; petiole very fleshy, thick, 2 cm long by 3 mm. Hoya puber Blume 1826. leaves (small) veinless oval-oblong acute fleshy puberulous. Hoya pubescens Reinecke 1893. leaves fleshy-coriaceous, flat, penninerved, oval or ovate, very shortly and abruptly acuminate, bases rounded, or somewhat cordate, petiole long, pubescent, blades on both sides shortly lanate. Hoya pubicalyx Merrill 1918. Leaves opposite, fleshy, when dry coriaceous, pale and shining on both surfaces and more or less wrinkled, oblong to oblong-obovate, 10 to 14 cm long, 3 to 6 cm wide, base obtuse, apex distinctly acuminate; lateral nerves pinnately arranged, indistinct, about 6 on each side of the midrib, ascending, scarcely more evident than are the lax, indistinct reticulations; petioles about 1 cm long. Hoya pubifera Elmer 1938. Leaves opposite, widely scattered, thick or leathery, curing grayish to yellowish brown, entire, the edges sometimes pressed with involute margins, narrowly oblong or oblanceolate to narrowly spathulate, velutinous beneath as well as densely or numerously papillose, the upper surface very similar but only slightly pubescent, the average blades 12 cm long by nearly 3 cm wide at or above the middle, the base attenuate, apex obtuse to subacute, a few of my detached leaves are only 6 cm long by 2 cm wide and with both ends obtusely rounded; midrib not conspicuous, lateral nerves none; petioles 1 to 3 cm long, dark yellowish brown, leaving circular scars upon falling. Peduncle stout, erect or nearly so, 4 cm long, similarly papillose and of a rich yellowish brown tomentum on my specimen. Hoya pulchella Schlechter 1913. loosely leaved, round, minutely pubescent. Leaves erect, spreading or outspread, petiolate, small, ovate, obtuse, slightly leathery, both sides glabrous, petiole minutely pubescent, above slightly grooved.

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Hoya purpurascens Teijsmann & Binnendijk 1863. leaves ovate, acuminate, base rounded, three nerved, above almost lacunose, reddish or purplish, margins revolute. Hoya purpureofusca Hooker 1850. Leaves opposite, on very thick, brownish petioles, four or five inches long, exactly ovate, acute, or shortly acuminate, thick, fleshy, five-nerved, the nerves all diverging from the base, and having a gland at the base where set on to the petiole. Hoya pusilla Rintz 1978. LEAVES very similar to those of H. lacunosa; fleshy, oblanceolate with acuminate bases and thickened margins; up to 8cm long by 3cm wide. PEDUNCLE reflexed, rigid, up to 5cm long. Hoya pusilliflora S. Moore 1916. Leaves drying pale greenish grey, nervation well marked on the upper side, less clear on the on the lower; petioles 2-3 mm. long, stoutish, channeled above. Hoya pycnophylla Rechinger 1908. Leaves thick leathery, margines (when dry) not at all or barely revolute, opaque conspicuously 5 nerved, nerves prominently visible on both sides, shortly petioled. Petioles 1 cm long. Hoya querinoensis Kloppenburg & Siar 2007. Leaf blades: elliptic acuminate, quintuplinerved, glabrous 14.9 – 21.2 cm. long x 4.9 – 7.6 cm. at the widest in the middle. Petiole: 2.1 – 3.0 cm. long curved, glabrous. Hoya quinquenervia Warburg 1904. coriaceous foliage, standing out, broadly ovate 5.6 cm. long by 4 cm. wide, apex shortly acuminate acute, base rounded, 5 nerved, the innermost about reaching the apex, reticulations subdistinct, edges revolute, equally green on each side. Petiole 1.5 cm. long, 0.3 cm. in diameter, glabrous. Petiole: curved to straight mostly 2-2.5 cm. long, round not grooved above and merging with the leaf base. On the new growth there is a fine puberulous indumentum not present on older growth. Hoya quisumbingii Kloppenburg 1992. Leaf Blade: 8.f1-12.3 ctn. long; 2.0 -2.3 cm. at the widest, near the middle, tapering to a narrow base and with a long drawn out sharp somewhat rigid apex, which in many instances is curved a little downward; with pinnate venation 6-8 pairs on either side of the midrib area, which is more evident above, but also prominent below and on the Side with a few stiff hairs along its straight length from the petiole to the apex. Hoya radicalis Tsiang et P. T. Li 1974. Leaves fleshy, spatulate or oblanceolate 10 to 21 cm long 3 to 4 cm wide, apexes shortly acuminate or acute, bases cuneate running into the petioles margins noticeably

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narrowing all the way down, nervation on both sides conspicuous, nerves on both sides about 10 or more reticulate veins, petiole, when dried, almost flattened to 3 to 6 cm long, with obsolete glands. Hoya ramosii Kloppenburg & Siar 2007. leaf blade is narrow, not elliptic-lanceolate. Hoya reticulata Costantin 1912. Leaves oblong, in corner or rarely a little rounded at the base, attenuate progressively from the third lower portion, very sharp points at the apex (acuminate of 56 mm.), glabrous, thin, transparent; secondary nerves 9-10 pairs, prominent, almost winged underneath, at about 40°, right, divided in regular dichotomies, coufluentes in arches of 4-5 mm. from the edges, very visible on the faces; visible tertiary and quaternary networks; limbe length of 11-18 cm. by 3-4.5 cm.; petiole length of 1.5-2.5 cm. on 2 mm. Hoya rasarioae Kloppenburg & Siar 2009.

Hoya recurvula Kloppenburg 2000. well leaved with pairs of lanceolate foliage, 8.0 - 12.0 cm. long by ca. 3.5 cm. wide held mostly upright along the stem; flat to slightly curved, apex acute. Petiole, terete, at first bronze-green becoming corky with age, terete, curved, no groove, about 5 mm. in diameter. Hoya reticulata Merrill 1920. Leaves fleshy when fresh, coriaceous when dry, ovate to oblong-ovate, shining, 8 to 12 cm long, 4 to 7 cm wide, the base broadly rounded, the apex slenderly and sharply acuminate, the epidermis on both surfaces coarsely reticulate-wrinkled when dry; petioles 1 to 2.5 cm long; lateral nerves about 4 on each side of the midrib, somewhat spreading. Hoya reticulata Schlechter 1913. Leaves spreading or outspread, shortly petioled, elliptic, somewhat obtuse, fleshy coriaceous, both sides glabrous, reticulately nerved below, petiole short, lightly grooved above. Hoya retusa Dalzell 1852. leaves shortly petioled, linear, three angled, apexes retuse, fleshy, pale…… Leaves 3-4 mm. long, 6-7 mm. wide, surface channeled above. Hoya retusa Warburg 1907. petioles 2 to 2.5 cm long, 2 mm thick, when young densely puberulous becoming glabrous, leaves leathery obovate 10 to 15 cm long, 7 to 10 cm wide bases rounded apexes retuse in sinus shortly apiculate, both sides glabrous, above on juvenile leaves pubescent, veins on both sides 5 to 6, spreading strait and long connected before reaching the margine indistinct, reticulations of nerves somewhat distinct.

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Hoya revolubilis Tsiang et P. T. Li 1974. Leaves with similar features to H. liangii Tsang which differs by the rounded exterior angle of the corona; in this species the leaves are thick fleshy elliptic-oblong 69 cm long 23 cm wide with the apex obtuse base wedge-shaped or rounded with margines revolute above shiny green and below papilose when dry pale yellowish, midrib above flattened and below prominent nerve with veinlets vanishing, petiole 1 1.5 cm long above on the petiole 2-3 glands. Hoya revoluta Wight ex Hooker 1883. Leaves 1 ½ - 3 in., midrib and nerves not visible, base acute; petiole 1/8-1/6 in., very thick. Hoya rhodostele Ridley 1923. Leaves thick; lanceolate acute, base cuneate: nerves invisible, 3 in. long, 1.5 in. wide; petioles thick .5 in. long. Peduncle axillary 4 in. long. Hoya rhodostemma Schlechter 1913. Leaves erect outspread, broadly elliptic, acuminate, base wedge shaped, leathery, both sides glabrous, above lucid, below opaque, petiole glabrous, above slightly grooved. Hoya ridleyi King & Gamble 1903. Leaves fleshy, coriaceous when dry, variable; ovate or ovate-oblong or oblong-lanceolate, acute at apex, rounded or narrowed or sometimes cuneate at base; both surfaces g1abrous; margins recurved; 3 to 5 in. long, 1.25 to 1.75 in. broad; midrib slender. Faint, as are the nerves generally, even when dry; main nerves about 5 to 6 pairs, the lowest pair starting from the base and curving along the margin, the other nearly at right angles, quickly branching and joining the marginal one; reticulations irregular netted; petiole very thick, .25 to .75 in. long, broad at the junction of the blade. Hoya rigida Kerr 1939. Leaves ovate or ovate-elliptic, bases rounded or broadly cuneate, apex long acutely acuminate, margins narrowly revolute, 12 to 14 cm. long, 4.5 to 6.3 cm. wide, when dry stiffly leathery, very hard, above black-brown, beneath dark, both sides opaque from the base 5-pli-nerved, nerves on both sides flat-convex, transverse nerves between prominent; petioles very thick, above flat-convex, 1.2 to 2 cm. long. Hoya rizaliana Kloppenburg 1991. The foliage is small essentially ovoid, glabrous and enervis with a short petiole…. Leaf Blade: 3.6 cm. x 1.2 cm., ovate tapering more toward the petiole, enervis, glabrous. Petiole: 1-1.1 cm. long, round, glabrous. Hoya rosea Schumann 1905. leaves shortly petiolate, lanceolate, acuminate, bases acute, dried leathery, rugose, both sides glabrous………..The petiole is 1 to 1.5 mm long; the leaf blade is 2.5 to 5 cm long and is 8 to 15 mm wide in the middle with veins that are clearly perceivable neither

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on the upper or under side; when dried it is leather yellow bordering on greenish or brownish. Hoya rotundifolia Siebold 1853. elliptic-round leaves, those above are elusively suggestively-plain stained. They are thick, succulent, approximately l0 centimeters broad and 12-13 long. Hoya rubida Schlechter 1901. loosely leaved; with leaves generally spreading, ovate or ovate-elliptic, shortly acuminate, both sides glabrous, texture thickly puberulous *,4-6 cm. long, 2-3 cm. wide below the middle, petiole glabrous above slightly grooved, about 1 cm long. Hoya rufolanata Ridley 1923. Leaves coriaceous; elliptic-ovate or obovate, acuminate at apex, cuneate or attenuate or rounded at base; upper surface shining, glabrous or slightly villous, pubescent on the midrib and main nerves; lower surface ferruginous-villous, especially on the nerves, dull; margins strongly recurred; 4 to 6 in. long, 2 to 3 in broad; midrib stout, impressed above, as are the 7 to 9 pairs of main nerves which, leaving the midrib at an angle of about 50°, are at first straight and then curve upwards to join a rather obscure intramarginal looped nerve, also impressed; secondary nerves few; reticulations transverse, irregular; petiole stout, .75 to 1 in. long, densely ferruginous-tomentose. Hoya rumphii Blume 1825. leaves oblong-lanceolate acuminate, with the base very obtuse glabrous, below flat and pale. Hoya rupicola K. Hill 1986. Leaves extremely fleshy, obscurely broadly pinnately veined, narrow-ovate to ovate, acute, basally tapered, pale gray-green, 14 cm long, 3.0-7.0 cm wide; petioles 1.02.0 cm long, 6-10 mm diem. Hoya ruscifolia Decaisne 1844. leaves ovate mucronate mucrone very acute somewhat sessile without veins very glabrous, peduncles short receptacle ovate reduced. Hoya salweenica Tsang & Li. 1974. . Leaves fleshy elliptic or nearly oblong, above opaque green below more pale in the dried state pale yellowish 1 to 12.5 cm. long 3.5 to 4.5 mm wide, apex very obtuse or nearly round, base rounded or broadly wedge shaped, nerves 6 to 8 pairs, small veins disappearing, an inconspicuous gland above on the petiole, petiole thick to 1 cm. long. Hoya samoensis Seemann 1866. leaves ovate-elliptic acuminate 5-tuplinerved, nerves prominent on both sides. Hoya sanae Bailey 1897.

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Leaves oblong to oblong-lanceolate, from 1 ½ to 3 ¼ in. long, thick, pale in colour, only showing a rather broad midrib, more or less covered on the under side with white hairs, often aciculate, petioles seldom exceeding ½ in. Hoya sarcophylla Ridley 1917. Leaves thick leathery lanceolate with the base obtuse, nerves invisible, midrib on the upper surface at the base obscure 8.5 cm. long 3 cm. wide, petioles thick 8 mm. long. Hoya schallertiae Burton 1982. (Not a hoya) Leaves sessile (or nearly so), in opposite pairs, bases cordate or truncate, apexes rounded, broader than long ( ¾" X 1" ± ), margins erose-wavy, convex above, and lumpy, puberulous, scabrous beneath. Leaves flattened and oppressed against tree trunk, covering numerous roots. Hoya schneei Schlechter 1921. loosely leaved, round, glabrous, leaves spreading or erect outspread, petiolate, elliptic, acuminate, with a wedge shaped base, leathery glabrous, above lucid. Hoya scortechinii King & Gamble 1903. Leaves thick, fleshy, drying coriaceous, ovate or ovate-lanceolate, acuminate at apex, rounded or cordate at base, both surfaces glabrous, lower minutely glandular; Margins recurved, 2 to 3.5 in. long, .75 to 1.25 in. broad; nerves only visible when dry; midrib thick; main nerves 5 to 6 pairs at an angle of 50° to 60° with the midrib, branching early and anastomosing near the margin; reticulations transverse; petiole stout. Hoya serpens Hooker 1883. very small suborbicular opaque papillose on both surfaces…….. Leaves ½ - ¾ in. diam., ovate or very broadly elliptic, base rounded or subcordate, nerves obscure; petiole very short, 1/16 - 1/10 in. Hoya shepherdii Short ex Hooker 1861. leaves 6 inches long and linear-lanceolate, curved downward, thickly-fleshy above dark green channeled, below more pale and semi-rounded, veins and petioles bent abruptly, petioles short round. Hoya siamaca Craib 1911. Leaves lanceolate, oblong-lanceolate or ovate lanceolate, apexes somewhat acute, bases obtuse, 3.7 to 9.5 cm. long, by 1.5 to 22 cm. wide, both sides glabrous, margins lightly recurved, when dried leathery, nerves entirely obscure; petioles 0.5 to 1.5 cm. long, glabrous. Hoya siariae Kloppenburg 2000. opposite lanceolate glabrous, deep green foliage ca. 14 cm. long x 3.5 cm. wide, tri-ply parallel veined, distinct on the upper surface less so below. Petioles heavy and long, 3.0 cm., diameter 0.09 cm. with deep groove on upper side.

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Hoya silvatica Tsiang et P. T. Li 1974. Leaves papery to fleshy elliptic or elliptic-oblong above almost glabrous and beneath pubescent, dense near to the nerves, 8 to 11 cm long by 2.5 to 4 cm wide, apexes shortly acuminate bases cuneate, costa below lightly elevate, lateral nerves 7 to 10 pairs with veins conspicuously reticulate 2 or 3 glands above petiole, petiole 1 to 3 cm long, pubescent. Hoya sipitangensis Kloppenburg & Wiberg 2004. Leaves opposite, green with a few silver spots….. Leaves and flowers larger than Hoya lacunosa Blume. Hoya sogeriensis Moore 1913. leaves relatively large elliptic or elliptic-oblong shortly acuminate apex obtuse bases obtuse 5-nerved fleshy-leathery glabrous setting upon moderately long thickened petioles, peduncles strong much shorter than the leaves. Hoya solaniflora Schlechter 1913. Leaves spreading or outspread, petiolate, lanceolate, acuminate, texture thinly coriaceous, both sides glabrous, petiole glabrous, slightly grooved above. Hoya sororia Schumann 1905. (1 cm. long) petioled leaves of 5 to 6 cm long and 1 cm wide. Hoya spartioides (Kuntz) Kloppenburg 2001. The reduced foliage (many times absent) and its prodigious production of green (chlorophyll) peduncles set this species apart as a fringe member of the genus. Hoya speciosa Decaisne 1844. leaves very broad ovate or ovate-elliptic with the base rounded apex shortly acuminate margins narrowly revolute both sides glabrous petiolate. Hoya stenophylla Schlechter 1913. . Leaves spread or outspread, briefly petioled, narrowly linear, base rounded, margins commonly recurved, leathery, both sides glabrous, with the petiole pubescent, short, above slightly grooved. Hoya stoneana Kloppenburg & Siar 2006. obtuse leaf bases not attenuata, and being pubescent not glabrous….. Foliage: linear lanceolate 5-8.5 cm. long and 1.5-2.5 cm. at the widest, waxy dark green, on both surfaces, lighter on underside. Dorsal surface with a few scattered hair cells and more along the edges, ventral surfaces covered with clear curved hair sells. Apex acute to apiculate base mostly obtuse. Petiole green colored pubescent mostly 1 cm. long and grooved above. Hoya subcalva Burkill 1901.

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Leaves ovate-elliptic, very shortly acuminate, bases rounded, 4 to 4.5 inches long 2 to 2.5 inches wide, glabrous, nerves on both sides moderately conspicuous; petioles 6 to 10 lines long. Hoya subglabra Schlechter 1913. Leaves spreading or outspread, petiolate, elliptic, acuminate, leathery textured, both sides glabrous, petiole glabrous, lightly grooved above. Hoya subquaterna Miquel 1856. leaves distinctly petiolate, opposite here and there almost shield shaped, rhomboid-ovate attenuate-acuminate, fleshy, veinless, petiole almost peltately inserted, 7-4 lines long. Hoya subquintuplinervis Miquel 1869. : Leaves elliptic-rounded apex here obsoletely somewhat apiculate, fleshy, glabrous, midrib almost 5-plinerved on both sides traversed 2 almost rising out of the base on the other side ½ of the second. Hoya suregaoensis Kloppenburg & Nyhuus 2011. A broad ovate leaf with a deep cordate base edges rolled under, apex apiculate. Nerves 5 triplinerved. Nerve angle 45˚. Hoya tauensis Kloppenburg & Siar 2008. Leaves glabrous, broadly ovate elliptic, edges undulate, glabrous, apex attenuate to apiculate acute, base obtuse, nerves 5 palmate visible on both surfaces central ones extend to near the apex, anastomosing, opposite; 6.7 - 9.7 cm. long 2.3 - 4.8 cm. widest near the middle. Petiole long medium heavy, glabrous, 1.6 - 1.8 cm. long ca 0.15 cm, in diameter, same color (buff) as the stem. Hoya tenggerensis Bakhuizen 1950. leaves ovate-oblong base rounded apex short acuminate glabrous 3 to 5 nerves, within a limited area, middle nerves with 2 basal precursor nerves laterals many, with thickness and thicker nerves similarly at the base, when dry manifestly thick leathery above, below veins not as manifestly coarse, above at the base of the midrib provided with thick kidney shaped cells ( lacking ? trachoma base) 10 to 12 long, petiole thick glabrous 1 cm long. Peduncles glabrous 1.5 to 4 cm long. Hoya teretifolia Griffith ex Hooker 1883. leaves 5-7 in. very slender cylindric quite terete. Hoya teysmanniana Miquel 1856. leaves at bases obtuse-rounded, somewhat obverse lanceolate-oblong (upwards a little wider) at the apex rounded or truncate or lightly emarginate, costa almost extended into a mucrone, thick leathery, veinless 53 inches long. Hoya thomsonii Hooker 1883.

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leaves beneath sparsely pilose, leaves 2-3 in. oblong or obovate-oblong acuminate very thick………… Leaves not very fleshy* , 2/3 - 1 ½ in. broad, midrib obscure, base obtuse or rounded, nerves very faint, arched; petiole ¼ - ½ in. Peduncle ½ -1 in., pubescent. Hoya tjadasmalangensis Bakhuizen 1950. leaves broadly ovate, base rounded, briefly acute and acuminate, glabrous. Not nerved above large warts when dry thickly leathery 19.5 cm long 22 cm wide. Hoya tjampeaensis Hochreutiner 1936. blades elliptic, apexes acuminate and acute, base attenuate acute, rarely a little rounded, margins entire, leathery but nevertheless nerves more or less prominent on both sides and conspicuous, middle nerves below very prominent, secondary nerves at the bottom small, oblique, meeting at the margins, over ¼ of blade long inconspicuous. Secondary nerves other than the principle about 6 to 7 on both sides, very irregular, more or less oblique or perpendicular and in this manner arching at an angle and noticeably irregularly anastomosing, i.e. more or less near margins or at a distance from the margins. Blade apparently glabrous, nevertheless very minute pilose very rarely adorned on both sides, mainly on the middle nerve. Hoya telosmoides Omlar 1996. Hoya tomataensis Green and Kloppenburg 2004. foliage is thick, lanceolate ovate, obscure nerved, base narrow…. opposite leaves and clear sap. Leaves are glossy deep green with a waxy surface, lanceolate-elliptic with a short apiculate apex, and hang essentially flat. The nerves are somewhat obscure but pinnate anastomosing the leaf base is rather long angustate. Typical leaf is 10.7 cm. long x 3.8 cm. at the widest near the center. The peduncles are curved, relatively long, terete and not grooved above 2.5 cm in length and 0.36 cm. in diameter. Hoya torricellensis Schlechter 1913. Leaves spreading or outspread, elliptic, acuminate, leathery, glabrous, petiole glabrous, slightly grooved. Hoya trinervis Trail 1837. leaves variable in size, oblong, sharply acuminate, pale green beneath, and mottled with small brown spots, light yellowish green above, with 3 prominent veins. Hoya treubiana Schlechter 1908. loosely leaved; with the leaves petiolate, oblong obtuse, glabrous, thickly leathery, nerves not at all conspicuous, with the margins revolute, 13-18 cm. long, in the middle nearly 4.5-5.5 cm. wide, petiole fleshy, 1-1.5 cm. long. Hoya trigonolobos Schlechter 1905. loosely leaved, producing aerial roots; leaves spreading ovate or ovate-oblong, acute or briefly acuminate, both sides glabrous, texture fleshy-leathery, about 14 cm long,

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in the middle or below the middle about 7 cm. wide, petiole thick, short, 0.5-0.7 cm. long; peduncle slender, round, glabrous about 6.5 cm. long. Hoya trinervia (Hort.) Mach. ex Regal 1865. Smaller 3—5 nerved, but not veined leaves,………….. Those of a short thick stalk support thick and leathery leaves, oval, before the pointed end, of 3 or 5 passed through under the long leaf apex anastomosing, the above side bright green, underside pale, glabrous throughout 3-4 inches long and 1 ½—2 inches broad. Hoya treubiana Schlechter 1908. loosely leaved; with the leaves petiolate, oblong obtuse, glabrous, thickly leathery, nerves not at all conspicuous, with the margins revolute, 13-18 cm. long, in the middle nearly 4.5-5.5 cm. wide, petiole fleshy, 1-1.5 cm. long. Hoya trigonolobos Schlechter 1905. loosely leaved, producing aerial roots; leaves spreading ovate or ovate-oblong, acute or briefly acuminate, both sides glabrous, texture fleshy-leathery, about 14 cm long, in the middle or below the middle about 7 cm. wide, petiole thick, short, 0.5-0.7 cm. long. Hoya trinervia (Hort.) Mach. ex Regal 1865. . Smaller 3—5 nerved, but not veined leaves,……………. thick and leathery leaves, oval, before the pointed end, of 3 or 5 passed through under the long leaf apex anastomosing, the above side bright green, underside pale, glabrous throughout 3-4 inches long and 1 ½—2 inches broad. Hoya trukensis Hosakawa 1937. Leaves outspread or erect-spreading, petioles about 1 cm long glabrous ovate, apexes acuminate or caudate-acuminate, bases rounded, papery, both sides glabrous, above lucid, 4 to 5.5 cm long by 2 to 2.5 cm wide. Hoya tsangii Burton ex Kloppenburg 1989. Leaves narrowly elliptic acute, 5-12 cm. long often 5 cm., near the middle 1-1.5 cm. wide, on both sides with out nerves; petiole short, glabrous 1 cm. long +/- above lightly grooved. Hoya tsiangiana P. T. L 1984. Same as nom. nov. Hoya retusa Warb. Hoya tsoi Merrill 1934. leaves oblong or narrowly oblong-elliptic, leathery, when alive fleshy, 6 to 8 cm long, 1.5 to 2 cm wide, pale, on both sides somewhat narrow, base acute, apex mostly acute acuminate, primary nerves on both sides 4 to 5, spreading, obscure, bending, anastomosing; petioles thick 8 to 15 mm long. Hoya uncinata Teijsmann & Binnendijk 1863.

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leaves acuminate, with the base rounded, now and then narrowed, acute, fleshy….. Leaves fleshy 0.09 long 0.035 wide, acuminate, with the base cuneate, above somewhat lacunose veined, below veinless, both sides with white dots…. Leaves (sometimes with the exception of the thinly pubescent midrib) glabrous or subglabrous on both sides. Hoya upoluensis Reinecke 1893. leaves leathery, when dry papery, briefly petiolate , often oblique, ellipticlanceolate, distantly acuminate, narrow based, definitely quintuple nerved, nerves prominent on both sides; inflorescence umbel-shaped, peduncles many times longer than the petiole; pedicels slender, shorter than the peduncles. Hoya vacciniiflora Schwartz 1931. Leaves oblong-lanceolate both ends acute very glabrous leathery when living probably thick, margins entire one-nerved, nerve impressed above, prominent below, veins throughout frequently uniting. Leaves shortly petioled petioles thick, channeled above, 0.5 to 1 cm long, blade 5.5- to 8.5 cm long and 1.51.7 cm wide. Hoya vaccinioides Hooker 1883. leaves ½ in. elliptic subacute fleshy nerves obsolete……….. leaves diverse when young ovate-orbiculatis, apex briefly acute, later elliptic lanceolate acute, base margine entire, other abbreviated, thick variegation. Hoya variegata De Vriese 1853. leaves white-variegated, margins often rosy fleshy, oval, elliptic oblong, somewhat acuminate, petioles reddish, 2 to 5 cm. long. Hoya variegata Siebold ex Morren 1846. leaves diverse, juvenile ones ovate-orbicular, apex short acute, becoming elliptic lanceolate acute, base margine one, the other abbreviated, thick variegated. Hoya variifolia Ridley 1926. leaves fleshy-leathery dimorphic elliptic obtuse, bases narrow, 1.5 cm. long, 5 mm. wide, and linear-lanceolate, both ends acuminate 4 cm. long, 2 mm. wide, petiole scarcely distinct; umbels 18-flowered; peduncles slender 2 cm. long. Hoya venusta Schlechter 1913. Leaves 3-5 cm. long, below the middle 0.9-1.7 cm. wide, petioles 0.2-.4 cm. long. Hoya verticillata (Vahl) G. Don 1837. leaves 4 in a whorl, petiolate, oblong, acuminated, acute, fleshy, glabrous, veinless. Hoya villosa Constantin 1912.

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Leaves oblong, elliptic, rounded at the basis and at the summit sometimes very slightly echancrees or very weekly mucronate, densely villose, yellow or rose colored underneath; a little villose on top; nerves four pairs of laterals, oblique, arched toward the edge, invisible underneath; limbe length of 7-11 cm. x 3.8-4. 8 cm.; petiole villous or glabrous, length of 1. 5-2 cm., 4 mm thick. Hoya vitellina Blume 1849. leaves oval oblong acute with base rounded or obtuse fleshy above veined, below no veins. Hoya vitellinoides Bakhuizen Brink f. 1950. leaves oblong base shortly obtuse and acutely acuminate on both surfaces very sparsely and very shortly pilose thickly coriaceous, penninerved, lateral nerves 4 to 6 parts on both surfaces conspicuously transverse all over by irregular veins without (?) trichomes 14 to 16 cm long, 7 to 7.5 cm wide, petiole thick 7.5 to 10 mm long. Hoya vitiensis Turrill 1915. Leaves elliptic-oblong or elliptic-ovate, with apex attenuate acuminate, base rounded or almost cordate, up to 10 cm. long and 7 cm. wide, entire, glabrous lateral nerves on both sides about 7 almost to the margins anastomosing when dry with the midrib flat prominent on both sides. Reticulations prominent in a transverse manner; petiole 0.5 to 1.5 cm. long, thick, glabrous. Hoya volubilis Griffith 1854. leaves ovate-lanceolate acute leathery glaucous at the base one callus , indistinctly 3 nerved. Hoya wallichiana Decaisne 1844. leaves broad ovate oval or obovate with rounded base or obsoletely emarginate apex short acuminate, with the limb arising from transverse veins transversely thickened and somewhat fleshy, below velvety with middle nerve very prominent glabrous, petiole scarcely 1 inch, thick. Hoya walliniana Kloppenburg & Nyhuus 2004. None Hoya warburgii Schlechter. Unpublished but herbarium sheet and drawing. Hoya wariana Schlechter 1913. Leaves spreading or somewhat outspread, shortly petioled, strap shaped, linear, acute or apiculate, base obtuse, thin leathery texture, both sides glabrous. Hoya wayetii Kloppenburg 1993. Leaf Blade: 5-14.5 cm. long x 1.5-2.7 cm. wide, thick, succulent, convex to nearly flat on the upper glabrous surface, triangular below and lighter green covered with

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granulose lighter colored punctations. Upper surface deep green with deeper green edging especially in younger leaves. Some rusty tones may be exhibited in newly formed leaves. Blades tapering to both ends, especially narrow toward the petiole which merges with the blade, leaf gland very small almost non-existent. Hoya waymaniae Kloppenburg 1995. Leaf Blade: thick, rigid, succulent, both surfaces finely pubescent especially so with the lower surface. 5-9 cm. long, margins very undulant 3-5 cm wide, enervis, edges rough with small bumpy thickened protrusions, much pinkish pigmentation. with a linear imbedded leaf gland present. Hoya weebella Kloppenburg 2000. Leaves: opposite, midrib prominent below, stem rusty, apparently no leaf gland, a few trichomes along the midrib, few hairs on lower midrib end of blade, upper surface rugose, base rounded (obtuse) tapering (lanceolate) to a narrowly rounded apex. 0.18 cm. long and widest near the base 0.09 cm. apex with small apiculation. Blade dark green underside glabrous, edges slightly recurved, thin. Above medium green, slight groove down the center above midrib, edges entire, below granulose punctate, glabrous. Petiole: curved, hirsute, hairs pointing toward the leaf blade, ca. 0.90 cm. long, clean 0.30 cm. in diameter.

Hoya whistlerii Kloppenburg 2002. opposite, thick lanceolate leaves. The longest leaves are 5.5 – 10.5 cm. long, mostly with acuminate apices. Hoya wibergiae Kloppenburg 2001. leaves opposite tend to be long narrowly elliptic curved along length apex acute to apiculate 6.5 cm. - 11.0 cm. long x ca. 4.2 cm wide, but some smaller and broader, triplinerved with faint anastomosing, only primary veins and midrib visible below surfaces glabrous, margins sharp and slightly curved under, blade not flat. Leaf upper surface with splotchy silvering, petiole terete, corky. Hoya wightiana Thwaites 1864. leaves glabrous, fleshy, linear-lanceolate, acuminate, obtuse, with narrow base. Hoya wrayi King & Gamble 1903. Leaves fleshy, thinly coriaceous when dry; lanceolate, obtuse and glandular-apiculate at apex, long-attenuate at base; upper surface glabrous smooth, lower minutely papillose, both grayish when dry; margins recurved; 3.5 to 4 in. long, 1 to 1.25 in. broad; midrib slender, broadened at base; main nerves very inconspicuous even when dry, about 4 pairs, at about 30° with the midrib; reticulations not visible; petiole very thick, rugose, jointed at the junction with the blade. Hoya yuennanensis Hand- Mazzetti 1936. Leaves almost circular continuously elliptic, 6 to 11 cm long, somewhat continuously longitudinal (rarely) doubly narrow.. briefly and obtusely apiculate, base broadly rounder truncate, very thick, flat, dry light green, both sides with twisted hairs, below when young almost velvety, below 5 to 7 nerves diverging at about 60° from the

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midrib straight to near the margins barely curved connecting loosely with the reticulate veins when dry prominent on both sides, at the base of the midrib is a yellowish gland; petioles long 3 to 7 mm, 2 to 3 mm wide densely hairy. Peduncles axillary, 2 to 3.5 cm long. Hoya zollingeriana Miquel 1857. Leaves with short petioles and bases obtuse, on the upper dorsal side of the petiole somewhat drawn out, elliptic or largely obovate-oblong, or obtuse-apiculate, thick leathery, beneath on both sides 4-5 obsolete veins (when dry, somewhat prominent) in pairs, 4-2.5 inches long.

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