Contents of Volume III Species

Page

Hoya coronaria Blume Hoya corona Ariadnes Blume Hoya costantinii P. T. Li Hoya crassicaulis Elmer ex Kloppenburg Hoya crassifolia Haworth Hoya crassifolia Ridley Hoya crassior Hochreutiner Hoya crassipes Turczaninow Hoya crocea Tuinbouw Hoya cumingiana Decaisne Hoya curtisii King & Gamble Hoya cystiantha Schlechter Hoya dalrympleana F. Muell Hoya darwinii Loher Hoya dasyantha Tsiang Hoya davidcummingii Kloppenburg Hoya dennisii P. I. Forster et D. J. Liddle Hoya densifolia Turczaninow Hoya dickasoniana P. T. Li. Hoya dictyoneura K. Schumann Hoya dimorpha F. M. Bailey Hoya diptera Seemann Hoya dischorensis Schlechter Hoya diversifolia Blume Hoya dodecatheiflora Fosberg Hoya dolichosparte Schlechter Hoya edanoi C. M. Burton Hoya edeni King ex Hooker Hoya eitapensis Schlechter Hoya elegans Kosteletsky Hoya elliptica Hooker Hoya elmeri Merrill Hoya elmeri Schlechter Hoya el-nidicus Kloppenburg Hoya endauensis Kiew Hoya engleriana Hosseus Hoya epedunculata Schlechter Hoya erythrina Rintz Hoya erythrostemma Kerr Hoya esculenta Tsiang Hoya esquirolii Leveille Hoya excavata Teijsmann & Binnendijk.

1 12 14 14 18 19 19 21 23 23 32 33 35 35 37 38 41 42 46 47 49 49 53 55 60 62 63 65 65 68 69 70 71 71 73 74 78 80 81 82 83 83

Hoya exilis Schlechter Hoya filiformis Rechinger Hoya finlaysonii Wight Hoya fischeriana Warburg Hoya flagellata Kerr Hoya flavescens Schlechter Hoya flavida P.I. Forster et D. J. Liddle Hoya flexuosa Spreng Hoya forbesii King & Gamble Hoya formosana Yamazaki Hoya fraterna Blume Hoya fungii Merrill Hoya fusca Wallich Hoya fuscomarginata N. E. Brown Hoya gigantanganensis Kloppenburg Hoya gigas Schlechter Hoya glabra Schlechter Hoya globiflora Ridley Hoya globosa LeConte Hoya globulifera Blume Hoya globulosa Hooker Hoya golamcoiana Kloppenburg Hoya gonoloboides Regel Hoya gracilipes Schlechter Hoya gracilis Schlechter Hoya grandiflora Sprengel Hoya grandiflora Blume ex Decaisne Hoya graveolens Kerr Hoya greenii Kloppenburg Hoya griffithiana Decaisne Hoya griffithii Hooker Hoya guppyi Oliver Hoya gymnanthera Wight Hoya hainanensis Merrill Hoya halconensis Kloppenburg Hoya halophila Schlechter Hoya hasseltii Miquel Hoya hellwigiana Warburg Hoya hellwigii Warburg Hoya heuschkeliana Kloppenburg Hoya hirsuta Moon Hoya hollrungii Warburg Hoya hookeriana Wight Hoya hypolasia Schlechter Hoya imbricata Decaisne Hoya imperialis Lindley

85 87 88 91 92 94 95 98 98 99 100 107 109 113 114 117 119 121 121 121 122 125 126 127 129 133 134 134 135 138 138 140 141 142 143 144 148 149 151 152 153 153 156 157 159 169

Hoya Literature Volume III Hoya coronaria Blume In Bijdragen tot de Flora van Nederlandsche Inde (1825) 1063. C. L. Blume. Hoya coronaria, Bl. H. foliis aveniis ovalibus acutis margine recurvis coriaceis infra villosis, corollis glabris (Corona Ariadna, Rumph. Herb. Amb. V. t. 172). (1064). Crescit: in montanis et ad ripas circa Rompien. Floret: per totum annum. Nomen: aroy Kilampahan. Translation: leaves veinless oval acute with margins recurved, leathery, beneath villose, corolla glabrous. (Corona Ariadna in Rumphia Herbarium Amboinensis 5. t 172). 1064. Growing: in the mountains and frequently on the stream banks around Rompien. Flowers the entire year. Local name “Aroy Kilampahan”. In Allgem Med-Pharm. Fl. 3 (1834) 1063. Kosteletsky. H. coronaria Blum. Groszblumige. H. Blätter eiförmig-oval, spitz, lederig, am Rande umgebogen, unterseits zottig und aderlos; Corollen kahl. - Rumph. 5.t.172. Schlingstrauch mit einen sehr langen Stengel und wenigen, kurzen Aesten. Blätter in groszen Zwischenräumen, auf kurzen, dicken Stielen stehend 3 – 4” long, 2” breit, dick und dicht, aber brüchig, am Grunde abgerundet ober etwas ausgerandet, vorn stumpf oder spisztich, oben kahl und glänzend, unterseits graulich und nur der Mittelnerve vorstehend. Dolden auf 2” langen, dicken Stielen, 6-10 blütig. Blüthen sehr grosz, 2” breit; auf 1” langen Stielchen, aussen feegrün, innen stark glänzend-braun, Abends und in der Nacht angenehm und fast ananasartig riechend. Corolle flach oder etwas zurück geschlagen; zipfel eiförmig, spitz. Balgkapsel grosz, 13-14” lang, at the base from the thick pulp finger and da an allmächlig verschmälert, blaszgrün, gerillt. - In Berggegenden und an Bächen in Java und auf den Melukken. Blüht fast stets (Symbol) - Die Blätter enthalten so, wie alle übrigen Theile, viel von einer dicken, etwas kletrigen, ober nicht scharfen Mitch und werden auf den Molukken als ein kühlendes Mirtel bei Gonorrhöen angewendet; auch bedient man sich dieser Milch äuszerlich bei solchen Wunden, welche von den spitzigen Stacheln einiger giftigen Fische herrübren, - Rumph beschreibt noch eine zweite Art (H. lutea *) bie der vorhen gehenden sehr ähnlich sehr soll, ober breitere, mehr oval - rundliche, blaszgrüne, unterseits aberige Blätter, eben so grosze, jedoch gelbe Blumen und nur 9-10” lange, auch dünnere, and der Spitze, etwas gekrümmte Balgkapfeln besitzt, übris gens aber bieselbe Anwenbung findet. Translation: Large-bloomer. Hoya. Leaves oval-oval, pointed, leathery, revolute at the edge, unterseits shaggy and but-free; Corolla glabrous. - Rumph. 5.t.172. Climber-shrub with a very long stem and few, short branches. Leaves intermediate in size, short flat stalks, 3 to 4 inch long, 2 inch wide, thick and dense, but brittle, at the base rounded upper some upright, in front dull or a little pointed, above glabrous and shiny, on the underside grayish and only that median-nerve prominent. Corymb on 2 inch long, fat peduncles, 6-10 flowers. flowers very large, 2 inches wide; on 1 inch long pedicels, outside sea-green, inside strongly shiny-brown, evening and in the night pleasantly and 3

satisfying pineapple smell. Corolla flat above, sometimes bent backwards; corners revolute, pointed. Seed capsule 13-14 inches long, at the base of the thickness pulpy finger and there it gradually narrows below, grooved. - In the mountains regions and at brooks in Java and on the Melukken. - Virtually always blooms, shrub – the leaves contain so, like all remaining parts, many of a fat, some climbing, otherwise not sharp milk and is applied in the Moluccas as a chilling Mirtel for gonorrhea; also one uses this milk externally with such wounds, which originates as a result of the pointed thorns of some poisonous fish, - Rumph describes in addition a second type (H. lutea *) with the previous going very similarly very positive, otherwise wider, more oval - roundish, pale– green, once again the underside of the leaf, even so large, pale yellow flowers, and only 9-10 inches long, also brownish, and of the top, possesses somewhat bent seed capsule, but otherwise seldom found attached. In General System of Gardening and Botany (1837) 127. G. Don. 24. H. coronaria (Blum. bijdr. p. 1063) leaves veinless; oval, acute, with recurved edges, coriaceous, villous beneath; corolla glabrous. Woody perennial shrub. Native of Java, on mountains and banks about Rompien, where it is called Aroy Kilampahan. Coronaria Ariadna, Rumph. amb. 5. t. 172. Garland Hoya. Fl. year. Shrub tw. In Synopsis Plantarum 6 (1840) 892. D. N. F. Dietrich. 21. H. coronaria Blume: fol. aveniis ovalibus acutis margine recurvis coriaceis infra villosis; corollis glabris. Rumph. 5. t. 172. In Java, woody shrub. Translation: leaves without veins, oval acute, margines recurved leathery below velvety; corolla glabrous. In Rumphia 4 (1848) 51. C. L. Blume. H. coronaria Tab. 182. fig. 2, Tab. 185 (analysis) et 184 (fructus): caule scandente; foliis ellipticis v. oblongo-ellipticis breviter acuminatis basi rotundatis marginibus recurvis subcarnosis supra obsolete venulosis glabratis subtus velutinis; racemis umbelliformibus multifloris; corollae magnae laciniis triangularibus extrosum pubescentibus intus glabris; coronae staminae foliolis angulo exteriore rotundatis Bl. Bijdr. Flor. Nrd. Ind. p. 1065 (excl. Syn. Herb. Amb.!). - Hoya velutina Wight Contr. bot. of India p. 53. De Caisne in De Cand. Prod. Syst. Veg. VIII. p. 635. 5. H. grandiflora Bl. mss. et De Caisne l.c. 2. - Aroy kilampahan Sundaiee. - Habit. Frequens in Javae occidentalis sylvis humilioribus et ad ripas umbrosas. latissime per Asiam tropicam dispersa, quippe quae et in Bengalia inveniatur. Succus lacteus aliquantum acer, qui omnibus partibus stripis continetur, facile nauseam excitat. Foliis tamen crudis Capsico (Tjabeh) et sale mixto subinde vescuntur montani Javae, ut digestionem infirmam et nimis tardam stimulent. - Suffrutex caule tereti, carassitie pennae anserinae, tanaci, superne pubescente, circum arbores crassissimas serpente. Folia remote opposita, petiolis ½ - 1 poll. longis teretibus intedum tortis subpubescentibus instructa, 3 - 5 poll., 4-plus 2 poll. lata, apice breviter abrupte acuminata, basi rotundata, marginibus subrecurvis, firma, supra nitidiuscula, transverse venulosa (interdum plane avenia) ac pube rara appressa obsita, subtus flavo-virentia nervo medio valde elevato et dense velutina, juniora utrinque dense pubescentia. Racemi umbelliformibus axillares v. extraaxillares, solitarii, multiflori, pedunculo suffalto petiolo paulo longiore, pube densa plane tecti. Calycis foliola ovata- rotundata, corollae appressa, velutina, accrescenti-

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persistentia. Corolla 1½ poll. diametro, carnosa, externe pube sericca e virescenti incarnata, intus glabra, eburnea punctisque purpureis aspersa; laciniis triangularibus, breviter acuminatis, convexis, marginibus subrevolutis. Coronae stamineae eburnea foliola horizontalia, supra depress cum umbone, subtus alveolis quibusdam exsculpta, angulo exteriore rotundata, interiore ex umbone in dentem obtusiusculum stigmati incumbentem et intus membranae antherarum membranaceae connatum continuata. Retinacula emarginatis stigmati inter dentes, quos modo dixi, innexa, cuneata, ima bifida, extus sulco longitudinali inscripta, atro-brunnea, cornea, nitida, latere utrinque in cornu horizontaliter divergens explanata eujus apici recurvo pollinia elavata ope pedicelli brevis elevati arcuato-incurvi membranacei ad basin affixa. Ovarium sub corona staminea reconditum, e carpellis duobus oppositis cylindriceis ad basin annulo carnoso brevissimo cinctis uniloculatis vulgo aborientibus. Styli 2, brevissimi, in stigma commune magnum discoideum medio subapiculatum conjuncti. Folliculus cucumerinus, 8-10 poll., cylindricus, aliquantum curvatus, sulco longitudinali exaratus. ad basim latiorem foliolis calycis adnatis obsessus, obtusissimus retusus, glaucescens, sub lente visus pube rara obsitus, aciculatus, intus laevigatus, unilocularis, exsuccus, textura crassa spongiosa, sulco longitudinali tandem dehiscens. Semina plurima, spermophorum laterale convexum spongiosum undique obtegenta, deorsum imbricata, ovato-oblonga v. subrhomboidalia, compressa, hinc convexiuscula, inde concaviuscula, extremitate superiore fasciculo capillorum simplicium seicco-albicantium longorum comata. Translation: 5. H. coronaria Tab. 182 fig. 2, Tab. 183 (analysis) and 184. (fruit); stem climbing; leaves elliptic or oblong-elliptic briefly acuminate, base rounded margins recurved somewhat fleshy, above without veins, glabrous, below velvety; with multiflowered umbel-shaped racemes; corolla large, lobes triangular outside pubescent, inside glabrous; leaflets of the staminal corona exterior angle rounded, Bl. Bijdr. Flora van Nederlandsch Indie Batavae p. 1065 (exclude Synonym in Herbarium Amboinensis !). - Blume manuscript and DeCaisne quoted above 2. - Aroy kilampahan Sundanese Habitat. Frequents Western Java in the humid forests and on the shaded stream banks , dispersed widely in tropical Asia, and which to some extent found in Bengal. A bitter sappy latex in some quantity which easily excites nausea. I was informed nevertheless that the Tjabeh, in the mountains of Java mix the leaves with Capsicum and salt, immediately after to aid digestion and is an excessively slow stimulant. Slightly woody round, densely covered with a fine down, above pubescent, creeping, very thick somewhat tree-like. Leaves far apart opposite. Petioles ½ - 1 in. long, round, somewhat twisted somewhat pubescent, 3 - 5 inches, 1-2 or more inches wide, apex briefly and abruptly acuminate, base rounded, margines somewhat recurved, firm above very shiny, transverse veins (between clearly veinless) and scattered appressed pubescence, below yellow-green, center nerve decidedly elevated and with dense velvet, when young both sides densely pubescent. Umbel shaped raceme, axillary or extraaxillary, solitary, many flowered, peduncle long supported by the petiole, densely pubescent distinctly concealed. Calyx leaflets ovate-rounded, appressed to the corolla, velvety, more or less persistent, appressed to the velvety corolla. Corolla 1½ in. in diameter, fleshy, outside pubescent silky and becoming greenish flesh-colored, inside glabrous, ivory-white scattered dots of purple; leaflets triangular, briefly acuminate convex, margines somewhat revolute, leaflets of the staminal corona ivory-white, horizontal, depressed above, with a umbo, below a cavity sculptured out, exterior angle round, interior with

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rounded tooth like projections, somewhat obtuse laying on the stigma and fused on the inside to the anther membrane continuously. Retinacula on the Stigma margins between the teeth, quos modo dixi, innexa, wedge shaped, appearing bifid, outside with a longitudinal furrow, dark brown, horny, shiny, both sides horizontally flattened and horny, divergent, pollinia affixed at the base whose apex is recurved elevated by means of a short club shaped erect incurved pedicel membrane. Ovaries situated below the staminal crown with 2 opposed cylindrical carpels, at the base a very small fleshy ring encircling the common ovaries. Stigma paired, small connected to the very large discoid shaped staminal crown. Follicles cucumber shaped, 8 - 10 inches long cylindrical,, somewhat curved, with longitudinal furrow with the calyx adnate to the base occupied with broad lobes obtuse, rounded, glaucose, below I have seen it covered with a sparse pubescence, obtuse marked with fine irregular streaks, inside smooth and polished, one cavity, juiceless, thick and spongy textured, finally splitting longitudinally along the groove. Seeds with plumes, seeds laterally convex spongy on all sides covered ornately imbricate, ovate-oblong or sub rhomboid compressed after this very convex, later very concave, upper extremity with a bundle of thread-like simple long straight long white silky-tuft of hairs. In Museum Botanicum Lugduno-Batavum 1 (1849) 45. C. L. Blume. 100 Hoya coronaria Bl.: caule scandente; foliis; subcarnosis ellipticis v. oblongi-ellipticis breviter acuminatis basi rotundatis marginibus recurvis supra obsolete venulosis glabratis subtus velutinis; corollae magnae laciniis triangularibus extrorsum pubescentibus intus glabris; coronae stamineae foliolis angulo exteriore rotundatis. Bl. Bijdr. Flor. Ned. Ind. p. 1063 (excl. Syn. Herb. Amb.!). H. velutina Wight. Contr. bot. of Ind. p. 35. Decaisne in De Cand. Prod. Syst. Veg. VIII. p. 635.5. - H. grandiflora Bl. Mss. et Decaisne l.c. 2. - Aroy kilampahan Sundaice. - Frequens in Java occidentalis sylvis humilioribus et ad ripas umbrosas et latissime per Asiam tropicam dispersa. Translation: stalks climbing, leaves somewhat fleshy elliptic or oblong-elliptic briefly acuminate with base rounded margines recurved veins above invisible glabrous below velvety; corolla large leaflets triangular outside pubescent inside glabrous; leaflets of the staminal corona exterior angle rounded. ........... Frequents Western Java in the humid forests and on shady river banks widely dispersed in tropical Asia. In Annales Botanices Systematicae 3 (1852) 64. W. G. Walpers. H. coronaria Blume (bijdr, 1063), Rumphia IV tab. 182. fig. 2. & tab. 184. (H. grandiflora Dcne. in DC. l.c. no. 2 - H. velutina Wight. - Dcne. in DC. l.c. no. 5. In Flora van Nederlandsch Indie Bataviae 1 (1856) 517. Miquel. 3. Hoya coronaria Bl. Scandens, folia e basi rotundata elliptica vel oblong-elliptica breviter acuminata subcarnosa, subtus velutina, 5-4 poll. longa. corollae magnae laciniae triangulares extus pubescentes intus glabrae, coronae stamineae phylla extrosum rotundata. Hoya grandiflora Bl. mss. Decaisn. in Dc. Prod. VIII. p.635. Hoya velutina Wight Contr. p. 35. Decaisn. l. c. - Hoya coronaria Bl. Bijdr. p. 1063, excl. syn. Herb. Amb. - Rumphia IV p. 31, tab. 182, fig. 2 tab. 104. In spec. Sumatr. corona stam. basi ima hirtella, qua de re tacent auctores.

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In Zuid-Azië niet zeldzaam. - Voor-Indie, Java, op belommerde plaatsen. Sumatra, bij Siboga (Teysm.) - Aroi Kilampahan sund. of Teboel Tiboel. Translation: climbing, leaves with base rounded elliptic or oblong-elliptic briefly acuminate, somewhat fleshy, below velvety, 5-4 inches long, corolla large, leaflets triangular, outside pubescent inside glabrous, leaflets of the staminal corona outside rounded, ………... etc. Corona staminal base hairy, from which the authors retained. In Botanical Magazine (1857) t. 4969. Curtis’s. Hoya coronaria, Bl.; caule scandente, ramis robustis, petiolis pedunculis calycibus extus foliisque subtus pills curvatis magis minusve velutinis, foliis sublonge petiolatis crasse carnosis apiculatis basi obtusis obsolete transversim venosis marginibus subrecurvis, umbellis pedunculatis multifloris, corollae majusculae laciniis triangularibus extus pubescentibus intus glabris, coronae stamineae foliolis angulo exteriore rotundatis obtusis. Hoya coronaria. Bl. Bijdr. Fl. Ned. Ind. p. 1063 (excl. syn. Herb. Amb.), Rumphia, vol. 4. p. 31. t. 183.f. 2, et t. 184.f. 2. Hoya grandiflora. Bl. MSS. et Decaisne in De Cand. Prodr. v. 8. p. 635. Hoya velutina. Wight, Contr. Bot,. of India, p. 35. Wall. Cat. n. 8150. Aroy Kilampahan, Sundaice. This fine Hoya was introduced to our stoves by the Messrs. Veitch, of the Exeter and Chelsea Exotic Nurseries, from Java through their collector, Mr. Thomas Lobb, and it proves to be the Hoya coronaria figured and described by Blume, l. c. It is next to Hoya imperialis, Lindl., undoubtedly the largest-flowered species of the genus yet cultivated in our gardens. The color of the corolla, though exhibiting no bright or gay tint, has nothing of the lurid character of the species last mentioned; it is pale sulphur-yellow, prettily relieved by five red spots immediately around and at the base of the corona stamineae. Blume appears to have been the first to discover this plant in moist woods and shady banks of Western Java, and was then disposed to refer it to the H. Corona Ariadnes punicea" of Rumphius, under the name of H. grandiflora, in which Decaisne has followed him in De Cand. Prodr.; but Blume afterwards corrected himself, and gave of this species a good figure and description under the name we here adopt It is not however peculiar to Java: Dr. Wallich detected it in Silhet. The milky juice of the leaves, Blume observes, is acrid, and excites nausea; nevertheless the mountain tribes eat the plant uncooked, mixed with capsicum and salt, and consider that it assists digestion. Our drawing was made from Messrs. Veitch's plant in November, 1856. Descr. Somewhat woody climber; branches rather thick, terete, downy, green. leaves opposite, distant, petiolate, elliptical, apiculate, thick, fleshy, pale-green, three to five inches long, obtuse at the base, the margin a little recurved, penninerved; nerves obscure, nearly horizontal: above glabrous, the costa on the upper side, and the whole leaf beneath, as well as the thick, terete petiole, about an inch long, are subvelutinous, the hairs (and wherever they exist on the plant) curved. Peduncle arising from between the pairs of leaves, an inch or more long, downy, bearing, an umbel of many flowers: pedicels (which are longer than the peduncle) downy. Calyx of five, downy, ovate, rather acute lobes Corolla rotate: the lobes triangular, acute, moderately carnose, pale sulphur-yellow, with five small red spots at the base of the tube. Staminal crown large, conspicuous, of five blunt lobes or rays. Blume describes the fruit as “follicles

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cucumerinus, 8 -10 poll., cylindricus, aliquantum curvatis, sulco longitudinali exaratus, textura crassa spongiosa, sulco longitudinali tandem dehiscens. Translation of above first section: stems climbing, branches robust. Petioles peduncles and calyx on the outside minutely velvety hairs and leaves below hairy, leaves somewhat long petioles thick fleshy apiculate base obtuse transverse veins not visible, margins somewhat recurved, umbels pedunculate of many flowers, leaflets of the corolla large triangular outside pubescent inside glabrous, leaflets of the staminal corona with the exterior angle rounded obtuse…………………………. Translation: Follicles cucumber shaped, 8-10 inches. cylindrical, somewhat curved, grooved longitudinally with a furrow, - texture thick spongy, eventually dehiscing along the longitudinal groove. In The Cottage Gardener (1857) 417. New and Rare Plants. Hoya coronaria (Garland Hoya). “This fine hoya” is a native of Java , whence it was sent to Messers. Veitch by their collector, Mr. Thomas Lobb. Its flowers are pale sulphur yellow colorued. Dr. Wallich found it in Sylhet and called it , as had Mr. Wight, Hoya velutina. It bloomed in November, 1856, at Messers Veitch’s. In Java it is called Aroy Kilampahan, - (Botanical Magazine, t. 4069.) In Flora of British India 4 (1883) 58. J. D. Hooker. 26. H. coronaria Blume bijdr, 1063, and Rumph. iv.31, t. 182 f. 2 and t. 104; leaves 3-4 in. elliptic or elliptic-ovate sharply acuminate very thick pubescent beneath, base rounded, margines recurved, peduncle stout and pedicels large, sepals pubescent, corolla coriaceous pubescent without glabrous within, follicles very thick. Miquel Fl. Ind. Bat. ii 616. H. velutina, Wight Contrib. 35; Wall. Cat. 8150. Dcne. in DC Prodr. viii 635. Malacca, Griffith, Maingay (Kew Distrib. 1125), Penang ? Wallich. - Distrib. Sumatra, Borneo. Stem stout, twining; young shoots, petioles, leaves beneath and inflorescence finely tomentose. Leaves 1-3 in. diam., nerves very obscure; petiole thick, ¼ - ½ in. Peduncles ½ - 1 in., pedicels as long. Sepals rounded, thick. Corolla 1 - 1½ in diam., thick, “cream white or yellowish speckled with purple,” Maingay; lobes erect or incurved. Coronal-processes small, waxy, white, nearly hemispherical, very concave above and convex beneath, the inner angle shortly horizontally produced, obtuse. Follicles 6-8 in. long by 1½-2 in. diam., subcylindric, obtuse, base gibbous, walls very thick, endocarp hard. Seeds ¼ in. long. The smaller shorter leaves with revolute margines distinguish this at once as H. obtusifolia; Wallich’s habitat of Silhet is probably in error for Penang from which island there is a drawing of this species in Herb. Kew. Perhaps two species mixed here, a purple and white-flowered. In Dictionary of the Royal Horticultural Society 2 (1884) 155. George Nicholson. H. coronaria (crowned). fl. yellow. November. l. oval, acute, with recurved edges, coriaceous, villous beneath. Java 1856 (B. M. 4969.) In Transactions of the Linnean Society of Botany 3 (1888) 321. H. Ridley. H. coronaria, Blume. Kwala Pahang, Kwantan. This is the commonest species of Hoya in the Straits. It very rarely produces fruit, but I have found the capsule twice in the same place, viz. at

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Pualau Tekong, between Singapore and Johore. Sir J. Hooker suggests, from the different colors of the flowers as described by Maingay and others, that there may be two species mixed here, a purple and a white-flowered one; but the diversity of color as described is only the effect of differences of age of the flower. The blossoms, when they first open, are greenish white; they then become of an ivory-white, with somewhat creamy tint; a little later irregular madder-pink spots appear here and there, after which the whole flower becomes dark pink and then commences to fade or falls off. In Queensland Agricultural Journal 3 (1898) 156-157. F. M. Bailey. Hoya coronaria var. papuana (n. var.) A robust tall twiner. Leaves oblong-ovate, thick-coriaceous, 4 to 5 in. long, slightly petiolate, the apex with a rather long point, 2 to 2½ in. broad at the centre, upper side glossy, the underside clothed with a close, short, stellate tomentum of a dirty-white colour; lateral nerves distant, nearly horizontal, only visible on the upper side, the gland indicating the attachment of the petioles, beneath scarcely visible. Petiole about 1 in. long. Flowers in almost sessile short racemes; the peduncle thick. stellate-tomentose, about 5 or 6 flowers open at one time. Pedicels tomentose, about ½ in. long. Flower-bud a thick 5-raayed star, the rays recurved, expanded flower about 1¼ in. in diameter, at first somewhat greenish, ultimately almost ivory, white or pale-yellow. Calyx-segments very obtuse, 2 lines long, tomentose. Corolla-tube swelled, very short, the inside lined with simple silky hairs; lobes broad with strongly reflexed margines, glabrous and glossy above, stellate-tomentose beneath. Corona-segments colour of the corolla, about 1 line in diameter, 2-ribbed on the underside. The flowers seem very near H. coronaria, Blume, but without the scattered purple dots on the flowers of that species. Hab.: Twining up trees on the margines of a small bay, foot of Mount Trafalgar, New Guinea. In Flora of the Malayan Peninsula (1903) 567. King & Gamble. 7. Hoya coronaria Blume Bijd. 1063 (1825), and Rumph. IV. 31. t. 182, fig. 2 and 184. (fruit). A stout twining shrub; the upper branches giving out roots on to the bark of trees and probably therefore more or less epiphytic; branches thick, velvety pubescent, the innovations especially so. Leaves thickly coriaceous; elliptic; elliptic-ovate or ovate-oblong, shortly acuminate at apex, rounded at base; upper surface when dry shining, wrinkled, puberulous especially on the midrib; lower surface pale brown, velvety pubescent; margins strongly recurved; 3 to 4 in. long, 1 to 2 in. broad; midrib stout, impressed above, raised beneath; main nerves very obscure, about 5 to 7 pairs nearly horizontal; reticulations not visible; petiole thick, velvety; .25 to .75 in. long. Umbels racemose, rather few-flowered, on thick rachises with large rounded .5 to 1.5 in. long at the ends of thick lateral velvety peduncles .5 to .75 in. long; pedicels velvety, curved, thick .5 to 1 in. long; buds depressed at top, the lobes decurved in a hook; flowers large, fleshy, about 1.5 in. in: diam., greenish-white changing to ivory-white, then apparently to pink which darkens and becomes speckled with purple (Ridley); the corona white. Calyx thick, becoming velvety without; lobes orbicular, .15 to .25 in. long; scales subulate. corolla very thick, glabrous within, villous without, lobes triangular acuminate, cordate at base .4 in. long. Corona of 5 spreading processes, horny when dry; the lower lobe nearly hemispherical, concave above, convex beneath, about .25 in. long; the upper acuminate, curved far over the anthers. Staminal-column short, thick, inserted at the base of the corona-tube; anthers incurved, their cells parallel above, divaricate and empty below;

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appendages prominent, scarious, ovate-acute; pollen-masses brown broadly clavate, attached by means of long slender twisted caudicles to the rather large pollen-carriers which are rounded above and bifid below. Style-apex circular, saucer-shaped with a small central umbo and 5 slender rays. Follicle l, 6 to l0 in. long, 1.5 to 2 in. in diam., gibbous at base, subcylindric above,-ending in an obtuse point; pericarp very thick, puberulous striate; the endocarp woody and smooth; the follicle at length opening nearly flat. Seeds obovate, .4 in. long, truncate at tip and bearing a copious white silky como 2 to 2.5 in. long; cotyledons elliptic, .2 in. long, fleshy; radicle cylindric, .1 in. long. Miq. Fl. Ind. Bat. II. 517; Hook. f. Fl. Br. Ind. IV. 58; Ridley in Trans Linn. Soc. and Ser. III. 321. Hoya grandiflora, Bl. MSS.; Decne. in Prodr. VIII. 635. H. velutina, Wight Contrib. 35; Wall. Cat. 8150; Dcen. l. c. Penang: ? . Wallich 8150 (marked “Si1let.” but probably in error); Curtis 236. Dindings: Curtis; Scortechini 1053. Perak: at Goping, Scortechini 2009; at Matang, on sea coast, Wray 2710; at Larut, Kunstler 3865 (leaves only); King’s Collector 4116, 6954. Malacca: Griffith 3788; Maingay (K.D.) 1125. Pahang: at Kwala Pahang, Ridley. Singapore: at Serangoon, Ridley 2731a. - Distrib. Java, Sumatra (Teysmann 1171), Borneo (Motley 871, Burbidge), Celebes, New Guinea (H.O. Forbes 516, flowers 2 in. in diam.). In Systematisches Verzeichnis Family 1 (1911) 6. Koorders. Hoya coronaria Bl. Kletterstrauch, bis 15 m. hoch. West-Java: Batavia: Depok: Region I: Kds 31172  (1141 - 14 XI 1898). Mittel-Java: Banjumas: Nusakambangan: Region I: Kds. 22126  (10. 23. XII. 1895 Ojot kapal. Jav. Auch Alcoholmaterial gesammelt). Kds 26847  (767 ‘ 25. I. 1897. In Exkursionsflora von Java (1912) 97-98. “Flora of Java” S. H. Koorders. (Key) 5b.Korolla mittelmaszig oder sehr grosz (sets über 1 cm im Durchm. haltend ................6 6a. Blätter nicht über 3 cm lang .............................................................................…....7 6b. Blätter 10-15 cm lang, elliptisch oder oblong-elliptisch, am Grunde abgerundet, oben kurz zugespitzt, ± fleschig, unterseits sammethaartig. Blattstiel 1 ½ -2½ cm. Korolle 2½ 3 cm breit, mit auszen flaumhaarigen, innen kahlen Zipfeln, Coronazipfel auszen abgebrundet, Follikel 22-25 x 6-8 cm, dickzylindrisch, ± gedunsen, gerade, oben ± stumpf. Blume, Bijdr., 1063 (exkl. syn.); Miq; l.c. 517; Blume, Rumphia IV. 31 tab. 182, fig.1 und 2. West und Mittel-Java: in der Ebere im Regenwald sehr zestreut (Kds n. 31172 , 22126 , und 26847 ,). Einh. Name: Aröj ki-kandel lakaki, Sund. ……H. coronaria Bl. Translation: 5b. Corolla middle-sized or very large ( fixed under 1 cm, in diameter) ......................6 6a. Leaf not over 3 cm. long ..................................................................................…7 6b. Leaf 10-15 cm long, elliptical or oblong-elliptical, at the at the base somewhat rounded, above a short sharp point, ± fleshy, on the underside velvety. Petiole 1 ½ -2½ cm. Corolla 2½ - 3 cm in diameter, with outside puberulous, inside tip glabrous, Corona tip outer somewhat rounded, Follicle 22-25 x 6-8 cm, thick cylindrical, ± bloated, straight, up ± blunt. Flower, Bijdr., 1063 (exkl. syn.); Miq; l. c.517; Blume, Rumphia IV.31 tab.182, fig. 1 and 2. West and Middle-Java: very scattered in the plane and in the rain forest. (Kds n.31172 b, 22126 b, and 26847 b,). Native name: Aröj ki-kandel lakaki, Sund.……H. coronaria Bl.

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In Journal of the Linnean Society 42 (1914) 22. ...... while in the forest Cratoxylon arborescens, with its quaint red stems, flowers, and fruit, was veiled in Hoya coronaria, whose branches hung down in curtains bearing corymbs of its green flowers.: 110 under Asclepiadaceae: Hoya coronaria Blume, Bijdr. 1063 Tambunan, 2000’, on swampy ground in sec. forest. Fl. Fr. Feb. 3002. Distrib. Borneo (D. Borneo). Malay Peninsula, Sumatra. A stout twiner on small trees, sending down curtains of long perpendicular shoots. Flowers green. Latex abundant. In Linnean Society Journal. 43 (1915) 110. “The Floral and Plant Formations” Miss L. S. Gibbs. Hoya coronaria, Blume, Bijdr. 1063. Tambunan, 2000', on swampy ground in sec. forest. Fl. Fr. Feb. 3002. Distrib. Borneo (D. Borneo). Malay Peninsula, Sumatra. A stout twiner on small trees, sending down curtains of long perpendicular shoots. Flowers green. Latex abundant. In Journal of the Royal Asiatic Society, Straits Branch Special # Sept. (1921) 506. H. Ridley. 4. Hoya coronaria Blume Bijdr. (1826) 1063; King & Gamble in Jour. As. Soc. Beng. 724 (1907) 566; Gibbs in Jour, Linn. Soc. Bot. 42 (1914) 110; Stapf in Trans Linn. Soc. Bot. 4 (1894) 208. Dutch Borneo, Motley 871; British North Borneo, Gibbs 3002, Burbidge. Malaya Peninsula and Sumatra to New Guinea. In Flora of the Malayan Peninsula 2 (1923) 400. H. Ridley. 19. H. coronaria Bl. Bijdr. 1063; Rumphia IV 31. t. 182, t. 2, and t. 184. A long twiner, velvety pubescent, Leaves thick coriaceous light green elliptic to ovate-oblong short acuminate base rounded velvety beneath 3 - 4 in. long. 1-2 in wide; petiole .75 in. long. Peduncle .75 in long, velvety, rachis thick with large tubercles .5 to 1.5 in. long, pedicels 1 in. long, flowers few in an umbel. Calyx thick velvety, lobes orbicular. Corolla fleshy star-shaped, villose outside, lobes triangular acute, waxy white then becoming spotted with pink, finally all pink, 1.5 in. across. Corona flat spreading. Follicles oblong blunt; pericarp very thick 610 in, long. Hab. Very common in mangrove swamps, river banks, etc. Singapore, Kranji; Jurong; Serangoon; Pulau Tekong, Jahor, Scudai River; Kwala Batu Pahat (Kelsali ) Pahang, Kwantan, Kwala Pahang. Selangor, Sepang. Dindings, Lumut (Curtis), Perak, Matang (Wray); Goping (Scortechini), Penang, Telok Bahang (Curtis). Trengganu, Bundi (Rostados). Distrib. Malay isles, Sumatra to Papua. Native name: Akar Setebal. In De Nuttige Planten von Nederlandsch Indie 2 (1927). K. Heyne. H. coronaria Bl. Volksnamen, Mal. Malakka: Akar Setabal – Soend.: Areuj ki kandel lalaki Corona Ariadnes lutea wordt door Rumphia beschreven (V. bl. 465) als een klimplant - 5 vadm. lang, groetend in het kreupelhout, Blume bericht (Rumphia IV bl. 31), dat het melksap een weinig bitter is en braakwekkend; op Java zou het schler met rauwe tjabe-bladeren en zout door de inlanders inwendig worden gebruikt om een gebrekkige, al te trage, spijsvertering te stimuleern. Translation: Akar Setabal - Soend.: Areuj ki kandel lalaki Corona Ariadnes lutea was described by Rumphia (V. bl.465) as a small plant 4-5 vadm long, describing it as a week plant, Blume mentioned (Rumphia IV bl. 31), that its milk-sap is somewhat bitter and an

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emetic; on Java shall it is mixed with raw tjabe-leaves and salt by the inlanders visceral becomes used about a defective, every to dilatory, and for a digestion stimulent. In Blumea “ Notes on the Flora of Java” 6 (1950) 378. R. C. Bakhausen van den Brink. Hoya coronaria Bl. Bijdr. (1826) 1063; Hooker in Bot. Mag. (1 March 1857 t. 4669). In Flore Siamensis Enumeratio 3 (1951) 35. W. G. Craib & A. F. G. Kerr. Hoya coronaria Blume, Bijdr. 1063 (1826); F.B.I. iv. 58: Mat. F.M.P., No. 19, 566 (776); F.M.P. ii.400, fig.108. Surat. Surat, Kao Ken. c. 50 m., pendant from limestone cliff Kerr 12447 ! Nakawa Sritamrat. Scichonm under 50 m., in light evergreen, kerr 15678 ! Ta Samet, under 50 m., evergreen forest, Kerr 14325 ! Pattani, Tomo, 210 m., Lakshnakara 710 ! Dist. Pen. Mal.! Sumatra (ex F.B.I.), Java ! (type), Borneo ! Celebes ! Local name: Ta bu (script), Malaya. Pattani (ex Lakshnakara). In The Gardens Bulletin, Singapore 20/2 (1963) 191. 1. Hoya coronaria Bl. Brunei: Tutong 4140. Small climbing epiphyte in understory of P.C. 6. Large fruits used by native people in the preparation of a stomach medicine. In Flora of Java 4 (1965) 267. C. A. Backer & R. C. Bakhausen van den Brink. b. Leaves 7-14 cm by 3½- 6 cm, oblong with subparallel margins, from an obtuse-rounded base, very shortly acuminate from broadly rounded or slightly narrowed top, thickly coriaceous, with a strong midrib and faint (on the upper surface somewhat more distinct) nerves, when young thinly pubescent above, subglabrescent, densely shortly pubescent beneath (hairs not inserted on tubercles) and beset with numerous minute papillae, with few thick trichomes; petiole short-hairy, 1½-2½ cm long; stem less densely hairy than leaves. Peduncle thick, pubescent; pedicels densely pubescent, under the fruit 3- 3½ cm long; calyx densely canescent, totaling c. 6 mm; segments ovate-orbicular; corolla c. 3¾ cm diam.; tube widely campanulate, short-hairy outside, c. ¾ cm high; segments broadly ovate-triangular, with strongly deflexed, acute, rather large lobes at the base, shortly acuminate, pubescent, green outside, glabrous, yellowish white, sparingly violet-dotted inside, very thickly fleshy; corona-scales yellow white, at the inner angle short, at the outer angle more than twice as long; oval-ovate, obtuse-rounded; pollinia lanceolateobovate, c. 1 mm long; caudicles curved; translators somewhat longer than the caudicles, c. ¾ mm; follicles rather broadly lanceolate, obtuse, shortly pubescent, 20-22 cm long. I XII: W,-half; below 1000; forest. H coronaria Bl. In Dictionary of Gardening the Royal Horticulture Society 2 (1965) 1015. H. coronaria l. oval, acute, leathery, hairy beneath, margins recurved; stalks terete, hairy. fl. yellow with 5 red spots at base of corona, large, in axillary umbels. Java. 1856. (B.M. 4969.) Syn. H. grandiflora. In Journal of the Royal Horticulture Society (1973). mature, sweet, cloying scent.

A. J. Wills. H coronaria... a

In Malayan Wildflowers (1974) 229. M. R. Henderson. 3. H. coronaria (Crowned Hoya). Fig. 281. Stems velvety hairy, leaves thick, light green, oblong, or sometimes oval

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or elliptical in outline, tip with a short, abrupt point, base more or less rounded, 2-4 in. long, ¾ - 2 in. wide, undersurface velvety-hairy, leafstalk ½ - ¾ in. long; inflorescence stalk short and stout, very rough with tubercles, flowers large, not many in each inflorescence, sepals rounded and velvety hairy, corolla fleshy and star-shaped, white at first, then pink spotted, then pink, about 1¾ in. across, hairy outside, processes from the short stamen column flat and spreading, spoon-shaped; fruit pods large and flat, up to about 10 in. long, with a thick rind, seeds oblong, with a plume about 2 in. long. Common by the sea on rocks and trees, in mangrove and on tidal river banks. Another similar, but much rarer, mangrove species is H. imperials (Royal Hoya). This has larger leaves which are not hairy, flowers about 3 in. across, whitish outside and brown-purple inside. In Hortus (1976) 574. Hoya coronaria Blume. Twining climber, sts. Thickish, puberulent; lvs. Oblong, to nearly 6 in. long, short-acuminate, leathery, with prominent midrib; umbels many-fld.; corolla open-campanulate, to 1 3/8 in. across, yellowish with reddish or violet spots toward the base, corona yellowish. Java. In Malayan Nature Journal 30 #3/4 (1978) 297. “The Peninsular Malaysian Species of Hoya” R. E. Rintz. 16) Hoya coronaria Blume, Bijdr. (1826) 1063. Type: Java, Blume (L). —FIG. 20. Distinguishing Features: Entire plant densely hirsute except on very old parts. STEMS stout. LEAVES coriaceous, elliptical with revolute margins and mucronate tips; densely pubescent below, less so or often glabrous above with age; up to 10cm long by 5cm wide. PEDUNCLE horizontal, rigid, 2 - 3cm long; often short-lived and bearing only one or a few umbels before dehiscing. UMBEL negatively-geotropic, convex with rigid, uniform pedicels 2—3cm long; 1 - 10 flowers, strongly scented at night. COROLLA fleshy, shallowly campanulate; glabrous inside, puberulous outside; base of corolla indexed to form it large gland encircling the base of the gynostegium; 2 - 3.5cm diam; white or yellow, often turning purple before dehiscing. CORONA upper lobe broad and flat; lower lobe with a broad, open groove below; entirely yellow. POLLINIA wingless with long, jointed caudicles and a large triangular corpuscle. FOLLICLE glabrous, up to 22cm long by 5cm diam; the thickness of the fruit being due to the spongy, 1.5cm thick pericarp. Ecology: Common throughout the peninsula on limestone hills and quartz ridges, along the coasts and in mangroves and occasionally along rivers in the lowlands. Distribution: S. Thailand, Sumatra, Borneo, Java, Sulawesi. In Exotica 3 (1978) 1631. Alfred B. Graff. Hoya coronaria (Java) long vining climber with waxy, oval, fresh-green leaves with margins recurved, hairy beneath; waxy flowers pale yellow whit 5 red spots at base of crown. In The New York Botanical Garden Illustrated Encyclopedia of Horticulture 5 (1981) 1726. H. coronaria is Javanese. It has short-haired, thick stems and short-pointed, oblongish to roundish leaves up to 6 inches long and with a conspicuous midrib. Smooth above, they have hairy undersides. The open-bell-shaped flowers 1 inch to 1¼ inches across and white to yellowish spotted with red or purple around the yellow corona, are in umbels of many.

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In Tropica (1992) 1020. A. B. Graf. Hoya coronaria (Java); long vining climber with waxy, oval, fresh green leaves with margins recurved, hairy beneath; waxy flowers pale yellow with red 5 spots at base of crown. Tropical. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 86 “The Genus Hoya in Thailand” O. Thaithong. 2. H. coronaria Blume, Bijdr. (1826) 1063. Rumphia: 4 (1883) 58; Fl. Java 2* (1965) 267; FMP 2 (1923) 400, Fig. 108; Fl. Siam. Enum. 3(1) (1951) 35; Malay. Nat. J. 30 (1978) 505, Fig. 20. Occurrences: (PEN): Surat Thani, Nakhon Si Thammarat, Pattai, Songkhla.

Hoya corona ariadnes Blume In Rumphia 4 (1848) 51. Blume. 2. H. corona Ariadnes tab. 182 (analysis) et 185; caule scandente; foliis ovato-ellipticis oblongisve apice in acumen breve recurvam constrictis basi rotundatis marginibus recurvatis carnosis subvenosis glabris; umbelis 6-11 floris; corollae maximae laciniis late triangularibus-convexis intus papilloso-subvelutinis; coronae staminae foliolis angulo exteriore subrotundatis Bl. Cent., Plant. nov. - H. speciosa et H. Ariadna De Cand. Prod. Sys. Veg. VII. p. 634 1 ed. 3. Corona Ariadnes punicea Rumph. Herb. Amb. V. p. 464. tab. 182. - Habit. in Moluccis, Celebes et Borneo in sylvis humilioribus, floribus magnis puniceis, sub vesperam odorem dulcem Ananassi maturi instar exhalantibus, omnium Specierum cognitarum facile pulcerrima et idcirco nuperrime caldariis nostris importata ab horticultoribus nomine H. imperialis salutata. Caulis fructicosus, inferne lignosus, teres, alte scandens et floribus suis magnificis ornatus de arboribus pendulus. Folia opposita, petiolata, patentissima, ovato-elliptica v. ovato-oblonga superne in acumen breve acumen recurva, basi rotundata, carnosa, firma, supra lucida ac subvenosa, subtus pallidiora nervo medio crasso valde prominente. Petioli teretiusculi, ecanaliculati, 8-11 lin. longi; folia 2.5- 5.5 poll., 1.5-2 poll. lata. Pedunculi axillares v. exteaaxillares et terminales, solitarii, 1-1.5 poll. longi, umbellato 6-11 flori. Pedicelli 1.5-2 poll. longi, superne aliquanto crassiores, ut calycis et alabastra pentagona viriduli. Calycis foliola ovato-rotundata, extus convexa, margine membranacea, obsolete ciliata. Corolla aperta plus 2 poll. diametro, coriacea, extus viridis et circum basim glabra, limbum versus pube rarissima appressa, intus e puniceo in atro-purpureum vergens et inconspicue papilloso-velutina; laciniis limbi patents lato-triangularibus, acutis, convexis, margine subrevolutis. Corona staminea lutescens, nitida; foliolis horizontalibus, carnosis, supra paulum depressis, subtus lato-sulcatis, angulo exteriore obtusissimis, interiore acutis illic intus connectivo membranaceo antherarum ovato obtusiusculo ad marginem interiorem carneo stigmati incumbenti conjunctis. Retinacula emarginaturis stigmatis cum foliolis coronae stamineae alternantibus affixa, cuneata, basi subbifida, sulco longitudinali inscripta, superne bicornuta, coronea, alto-purpurea. Pollinia clavata, subarcuata, duriuscula, flava, basi per stipitem brevem arcuatum cornibus retinaculi affixa. Ovaries 2, oblonga. Stigma magnum, peltatum, e papilla centrali radiis quinque elevatis et reticulis oppositis. Folliculi ab ortu solitarii, plus ventrali inscripti, quo tandem dehiscunt. Translation: stems climbing, leaves ovate-elliptic oblong, apex briefly sharpened, recurved constricted, base rounded margines recurved fleshy somewhat veined glabrous;

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umbels 6-11 flowers; corolla large lobes wide triangular inside papilose somewhat velvety; leaflets of the staminal corona with the exterior angle somewhat rounded, Blume Cent. Plant. nov. - H. speciosa & H. ariadna DeCaisne in De Cand. Prod. Sys. Veg. 8. p. 634. 1 and 3. Corona Ariadna punicea in Rumph. Herb. Amb. 5. p. 464. tab 182. Habitat: in Moluccas, Celebes and Borneo, in the humid forests, flowers large crimson, somewhat an evening perfume like inhaling from the pineapple, up to this time easily the only known specimen, a very beautiful and therefor recently imported into hot houses from nature namely H. imperialis. Stems woody, lower part ligneous, round, high climbing and flowers of themselves magnificently ornate with the arbor pendulous. Leaves opposite, petiolate, very flat, ovate-elliptic or ovate oblong, above surface briefly acutely recurved, base rounded, fleshy, firm, shinny above and somewhat veined, below more pale with the central nerve thick strongly prominent. Petioles continuously round, without a longitudinal channel, 8-11 lines (line=1/12 of inch) long; leaves 2.5-5.5 inches, 1.5-2 inches wide. Peduncles axillary or extra axillary and terminal, solitary, 1-1.5 inches long, 6-11 flowered umbels. pedicels 1.5- 1.75 inches long, above somewhat fleshy, calyx and flower bud truly 5 sided. Calyx leaflets ovate-rounded, outside convex, margines membranaceous, with a few cilia. Corolla open more than 2 inches in diameter, leathery, outside green and around the base glabrous, toward the limb very rarely with appressed down, inside crimson tending toward dark purple and inconspicuously velvetypapilose; limb lobes flat broadly triangular, acute, convex, margines somewhat revolute. Staminal corona yellow, shinny, leaflets horizontal, fleshy, above a little depressed, below longitudinally grooved, exterior angle obtuse, interior acute and their connected to the ovate obtuse anther membrane, connected at the interior margine and incumbent on the flesh colored stigma. Retinacula affixed on the margines of the stigma and alternate with the staminal corona. wedge shaped, base somewhat bifid, inscribed with longitudinal furrows, rising above the 2 crowns, crown dark purple. Pollinia club shaped, somewhat curved, somewhat hard, yellow, affixed at the base by means of the short slightly curved stem from the horny retinacula. Two oblong ovaries. Stigma large, shield shaped, with a central nipple radially 5 elevations opposite the retinacula. Follicle arising solitarily, more or less long, pendant, conical elongate, above attenuate and by no means curved, glabrous, green, longitudinally striate, with a ventral groove, indeed at length dehiscing. In Museum Botanicum Lugduno-Batavum 1 (1849) 43. C. L. Blume. (9. Hoya corona Ariadnes Bl.: caule scandente; foliis carnosis ovato-ellipticis oblongisve apice in acumen breve recurvum constrictis basi rotundatis marginibus recurvatis subvenosis glabris; corollae maximae laciniis late triangularibus convexis intus papilloso-subvelutinis; coronae stamineae foliolis angulo exteriore subrotundatis. Bl. Cent. Plant. nov. et Rumphia IV. tab. 183. - H. speciosa et H. Ariadna Decaisne in De Cand. Prodr. Syst. Veg. VIII. p. 464. tab. 182. - In Moluccis, Celebes et Borneo in sylvis humilioribus. Translation: stems climbing, leaves fleshy ovate-elliptic, apex briefly sharpened recurved constricted, base rounded, margines recurved, somewhat veined, glabrous; leaflets of the corolla large broadly triangular, convex, inside papilose-somewhat velvety; leaflets of the staminal corona with exterior angle somewhat round. ........ ...... In the Moluccas, Celebes and Borneo in the humid forests.

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In Tuinbouw Flora 1 (1853) 70. DeVriese. 9. H. Corona Ariadnes Bl.. De kroon van Ariadne is in de Rumphia afgebeeld IV. pl. 183. Deze prachtige Hoya komt voor op the Celebes en Borneo in lage, boschachtige streken. Zij geeft, volgens Blume, des avonds een zeer sterken geur als van ananassen. Het kan zijn, dat wij ons bedriegen, maar, te oordeelen naar de van H. imperialis tot onze kennis gekomen af beeldingen en de prachtige bloemschermen, die wij in 1851 zagen in de kassen van Sion House, kunnen wij nog niet met volle overtuiging aannemen, dat zij dezelfde zou zijn met H. Corona Ariadnes, ‘t geen ik meen als ‘t gevoelen van Dr. C. L. Blume te moeten houden (t. a. pl. IV. 31). Intusschen moet ik bekennen, dat er hier eene groote moeijelijheid is om de zaak te beslissen. De bloemen van Blume’s afbeelding van H. Corona Ariadnes zijn donkerrod in ‘t bruin overgaande. Die van de Engelsche afbeeldingen van H. imperialis zijn meer in het paars overgaade; die van Sion House waren bleek paars in’t rozenkleurige van H. imperialis, zoo als ze is afgebeeld en zoo als ik ze zag, is een ware, zeer rijkbloeming scherm; dit is het geval met H. corona Ariadnes niet. Deze is niet de plaats voor een dergelijk onderzoek. Welligt is van den schrijver der Rumphia te wachten, dat hij deze twijfelingen zal oplossen. Translation: (by Albert Hofman). The crown from Ariadne is pictured in Rumphia IV. pl.183. These magnificent Hoyas can be found on the Celebes and Borneo at low elevations, in woody regions. It produces according to Blume, in the evening a very strong odor as from pineapple. It can be that we are mistaken, but, to judging after having seen the pictures and beautiful umbels in the greenhouses of Sion House, of Hoya imperialis, we are not completely convinced yet that it would be the same as H. Corona Ariadnes, which I think is the feeling of Dr. C. L. Blume. (t. a. pl. IV. 31). Therefor I must confess in the meantime, that it is difficult to decide in this case. The flowers from Blume's picture of H. Corona Ariadnes are dark red with brown overtones. Those from the English pictures of H. imperialis are more towards purple, those from Sion House were pale purple, nearly rose colored as is H. imperialis, as it has been pictured and as I saw it, is a true, very rich-flowering umbel; this is not the case with H. corona Ariadnes. This is not the place for a such, investigation. Perhaps we might expect the author of Rumphia to solve these doubts.

Hoya costantinii P. T. Li In Bulletin of Botanical Research 4 (1984) 120. P. T. Li. Hoya costantinii P. T. Li, nom. nov. Hoya reticulata Cost. in Lec., Fl. Gen. Indo-chine 4:1912, non Moon (1824), nec Merr. (1912), Schlechter (1913) Distr.: Cambodia. Editors Note: See Hoya multiflora.

Hoya crassicaulis Elmer ex Kloppenburg In Leaflets of Philippine Botany 10 (1938) 3579-3580. A. E. D. Elmer. Hoya crassicaulis Elm. n. sp. Climbing along the interlaced stem of a strangler fig species. Old stem the size of a small finger, subligneous, terete, crooked, repeatedly branched, very tough but stout, reaching up to 7 m. above the ground, more or less gnarled at the point of branching, the main stems curved and occasionally twisting, that portion on my dried specimen only 5 mm thick, barely bendable, smooth and glabrous, curing a dirty

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yellowish brown; lateral branches short, many of then thick and roughened with large leaf scars. Leaves deciduous, opposite or in clusters, descending, exceedingly thick and consequently very rigid, shallowly conduplicate, strongly recurved toward the apex, the fresh ones yellowish green but paler beneath, glabrous on both surfaces, oblong, gradually tapering into the rather sharply acute to acuminate point, base obtusely rounded, varying from 10 to 17 cm long without the petioles, from 5 to 8 cm wide across the middle or a trifle below it, oblong but occasionally ovately oblong, the entire margins often pressed minutely involute, the upper surface sublucid, curing similarly dull yellowish brown on both faces; midrib region thickened especially toward the base, longitudinally wrinkled in the dry leaves; nerves several from each side, oblique, not very evident but more so on the upper surface, otherwise wrinkled not reticulated; petioles brownish on the live plants, glabrous, much wrinkled or rugose and yellow when cured, channeled along the upper side, 1.5 to 5 cm. in length, thick and fleshy, leaving very large and circular scars upon falling, persistent to the blades, a trifle thicker toward the base which is frequently curved. Flowers fragrant, forming a globose cluster from the terminal leaf axils of the stems and branchlets, dull except the purplish brown perianth tips; peduncles 5 cm long more or less, appearing minutely papillate or punctate, earth colored, ascending; pedicels 1.5 to 2.5 cm long, narrowly oblong, subhyaline except the reddish brown acute tips, besides the 5 outer lobes there are a pair of inner ones; corolla rotate, united at the base, also hyaline, glabrous, membranous, 5 mm long and as wide across the middle, the upper half strongly reflexed, the acute apex prominent; horns of the corona 5 and alternating with the corolla lobes, wheel like, ovately oblong and a pair of pockets in the sinuses. Follicles dry, dehiscing along the upper side, terete, linear 12 cm long on ours; seeds 3 to 5 mm long, compressed, pale brown, obtusely rounded at the base, at the truncate apex bearing a dingy coma. Type specimen number 14440, discovered by A. D. E. Elmer climbing a Balete in the hemp region at 250 feet elevation, Irosin (Mt. Bulusan), Province of Sorsogon, Luzon, October 1915. Our specimen is probably most closely related to Hoya incrassata Warburg. Compare it with my number 16718 from this same locality and which number I now consider that of Warburg’s species. Our new species here described should also be critically examined with Bureau of Science number 834 collected by Foxworthy on Palawan. The shape, size and venation of this small but critical group of Hoya species with thick leaves must be carefully considered. There are some plants almost identical in foliage yet so different in flowers, and we can only imagine what ripe fruits will reveal. In Fraterna “International Hoya Association” 3rd. Qtr. (1995) 10-13. Dale Kloppenburg. Hoya crassicaulis Elmer ex Kloppenburg n. sp.. Hoya crassicaulis Elmer nomen invalid. Holotype Elmer #14440 (BO) discovered by A.D.E. Elmer climbing on a belete in the hemp region at 250' elevation, Irosin, Mt. Bulusan, Sorsogon, Luzon, Philippines in October 1915. Isotypes at UC and A/2. Planta epiphytica, scandens, ramulis crassis 0.5 cm. diametro, laevis et glabris, foliis coriaceis ellipticus ad oblongo-ovatis, nitidis, 8-17 +/- cm. longis medio fere 5-8 cm latis, basi late rotundatus vel paullo subcordatus, apice tenuiter acute acuminatis ad subacuminatus et recurvis, marginibus subrevolutis, costa subtus valde prominente, venis utrinque ca. 4 patalis ante marginem arcuate confluentibus, umbellis multifloris, 60-125

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florae; petiolis glabris, superne sulcato 1.5-5 cm. longis carnosis et incrassatis; pedunculo usque 5 cm. longo; pedicellis tenuiter filiformibus 1.5-2.5 cm. longis; calycis segmenta 0.15 cm. longis ovate-triangula, obtusiuscule, sub hyaline, extus punctate et sparsissime ciliolata vel glabris; corolla rotata apicibus reflexis, usque ad tertiam partem basilarem 5fida, lobis rhomboideo ovatis, acutis, glabra, intus punctate papillosa, ca. 1 cm. + diametro; coronae stamineae lobis late ellipticis patulis navicularibus, supra concavis, utrinque 0.37 cm. longis 0.17 cm. latis; folliculis 9-11 cm. longis, usque ad 0.4 cm. diametero. This species is closest to H. incrassata Warburg and has been confused with and lumped with this species in commerce as well as mislabeled on many herbarium sheets. After studying Elmer's type sheet #14440 at (UC) and (BO), examining 23 clones, most of which I now consider H. crassicaulis Elmer ex Kloppenburg, I now consider this a distinct species and not part of a (cline), a continuous series of variation between two extremes. H. incrassata Warburg is relatively rare in the Philippines in comparison to this new species. There are many existing clones (23 or so) in commerce which exhibit clonal variations in a number of characteristics including umbel flower number, umbel diameter, pedicel length, and foliar characteristics. None approach the smaller diameter flower cluster of Hoya incrassata Warburg, the smaller calyx size (also usually more ciliate), longer peduncle, shorter pedicels, smaller corona among other differences. The pollinaria of the two species are very distinct and I have not found any intermediate types. The retinacular head of H. crassicaulis is long when compared to the short winged head of H. incrassata's. The attachment of the translators are just below the short broad (winged) head in this latter species whereas H. crassicaulis has a elongated more rounded narrow head, has translators attached well down (below the waist) of the retinaculum. The remanding portion of the retinaculum in this species is broad hipped, reminding me of a tarantula in shape. The retinaculum is so long and the attachment of the translators and caudicle are so low down that on examining this structure the retinaculum often turns on this axis so as to present itself in an unnatural configuration. The corona scales of H. crassicaulis are longer, cupped above with the outer lobe elevated near the apex, giving a sway backed cross section. H. incrassata's scales are more bulky, shorter, thicker and essentially horizontal. Both species have paired anther wings, more prominent in H. crassicaulis. The two species are undoubtedly evolutionarily linked but they do not represent a cline. The type sheet for H. crassicaulis Elmer ex Kloppenburg is 14440 1915 Elmer (BO) also at (UC, 2/A). I believe #46166 1923 Edano (Pasal river, Panay, Philippines) and #21613 1927 Wentzel (BO, UC, A) and #2613 1927 Wentzel (BO,UC,A) also represent this species. Elmer believed his sheet #16718 (BS) was H. incrassata Warburg. Two co-types are listed by Dr. Warburg i.e. #12997 and #12998. In addition the following sheets labeled as H. incrassata Warburg should now be critically examined: #1003 Weber (BS), 834 Foxworthy, Palawan, #10431 McGregor, 14387 Warburg, #4082 Merrill (SI, BS), #10336 McGregor, 393 Bolster, #32384 McGregor (BS), #29323 McGregor (BS), #14223 Ramos (BS), #14730 Ramos (BS), 22822 McGregor, #3257 Ramos (BS), #34530 1919 Ramos/Pascaso and #83385 Ramos/Convocar. Merrill's #11591 1922 is also this species. Comparisons taken from photomicrographs follows: If you have purchased H. incrassata from Hill n' dale the labels should be

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changed to read H. crassicaulis. Those from Green Plant Research of Hawaii labeled H. crassicaulis are correct and those labeled H. incrassata from Rainbow Gardens of Vista CA are also correct. Dale Kloppenburg 20 Dec. 1993 Translation of Latin above: Plant epiphytic, trailing, branches thick 0.5 cm. in diameter, smooth and glabrous, leaves leathery elliptic or oblong-acute, shiny, 8-17 cm. ,ore or less large, in the center nearly 5-8 cm. wide, base rounded or a little somewhat acuminate and recurved, margines somewhat revolute, midrib prominent below, veins on both sides about 4, spreading curving confluent to the margines, umbels many flowered 60-125 flowers; petiole glabrous, above grooved 1.5 to 5 cm. long fleshy and thickened; peduncles at least 5 cm. long; pedicels narrowly threadlike 1.5 to 2.5 cm. long; segments of the calyx 0.15 cm. long ovate-triangular, somewhat obtuse, somewhat hyaline, outside punctate and sparsely ciliate or glabrous; corolla rotate apices reflexed. 5 parted all the way to 1/3 from the base, lobes reflexed ovate, acute, glabrous, inside punctate papilose about 1 cm. in diameter, lobes of the staminal corona elliptic outspread, boat shaped above concave, certainly .37 cm. long .17 cm. wide, follicles 9-11 cm. long, up to 0.4 cm. in diameter. In Philippine Hoya Species 3 (1996) 57-58. R. D. Kloppenburg. Hoya crassicaulis (Elmer) Kloppenburg in Fraterna 3rd quarter issue 1995. Type: Elmer 14440, climbing a belete in the hemp region at 250' elevation, Irosin (Mt. Bulusan), Province of Sorsogon, Luzon, Philippines, October 1915. Key #47 Elmer's English description follows. Climbing along the interlaced stem of a strangler fig species. Old stem the size of a small finger, subligneous, terete, crooked, repeatedly branched, very tough and stout, reaching up to 7 m. above ground, more or less gnarled at the point of branching, the main stems curved and occasionally twisting, that portion on my dry specimen only 5 mm. thick, barely bendable, smooth and glabrous, curing a dirty yellowish brown; lateral branches short, many of them very thick and roughened with large leaf scars. Leaves deciduous, opposite or in clusters, descending, exceedingly thick and consequently very rigid, shallowly conduplicate, strongly recurved toward the apex, the fresh ones yellowish green but paler beneath, glabrous on both surfaces, oblong, gradually tapering into the rather sharply acute to subacuminate point, base obtusely rounded to broadly cuneate or rounded, varying from 10 to 17 cm. long without the petioles, from 5 to 8 cm. wide across the middle or a trifle below it, oblong but occasionally ovately oblong, the entire margins often pressed minutely dull yellowish brown on both faces; midrib region thickened especially toward the base, longitudinally wrinkled in the dry leaves; nerves several from each side, oblique, not very evident but more so on the upper surface, otherwise wrinkled not reticulated; petioles brownish on the live plants, glabrous, much wrinkled or rugose and yellow when cured, channeled along the upper side, 1.5 to 5 cm. in length, thick and fleshy, leaving very large and circular scars upon falling, persisting to the blades, a trifle thicker toward the base which is frequently curved. Flowers fragrant, forming a globose cluster from the terminal leaf axis of the stems and branchlets, dull except the purplish brown perianth tips; peduncle 5 cm. long more or less, appearing minutely papillate or punctate, earth colored, ascending; pedicels 1.5 to 2.5 cm. long, filiform, pale yellowish

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brown; calyx segments 1.5 m. long, narrowly oblong, subhyaline except the reddish brown acute tips, besides the 5 outer lobes there area pair of inner ones; corolla rotate, united at the base, also hyaline, glabrous, membranaceous, 5 mm. long and as wide across the middle, the upper half strongly inflexed, the acute apex prominent; horns of the corona 5 and alternating with the corolla lobes, wheel like, ovately oblong and with a pair of pockets in their sinuses. Follicle dry, dehiscing along the upper side, terete, linear, 12 cm. long on ours; seeds 3 to 5 mm. long, compressed, pale brown, obtusely rounded at the base, at the truncate apex bearing a dingy white coma. Meaning: crassicaulis—L. crassus; caulis a thick stemmed plant.

Hoya crassifolia Haworth. In Supp. Plant. Succ. (1811) 8 Haworth. Hoya crassifolia. In Transactions of the RHS 7 (1826) 22-23. J. Traill. II. Hoya crassifolia. I find this plant only noticed by one writer, to whose work I add the references. H. crassifolia. Haworth in Supp. Plant. Succ. p. 8. A plant of this species was sent to Mr. Reginald Whitley, from China, in 1817, on board the Wexford East Indiaman, and has since been cultivated and increased in his nursery at Fulham. It is very distinct from, and much more robust growth than, any of the others. The stems are strong and woody, thickly covered with small irregular warts, whence the stem-roots are protruded. The leaves are obovate, very bluntly acuminate, upwards of four inches long, without veins, of a darkish green colour above, pale beneath. The petioles are thick and fleshy, of a purplish green colour, and much flattened above. The plant grows freely in strong heat, but has not yet produced flowers in Europe, it is therefore uncertain whether it belongs to the genus, although it may be presumed so, from its general habit not differing from Hoya. The specific name crassifolia, which had been given to the H. carnosa, by Baron Jacquin in 1811, has been applied to this plant by an inadvertence on the part of the author above quoted, he supposed Jacquin’s plant to be identical with this one, referring it to Jacquin’s figure in his Ecologae. Though the Hoya (Schollia) crassifolia of Jacquin, was published by him in 1811, and was stated in his Ecologae to have been sent to him in England in 1804, yet Mr. Haworth has refereed the plant first introduced by Mr. Whitley in 1817 to it, and has adopted Jacquin’s specific mane, and even stated the large spots on the leaves so conspicuously exhibited in Jacquin’s figure as distinguishing this plant from H. carnosa. The name is objectionable, inasmuch as it may occasion confusion in references, and when the plant blossoms so that it can be fully and perfectly described, it will probably receive some other. In General System of Gardening and Botany 4 (1837) 126. G. Don. 19 H. crassifolia (Haw. suppl. pl. succ. p. 8. exclusive of the synon..) leaves petiolate, obovate, obsoletely spotted, bluntly acuminate, very thick. (woody perennial) Native to the east Indies. Leaves 4 inches long, dark green above, pale beneath. Very like H. carnosa, but the leaves are thicker, more obsoletely spotted . Branches warted, producing roots from the warts, Flowers unknown, Thick-leaved Hoya. Fl.? Clt. 1817. shrub tw. In Decandolle, Prodromus Syst. Veg. 8 (1844) 60. Decaisne. (Species non statis notae)

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H. crassifolia (Haw, succl. pl. suppl. p. 8 ), foliis oppositis petiolatis subovatis obsolete maculatis praecrassis, floribus umbellatis, petalis truncatis obtussive. Affinis H. carnosae at foliis obtusioribus crassioribus, maculis marginis obsolentis pallescentibus. (woody perennial) in India. Schollia crassifolia Jacq. eclog. pl. t. 2. Traill l.c. p. 22. Translation: leaves opposite petiolate somewhat ovate obsoletely spotted, flowers in umbels, petals truncate obtuse. Related to H. carnosa with foliage more obtuse, much thicker spots paler disappearing near the margins. In Flora of British India 4 (1883) 62. J. D. Hooker. (Doubtful and Excluded Species). H. crassifolia, Haw. Succ. Pl. Suppl. 8; Dcne. in DC. Prodr. viii, 640 Schollia crassifolia, Jacq: Eclog. t. 2; Traill, in Trans Hort Soc. vi 22; from India; with leaves described as excessively thick, obsoletely spotted, petals truncate or obtuse, flowers unknown. Wight (Contrib. 38) refers it to the Chinese H. carnosa, Br. which it would supersede.

Hoya crassifolia Ridley In Journal of the Royal Asiatic Society, Straits Branch 61 (1912) 30-31. H. Ridley. Hoya crassifolia, n. sp. Stem rather stout with rugose often pale bark rooting, profusely. Leaves very coriaceous ovate acuminate slightly cordate at the base, margins undulate and minutely irregular dentate, nerves very inconspicuous 3-4 pairs at an acute angle, midrib inconspicuous 6 inches long 2½ inches wide, petiole very short and thick. Peduncle 1½ inches long rachis ½ inch long with elliptic scars, pedicles slender ¼ inch long, hairy with long rufous hairs. Sepals small linear lanceolate acuminate. Corolla half an inch across pink, lobes triangular cuspidate with a long point, villous with long white hairs on the edge, Corolla scales, lower lobes long thick blunt not smooth and spining, upper ones short erect about half as long triangular acuminate. Staminal column short thick appendages long linear acuminate thin membranaceous and translucent white. Perak: Tapah on trees on the Temoh Road (Ridley 14059). A most remarkable plant with its hard stiff leaves roughened with short tooth-like processes at the edge, hairy pedicels and long thin stamen appendages. Editors Notation: It seems to me Ridley or the publishers left out the corolla description and under corolla describe the corona. See H. caudata in Flora of the Malay Peninsula.

Hoya crassior Hochreutiner In Candollea 6 (1936) 480-481. Hochreutiner. Hoya crassior Hochr. sp. nov. — Planta epiphytica; caules ± cylindrici glabri, in nodis incrassati. Folia opposita, mediocri; petioli breves, glabri, ± carnosi, supra caniculati et in parte cum lamina uniti ± glandulosi; lamina glaberrima, crassissima, carnosa, ita ut nervus utrinque impressus videatur, centrum lanceolata, apice paululum attenuato-acuminata, basi acuta, margine integra, nervi secundarii inconspicui, vel in laminis foliorum supremorum axilla affixa vel ± extra axillam sed juxta eam deposita. Pedunculus mediocris glabra, quam pedicelli brevior; pedicelli filiformes, glabra, numerosi. Calyx glaber, profunde lobatus, lobis ovatis obtusis, margine pilos paucos minutissimos gerentibus. Corolla non profunde lobata et inter lobis, in sinubus, glandulosa; corolla tota glabriuscula glaberrima. lobis acutis sed intus, versus lobarum apicem, aream parvam villosam praebentibus. Corollae

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phylla cum staminibus concreta, ovoidea, facie suprtiore apiculum in stamina adpressum praebentia, apice dilatata ita ut in appendicem scariosam ovatum abeuntia; pollinia parva elliptica. Stigmatis caput applanatum sed medio breviter apiculatum. Ovarium breve, in gynostegii cavernula parva dispositum ± conicum. Petioli 3-4 mm. longi; lamina 6x1.6 4x1 cm. longa et lata. Pedunculus ca. 1 cm. longus, pars florifera 3-4 mm. longa; pedicelli 1.5- 1.7 cm. longi. Flos expansus minus quam 1 cm. latis, - siccus ca. 0.5 cm. latis. - Calyces lobi ca. 1.25 x 0.75 mm. longi et lati; corolla ca 4 mm. longa; coronae phylla sicca ca. 1.5 x 0.75 mm longa et lata. Samoa, ile d’Upolu, route du lac Lanuto, alt. 400 m. mais pas plus haut, epiphyte a fleurs d’un blanc jaunatre, 26 mars 1905 (n. 3411). Cette espece est tout as fait differente de celles que Reinecke a signalees a Samoa, dont l'une a des feuilles 5-nerviees avec une corolle de 1.5 cm. de diametre et l’autre des feuilles laineuses. Le H. bicarinata Gr. en differe aussi par ses feuilles ovees et les articles bicarenes de sa couronne. Rechinger a decrit egalement trois especes novelles se Hoya de Samoa dont l’une a des feuilles tres petites (3 cm.) et l’autre des fueilles tres minces qui ne cadrent pas avec notre espece; en revanche, la troisieme espece le H. pycnophylla Reich. resemble a note plante et la photographie qu’em donne Rechinger (Denkschr. Akad. Wien, math. naturw. Pl. LXXXV, t. XII, f. 1 (1910) nous a fait croire, a premiere vue, qu’il s’agissait de la meme espece. Cependant, quoique la description de la fleur soit rudimentaire, neanmoins la presence des feuilles montrant cinq nervures longitudinales sur toute leur longueur, malgre l’epaisseur du limbe (exprimee par le nom de pycnophylla) et l'indication: “nervi utrinque manifeste prominentes” alors qu’ici la nervation est purement pennee et les nervures completement noyees dans le mesophylle, montrent qu’une identification est impossible. On peut citer encore dans la diagnose de l’auter: “segmenta calyces... apice acuminata glabra” tandis que, chez notre espice, les sepales sont obtus et cilies. Il est vrai que les indications de Rechinger sont quelque peu sujettes a caution car, dans sa description il dit: petiole de 1 cm. , limbe de 9 x 2 - 10 x 3 cm., pedoncule 5-6 cm. et pedicelles 15 mm. alors que, sur la photographie de sun specimen. on measure des petioles de 5-7 mm., des limbes 3.8 x 1 - 5 x 1.5 cm., des pedoncules de 1-2 cm. et des pedicelles de 5-8 mm.; cependant, on ne peut pas se tromper sur la nervation foliaire, bein visible sur la photographe, c’est pourquoi une identification nous a paru impossible. Enfin. se nous comparons notre espece au H. diptera Seem. nous observons que, chez ce dernier, les tiges quadrangulaires. les feuilles ovees elliptiques, les decrit pour Samoa un Physostelma Betchei mais, si nous en croyons la description, cette plante differe de la notre par les dimensions de l’inflorescence et de la fleur et par la corolle velue interieurement. En l’apparance de la corolle, que est peu profundement lobee chez notre espece, pourrait induire les asclepiadologues a en faire un Physostelma crassior. Translation: Hoya crassior Hoch. species new. Plant epiphytic, stalks more or less cylindrical glabrous thickened at the nodes. Leaves opposite, midsize, petioles short, glabrous, more or less fleshy, channeled above and where united with the blade somewhat glandular; blade very glabrous, very thick and fleshy, the midrib it seems depressed on but sides, otherwise lanceolate, apex a little attenuate-acuminate, base acute, margines entire, secondary nerves inconspicuous, or in juvenile leaflet blade about 3, both sides arching and meeting and with reticulate nerves. Inflorescence ambulate somewhat terminal i.e. affixed in the highest leaf axils yet more or lass outside the axils. Peduncles moderately glabrous, but shorter than the pedicels; pedicels threadlike ,

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glabrous, numerous. Calyx glabrous. deeply lobed, lobes ovate obtuse, margines minutely sparsely ciliate, Corolla not deeply lobed and inner lobes at the sinus, glandular: corolla totally glabrous to very glabrous, lobes acute but towards the lobe apexes exhibiting a small area of villoseness. Scales of the corona with stamens grown together, ovoid, making the upper apex to the stigma. Stigma broad elliptic, more or less ovate, margines becoming scarious, apexes swollen with the appendages scarious tapering ovate; pollinia small elliptic. Stigmas head flattened but briefly apiculate in the middle. Ovaries small, and more or lass deposited in the gynostegium cavity. Petiole 3-4 mm. long; blades 6x1.6 x 4 x 1 cm. wide. Peduncle about 1 cm. long, flower bearing part 3-4 mm. long; pedicels 1.5 - 1.7 cm. long. Flowers expanded less than 1 cm. wide, - dried about 0.5 cm. wide. - calyx lobes anout 1.25 x 0.75 mm. long and wide; corolla about 4 mm. long; leaflets of the corona dry about 1.5 x 0.75 mm. long and wide. Samoa on the island of Upolu, the road to Lake Lanuto, alt.400 m. but not higher, epiphytic with yellowish white flowers, 26 March 1905. (number 3411). This specimen is completely different from those that Reinecke observed on Samoa, one of which has five-ribbed leaves with a corolla 1.5 cm. in diameter and the other with furry leaves. The H. bicarinata Gray differs from them also by its ovoid leaves and bicarinate articles of the crown. Rechinger likewise described 3 new species of hoya from Samoa of which one has very small leaves (3.cm.) and the other very thin leaves which do not tally with our species; on the other hand the 3rd species, H. pycnophylla Rech. resembles our plant and the photo that Rechinger gives of it (Denkschr. Akad. Wein, math Natur. pl. 85 , t. 12, f. 1 (1910) leads us to believe, on first glance, that is was the same species. However, although the description of the flower is rudimentary, nevertheless the presence of the leaves showing 5 longitudinal ribs on their whole length, despite the thickness of the leaf blade (expressed by the name pycnophylla) and the indication (nerves on both sides manifestly prominent) while here the ribbing is purely pinnate, and the ribs completely submerged in the mesophyll, shows that an identification is impossible. One can further cite in the diagnosis of the author (calyx segments .... apexes acuminate, glabrous) while in our species the sepals are obtuse and ciliate. It is true that Rechinger’s indications are somewhat subject to caution for in his description he says: petioles of 1 cm; leaf blade of 9x2 to 10 x 3 cm. peduncle 5-6 cm., and pedicels 5-8 mm. that is why an identification appears impossible to us. Finally if we compare our species to H. diptera Seemann, we observe that with this later plant the quadrangular stems, the ovate-elliptical leaves, the compressed peduncles, do not tally with our plant. Schlechter described for Samoa a Physostelma Betchei, but if we believe his description this plant differs from ours by the dimensions of the inflorescence and the flower and by the corolla which is hairy on the inside. On the other hand the appearance of the corolla which is not deeply lobed in our species could induce asclepiadologists to make is a Physostelma crassior.

Hoya crassipes Turczaninow In Bulletin de la Societe Imperiale des Naturalistes de Moscou 21 (1848) 261. N. Turczaninow. Hoya crassipes Turcz. (Hoyae verae): volubilis, foliis petiolatis ellipticis aut obovatis, ad basin attenuatis, ipsa basi obtusis, breviter acuminatis, carnosis aveniis

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glabris; pedunculis extra axillaribus brevibus crassis in receptaculum carnosum cylindricum, sulcis transversalibus irregulariter exaratum abeuntibus; umbellis in vertice receptaculi sessilibus multifloris; corollae laciniis triangularibus acutis glabris, intus papillosis, ad faucem 5-maculatis; coronae stamineae foliolorum parte inferiore ovata obtusa concaviuscula uninervia, subtus sulco notata, superiore acutiscula gynostegium paulo superante, Species forma receptaculi distinctissima. Flores fere duplo minores illorum H. carnosae. Habitat in Java, Zoll. coll. n. 2581. Translation: Same Latin and translation as below in Annales Bot. Syst. In Annales Botanices Systematicae 3 (1852) 9-10. G. C. Walpers. 4. H. crassipes Turcz. Bulletin de la societe imp. natural. de Moscou XXII. Part. I. Pag. 261. - Volubilis: foliis petiolatis ellipticis aut obovatis; ad basin attenuatis, ipsa basi obtusis, breviter acuminatis, carnosis aveniis glabris; pedunculis extra axillaribus brevibus crassis in receptaculum carnosum cylindricum, sulcis transversalibus irregulariter exaratum abeuntibus; umbellis in vertice receptaculi sessilibus multifloris; corollae laciniis triangularibus acutis glabris, intus papillosis, ad faucem 5-maculatis; coronae stamineae foliolorum parte inferiore ovata obtusa concaviuscula uninervia, subtus sulco notata, superiore acutiscula gynostegium paulo superante. Species forma receptaculi distinctissima. Flores fere duplo minores illorum H. carnosae. - Habit in Insula Java. Translation: Twining, leaves petiolate elliptic or obovate, the base attenuate, briefly acuminate, . fleshy veinless glabrous, peduncles extra axillary slightly thick rachis fleshy cylindrical, grooved irregularly transversely leaving a irregular groove, umbels whorled, rachis sessile, many flowered; leaflets of the corolla triangular acute glabrous, inside papilose, at the throat 5 spotted; florets of the staminal corona lower part ovate obtuse concave one nerved , marked below with a groove, acute above a little above the Gynostegium. Species with a distinct receptacle. Flowers almost two times smaller than H. carnosa. From the Island of Java. In Flora van Nederlandsch Indie Bataviae 1 (1856) 519-520. F. A. W. Miquel. 11. Hoya crassipes Turcz. Volubilis: folia petiolata e basi attenuata ima obtusa elliptica vel obovata breviter acuminata, carnosa avenia glabra: pedunculi extraaxillares breves crassi in receptaculum carnosum cylindricum, sulcatis transversis irregulariter exaratum abeuntes; umbellae in ejus vertice sessiles (?) multiflorae; corollae laciniae triangularibus acutae glabrae, intus papillosae, ad faucem 5-maculatae; coronae stamineae phyllorum pars inferior ovata concaviuscula uninervis, subtis sulco notata, superior acutiscula gynostegium paullo superans. Turcz. Bull. Mosc. XII.* 1, p. 261. Java (Zoll.). Note: This should read XXI. Translation: Twining, leaves petiolate with the base attenuate likeness obtuse elliptic or obovate briefly acuminate, fleshy veinless glabrous: peduncles outside the axles short thick with the receptacle fleshy cylindrical, grooved irregularly transversely, umbels at top sessile (?) many flowered, leaflets of the corolla triangular acute glabrous, inside papilose, with 5 spots at the collar; with the staminal corona leaflets outer angle ovate, somewhat concave, one nerved, notably grooved below, above somewhat acute, a little longe than the Gynostegium. In Systematisches Verzeichnis Family 1 (1910) 6. S. H. Koorders & Ann Schumacher.

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H. crassipes Turcz. Kletternder Strauch, bis 15 m. hoch. Mittel-Java: Perkalongan: Subah: Region I: Kds 27306  (500* 19. IV. 1897). Semarang: Kedungdjati: Region I: Kds 27217  (630* 15. IV. 1897 Ojot kapalan, Java).Ost-Java: Besuki: Puger: Region I: Kds 39891  (21 * 20. V. 1902 Auch Alcoholmaterial gesammelt), Kds 30221  (1860 * 19. III 1898. Ojot kapal, Java.) In Exkersionsflora von Java (1912). “Flora of Java” S. H. Koorders. (Key). 9b. Blätter 7-13 x 4.5 - 8.5 cm, ± obovate, oben abgerundet (Seltener apiculat), am Grunde ± keilförmig oder stumpf, kahl, oberseits an der Insertion des Blattstiels mit kleinen Drüsen versehen. Blattstiel dick, bis 1½ cm lang. Pedunculi selten bis 9 cm lang, meist 1 - 4 cm, oft einzeln, blattgegenständig oder einzeln an blattlosen Zweigen. Korolle rot. Stauch, bis 15 m hoch kletternd. Miq, l. c. 519. Mittel - und Ost-Java: in der Ebene im litchen Regenwald und im Djatiwald zerstreut (Kds. n. 30221 , 27306  usw.; Zollinger n. 2581). H. crassipes Turcz. Translation: Leaves 7 to 13 by 4.5 to 8.5 cm, more or less obovate, above not to speak of (usually apiculate) toward the base more or less otherwise obtuse, glabrous, on the surface near the insertion point of the petiole a small gland is visible. Petiole thick up to 1.5 cm long. Peduncle usually up to 9 cm long, usually only 1 - 4 cm long, often solitary, outside the axil or in the axil. Corolla red. Shrub up to 15 meters tall, climbing. From middle and east Java: in the open country and in the light rainforest and scattered near Djati forest.

Hoya crocea Tuinbouw In Tuinbouw Flora 1 (1853) 73. DeVries.

Hoya cumingiana Decaisne In Decandolle, Prodramus Syst. Veg. 8 (1844) 636. Decaisne. 15. H. cumingiana, scandens glabra, ramis foliosis, foliis ovato-cordatis obtusis subcarnosis supra glabris subtus venosis papilloso-velutinis brevi petiolatis, pedunculis plurifloris brevibus pedicellisque glabris, corollae laciniis triangularibus acutis reflexis, extrorsum glabris, introrsum papillosis, coron. stam. foliolis ovatis supra convexis marginibus revolutis angulo interiore porrecto, stigmati apiculato incumbente. Woody shrub in insula Philippinis (Cuming exsiccat. n. 1480). (v. s. in herb. Delessert.). Translation: climbing glabrous, branches leafy, leaves ovate-cordate obtuse somewhat fleshy above glabrous below conspicuously veined papilose velvety with short petioles, peduncles many flowered glabrous short pedicelled, leaflets of the corolla triangular acute reflexed, outside glabrous, inside papilose, leaflets of the staminal corona ovate above convex margins revolute, interior angle projected incumbent on the apiculate stigma. A woody shrub from the Philippine islands (Cuming’s specimen number 1480) (I have seen it in the Delessert’s herbarium). In Fleur des Serres 7 (1851) 176. 506. Hoya cumingiana, Decaisne. Liane grimpante de serre chaude, a feuilles denses et planes, a fleurs d’un jaune-verdatre groupees en ombelles axillaries presque sessiles, que se montrent dans le mois de mai.

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Cette curicuse espece, originaire des Philippines, a ete presentee par ses introducteurs MM. Veitch, a l’une des exhibitions de la societe d’horticulture de Chiswick. C’est un arbuste dresse, dont les feuilles denses, decussees, sessiles et cordees a la base, sont tres legerement pubescentes a la page inferieure et d’uet texture a peu pres parcheminee. Les fleurs peu brillantes, sont d’un jaune-verdatre rehausse par la teinte pourpre-vif de l’etiole centrale, L’espece est tres distincte de ......tes celles que l’on connait dans les jardines et, sans les fleurs, ne serait pas aisemead reconnue pour un Hoya. (Lindl. in Paxt. Fl. Gard, II, 114). Translation: Trailing liana for the greenhouse, has dense leaves and cover, has flowers of a yellow-greenish nearly grouped in umbels, axillary and sessiles, that appear in the month of May. This curious species, originally from the Philippines, has been presented by its initiators Messers. Veitch, at one of its exhibitions at the society of horticulture of Chiswick. It is a bush, with dense thick foliage, sessiles and rounded at basis, are very lightly pubescente on the bottom of the leaf and of a texture more or less parchment-like. Flowers somewhat brilliant, are a yellow-greenish heightened by the bright-crimson hue of the center of the flower. The species is very distinct from those that one knows in garden, without flowers, one would not be able to recognized it for a Hoya. (Lindl. Paxt in. Fl. Gard, II, 114). In Paxton’s Magazine of Botany 13 (1853) 263. Lindley. Hoya cumingiana . A beautiful creeper, with most foliage of rich green, producing clusters of yellowish-white flowers, about twenty in a bunch, each flower having a deep spot of purple at the base of the petals. They likewise have a delicious fragrance, which compensates for the lack of color. Messers Veitch and Son are the introducers and cultivators of it, in whose nursery we noticed it flowering in September last. In Fleur des Serres 23 (1853) 121-122. J. E. P. Hoya cumingiana, Dcne. in DC. Prodr., VIII, p. 636. — Hook. in Bot. Mag., tab. 5148 (hic iterata). — Ch. Lem. in Revue horticole de Paris. ann. 1860, p. 502 et Jardin fleur., vol. II p. 151 (miscell.) — Lindl. in Paxt. Fl. Gard., 11, gleanings, 114, n. 382, cum icone. Si modestes qu’elles soient, et celle-ci n’a pas le d’etre ambitieuse, les especes d’Hoya ont au moins les avantages d’un feuillage permanent, ferme de texture, de contour et de couleur, et servant de fond a des fleurs a la fois simples et bizarres, dans lesquelles une premiere etiole a surface bombee (corolla) encadre ou plutot sertit une etiole pentagonale (couronne des appendices staminaux) decoupee comme une gemme en brillant. Cela nous est bien connu par ce vrai joyau des serres, le classique Hoya carnosa: cela reparant moins vif et plus viole dans les fleurs de l'espece ici reproduite, ou la teinte buffle clair ou jaune verdatre du fond est rehaussee d’un lavis de pourpre au sommet des cinq pointes de la gemme staminale. C’est de Borneo ou Singapore que M. low, de Clapton, recut jadis les exemplaires de cette espece d’un genre essentellement asiatique (exception fait pour de rares representants dans l’Afrique tropicale). De leur cote, MM. Veitch l’avaient eue de leur collecteur Thomas Lobb. En le rapportant a l’Hoya Cumingiana de Decaisne, et, par consequent, aux echantillons d’herbier des Philippines, distribues par le collecteur Cuming sous le n, 1480, feu sir William Hooker fait pourtant cette reserve que la plante

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de M. low ne saurait etre dite glabre ses jeunes rameaux, ses petioles, ses pedoncules et ses calices etant revetus d’une fine pubescence. Mais, d’apres M. Decaisne lui-meme, le prototype decrit aurait les feuilles garnies en dessous de papilles veloutees, et l’on sait combien la pubescence et la glabrescence comportent, dans ce genere et ailleurs, de nuances de transition. Il est donc probable que la determination est correcte, d’autant plus que les caracteres de la fleur semblent repondre a la diagnose originale. La culture des Hoya est bien connue. On sait que, par une singularite remarquable ces plantes (l’Hoya carnosa du moins) produisent plusieures fois de suite, sur le meme pedoncule, un nouveau fascicule ou ombelle simple de supprimer, pares floraison, ces pedoncules sont la force productive n’est pas epuisee et qui pourront, sans prejudice des ombelles primipares venues sur d’autres rameaux, couvrir les rameaux anciens d’une nouvelle moisson de fleurs. Translation: So modest that they are, and this one doesn't have the to be ambitious, species of Hoya have at less advantages of a permanent foliage, farm of texture, contour and color, and serving bottom has flowers at a time simple and bizarre, in which a first blanched has rounded surface (corolla) frames either rather sets a blanches pentagonal (crown of appendixes staminal) cut up like a gem while shining. It is known us well by this true jewel of the greenhouses, the classic Hoya carnosa: it repairing less quick and more viola in flowers of the species here reproduced, or the hue buffalo color or yellow greenish some bottom is heightened of a purple lavis to the top of the five tips of gem staminal. It is Borneo or Singapore that Mr. Low, of Clapton, received copies of this species of a kind previously Asian essentellement (exception makes for rare representatives in the tropical Africa). Of quotes them, Messers Veitches had him for their collector Thomas Lobb. As bringing back it has the Cumingiana Hoya of Decaisne, and, therefore, to Curators of the Herbarium of Philippines, distributed by the collector Cuming under the n, 1480, fire sir William Hooker makes this reserve yet that the plant of Mr. Low would not know its young branches to be glabrous, stalked sexes, its stalks and its chalices stands clothed of a fine pubescence. But, of after Mr. Decaisne himself, the described prototype would have the garnished leaves below velvety papillous, and one knows the pubescence and glabrescence comportent, and how much in that generated and elsewhere, of transition nuances. It is therefore likely that the determination is correct, of as much more that characters of the flower seem to answer has the original diagnose. The culture of the Hoyas is known well. One knows that, by a remarkable oddness these plants (Hoya carnosa of the less) produce plusieures time in succession, on the same stalk, a new manual or simple umbel is suppressed, apires flowering, these stalked are the productive strength is not exhausted and that will be able to, without prejudice to primary umbels comings on other branches, to cover the ancient branches of a new crop of flowers. In Tuinbouw Flora 1 (1855) 71. DeVries. 13. Hoya cumingii* Decaisne. Eine klimmende en slingerplant, met groen-gele bloemen, en deze op bloemschermen, die bijna ongesteeld zijn. Deze plant bloeit in Mei. Wat hier zeer merkwaardig is, is de bladstelling, die altijd overkruis is. De bloemen zijn niet schon en men zou, zonder dezelve te zien, de plant niet voor eene Hoya houden. De Herren Veitch hebben haar ‘t

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eerste op eene tentoon stelling te Chiswick aangeboden. (Lindl. in Paxt. fl. gard. II. 114. Fl. des serr. VII. p. 176). Notation: * Species misspelled. Translation: . A climbing and twining-plant, with greenish-yellow flowers, and these on the flower stalk, almost stalkless. This plant blooms in May. What is here very remarkable is, that the leaf-position, these are always overlapping. The flowers his not beautiful and one shall, without to see therefor, holding a plant not in the form of a Hoya. The Mr. Veitch has here offered the first one presented to the Chiswick market. In Flora van Nederlandsch Indie Bataviae (1856). F. A. W. Miquel. 4. Hoya cumingiana Decaisn. in DC. Prodr. VIII. p. 636. Scandens, glaber, ramis foliosis, folia breviter petiolata ovato-cordatus subcarnosa, supra glabra, subtus venosa papillosovelutina, pedunculati pluriflori breves pedicellique glabri, corollae laciniae triangulares acutae reflexae, extus glabrae, intus papillosae, coronae stamineae phylla ovata, supra convexa marginibus revoluta, angulo interiore porrecto stigmati apiculato incumbente. Philippijnsche eil. (Cuming, n. 1480). Translation: Same as for Decaisne’s 1844 above. In Annales Botanices Systematicae 5 (1858) 506. W. G. Walpers “Icones Addendae”. Hoya cumingiana Decne. - DC Prodr. VIII 636 n. 15. - Paxton fl. gard. II m, 382 xylogr. fig. 192. In Botanical Magazine (1859) 5148. Hoya cumingiana. Mr. Cuming’s Hoya. Hoya cumingiana; scandens glabra ramis foliosis, folia breviter petiolata ovato-cordatus subcarnosa, supra glabra, subtus venosa papilloso-velutina, pedunculati plurifloris brevibus pedicellique glabri, corollae laciniis triangulares acutae reflexis, extus glabrae, intus papillosae, coronae stamineae phylla ovata, supra convexa marginibus revoluta, angulo interiore porrecto stigmati apiculato incumbente. Dcne. in De Cand. Hoya cumingiana. Decne in De Cand. Prodr. v. 8. p. 636. Flowering specimens of the pretty Hoya here figured were communicated to us by Mr. Lowe, of the Clapton Nursery, who received the plant from Eastern Archipelago, gathered either in Singapore or in Borneo. It probably is a native of both those islands, and the Malayan Islands generally, being unquestionably the H. Cumingiana of Decaisne in De Candolle’s Prodromus,’ from the Philippine Islands (n. 1480 of Mr. Cuming’s distributed specimens). There are, indeed some trifling discrepancies between the character in the ‘Prodromus’ and our specimens, almost wholly, however, depending on the more or less pubescent character, a circumstance extremely liable to vary. Description: A climber, with terete, green branches, slightly pubescent, as are the very short petioles, the peduncles, and the calyx. Leaves coriaceous, elliptical-ovate, cordate at the base, suddenly acute at the point, very indistinctly penninerved. Peduncles a little supra-axillary, three-quarters of an inch long, bearing an umbel of flowers, which are drooping; pedicels an inch long, slender. Calyx of 5 parts, the segments oblong-ovate, obtuse. Flowers tawny-yellow. Corolla with its five obtuse lobes reflexed. Staminal crown purple in the centre. Translation: trailing glabrous stems with leaves, leaves shortly petiolate ovate cordate somewhat fleshy, above glabrous, veined papilose-velvety, with a peduncle with very

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short glabrous pedicels, flaps of the corolla triangular acute reflexed, outside glabrous inside papilose, leaflets of the staminal corona ovate, above convex margines revolute, interior angle apiculate incumbent upon the stigma. In Revue Horticole de Paris Ann. (1860) 502. Hoya cumingiana, Decaisne; De Candolle, Prodrome, VIII, 636, Botanical Magazine, t 5148, November 1859. Decouvert dans les iles Philippines, par M. Cuming (1836 -1844) ce joli Hoya a ete retrouve recemment par M. Low fils aux ....rons de Singapour ou dans il’ la de Borneo, d’ou il l’a envove .... tant d’autres excellentes nouveautes, a M. Low pere, horticulture a Clapton. Bien que les branches, les petioles, les peduncles et les calyce du Hoya Cumingiana soient legerement pubescents, M. W. ...kerr n’hesite pas a rapporter cette plante a celle determinee .... par M. Decaisne, que dit tous ces organes glabres; la pubescence est du reste, comme le fait observer M. Hooker, fort sujelte a varier. Cette plante est grimpante comme ses congeneres; ses feuilles sont ovees-elliptiques, cordiformes a la base, brusquem... aigues au sommet, coriaces, indistinctement penninerves les pedoncules , inseres un peu au-dessus de l’aisselle, sont long.... plus de 0m.02 et portent une ombelle de fleurs penchees, s.....neus par des pedicelles equilongs; cells sont conformees com.... dans le genre, et d’un jaune de buffle, legerement macule de cramoisi sur l’appareil genital. On ne nous dit pas qu’elles ..... odorantes, ce qui est assez probable. Le Hoya Cumingiana reclame la serre chaude. Translation: this pretty Hoya was recovers recently by Mr. Low son to them... .rons of Singapore or in the island of Borneo, of or he has him envove.... so many other excellent novelties, has Mr. Low father, horticulture of Clapton. Although branches, leafstalks, peduncles and calyces of the Hoya Cumingiana are slightly pubescent, Mr. Walker doesn't hesitate to bring back this plant as the one determined by Mr. Decaisne, that so-called all these glabrous organs; the pubescence is the remainder, as the fact observed by Mr. Hooker, as strongly subject to variability. This plant is trailing as its fellows; its leaves are ovate-elliptic, with cordiforme basis and has the apex shortly sharp at the top, tough, indistinctly penninerved, the stalk inserted a little above of the axiles, are long.... more of 0m .02 and carry an umbel of flowers leaned, s... ..neus by pedicels equally long; cells related, conformed com.... in the kind, and of a buff-yellow, corona stained crimson with some appeal. One ne'noue doesn't say that they..... odornates, what is likely enough. The Hoya Cumingiana reclaims the hot greenhouse. In The Garden 18 (1880) 355. “The Hoyas or Honey Plants” Z. B.. Hoya cumingiana It is somewhat distinct from the majority of cultivated hoyas, being stiff and erect rather than climbing, as most of the Hoyas are. Forms a bush - indeed something similar to what is seen in the allied Cyrtoceras reflexum, The flowers, too, are not unlike those of the Cyrtoceras, but the colors are different, as in the Hoya they are yellowish-green, with a brown coronet. It is native of the Philippines, and was introduced by Messers Veitch about thirty years ago. The annealed woodcut shows the flowers of this species. Its distinctness in habit is perhaps the recommendation it has when compared with other Hoyas of which there are many beautiful varied kinds, though from some cause or other many are not much cultivated. The best known is Hoya carnosa, which despite its commonness, is one of then finest stove climbers we possess. Flowers in almost any position where warmth, moisture and a little light are afforded it, clinging by means of its

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stem-roots to a damp wall as ......, and never failing to produce com...... even-formed umbels of flesh-coloured, waxy ......ing flowers. etc. re. other species. In Paxton’s Flower Garden 2 (1883) 114 382. Hoya cumingiana. Decaisne. A stove shrub with dense flat leaves and short axillary umbels of greenish-yellow flowers. Native of the Philippines. Blossoms in May and June. Introduced by Messrs. Veitch and Son. (Fig. 12.) At one of the exhibitions in the garden of Horticultural Society this novelty was produced by Messrs. Veitch and Son. It is an erect bush with closely packed decussating sessile cordate leaves, very slightly downy beneath, and of a somewhat parchment-like consistency. The flowers are destitute of gay colours, the principle tint being yellowish- green, relived by a coronet of rich purplish brown. It is very distinct from any of the other species in cultivation, and before flowering would not be taken for a Hoya at all. In Dictionary of Gardening 2 (1885). G. Nicholson. H. Cumingiana (Cuming’s) * fl greenish-yellow or white, with a coronet of rich purplish-brown; umbels axillary, Short. Spring and summer. l. closely packed, flat, sessile, cordate, slightly downy beneath. Malay Archipelago. (B. M. 5148) In Plantas Filipinas (1886) 189. Vidal. H. cumingiana, Dcne (Miq. l.c. 517.) Cum. 1480 I. Mindoro. In Journal of Botany, British and Foreign 36 (1898) 416. James Britten. H. cumminghami (sic) is given by Teijsmann & Binnendyck (in Tijdschr. Ned, Ind. XXV. 408, and in Cat. Hort. Bogor. 130) as a name under which H. excavata had been cultivated in the Utrecht Gardens. It is not in Ind. Kew. In Philippine Journal of Science 1 (1906) 119. R. Schlechter. 1. Hoya cumingiana Decne. l. c. 636. (311 Whitford) May. An Epiphyte in forests at 700m. Endemic. This species is abundant in the region of Pinus insularis, northern Luzon, where it is always found on boulders, outcroppings, and cliffs. In Philippine Journal of Science 3 (1908) 351-352. “Enumeration of Philippine Flowering Plants” E. D. Merrill. Hoya cumingiana Decne. in DC. Prodr. 8 (1844) 636; Miq. Fl. Ind. Bat. 2 (1857) 517; F.- Vill. Novis. App. (1880) 135; Vidal Phan. Cuming. Philip. (1885) 127, Rev. Pl. Vasc. Filip. (1886) 189; Merr. in Philip. Jour. Sci. 1 (1906) Suppl. 119, 5 (1910) Bot. 379. Luzon (Benguet, Bantoc, Bataan, Rizal, Camarines), Mindoro, Ticao, Palawan, Cuming 1480, Merrill 6325, 9435, 11562, Elmer 6419, Whitford 311, B.S. 26221, 3256, 22214 Ramos, 8803 McGregor. Chiefly on limestone cliffs and boulders in open places from sea level to an altitude of 1600 m. Endemic. Local names Alisip (Ig.); barobagol (Bik.); laklakipan (Bon.); pilpilak (Bon.); pirpirak (Ilk.); puki (Bon.). In Philippine Journal of Science 5 (1910) 379. 1. H. cumingiana Dcne. in DC Prodr. 8 (1844) 636. On boulders in stream depressions in the pine region, altitude 1,200 m, C. M.

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Z. 16694, Merrill 6352. Endemic; very closely allied to H. densifolia Turcz., of Java, and the later species perhaps not specifically distinct. See Backer lc. Bogor. 3 (1908) 157. In Philippine Journal of Science 18 (1921). Merrill. Hoya cumingiana Decne. in DC Prodr. 8 (1844) 636; Miq. Fl. Ind. Bat. 2 (1857) 517; F. - Vill. Novis App. (1880) 135; Vidal Phan. Cuming. Philip. (1885) 127, Rev. Pl. Vasc. Filip. (1886) 189; Merr. in Philip. Jour. Sci. 1 (1906) Supp. 119, 5 (1910) Bot. 379. Luzon (Benguet, Bontoc, Bataan, Rizal, Camarines), Mindoro, Ticao, Palawan, Cumming 1480, Merrill 6352, 9435, 11562, Elmer 6419, Whitford 811, B.S. 26221, 3256, 22214 Ramos, 8803 McGregor. Chiefly on limestone cliffs and boulders in open places from sea level to an altitude of 1600 m. Endemic. Local names: Alisip (Ig.); barobagol (Bik.); laklakipan (Bon.); pilpilak (Bon.); pirpirak (Ilk.); puki (Bon.). In Sunyatsenia 3 (1936) 176. Tsiang. Hoya cumingiana Decaisne in DC. Prodr. 8:636. 1844; Gardening (London) 7: 609. 1885; Merrill, Enum. Philip. Fl. Pl. 3: 351. 1923. Philippine Islands: Mayon volcano, Albay Province, Luzon, M. Ramos & G. Edano 75716, Sept., 1928. Endemic. The plant matches exactly the illustration in Gardening. Series 2. Toxostemma Tsiang. Angulus exterior loborum coronae rotundatus. In Leaflets of Philippine Botany 10 (1938) 3581. A.D.E. Elmer. Hoya cumingiana Dcne. Number 6419 collected by Elmer at Baguio. - This species bears quite a characteristic foliage and was discovered by Cuming on Mindoro. In Dictionary of the Royal Horticulture Society (1965). Hoya cumingiana. l. close, flat, cordate, slightly downy beneath, sessile. fl. greenish-yellow or white; corona rich purplish-brown; umbels short, axillary. Spring, summer. Malaya. (B.M. 5148). In Philippine Hoya Species 1 (1989) 42-43. R. D. Kloppenburg. Hoya cumingiana Decaisne, in D.C. Prodromus, Vol. 8, p. 636 (1844) Ins. Philipp. Type: Cumin # 1480. Section: Plocostemma. A very adaptive species preferring limestone locations, outcroppings, cliffs, boulders, and roadcuts in open places from sea level to 2000 m. It is a scandent shrub with both pendulous and upward twining stems, glabrous leafy branches, green becoming woody, leaves ovate cordate. Variable in leaf size, shape and pubescence. Peduncle and pedicel length are also variable, but it is easily recognizable and distinctive. Leaves in general glabrous above, venous, papillose-velvety below, shortly petiolate. Petiole 0.5 + cm. long. Peduncles 2.8 + cm. long thickening toward the rachis. Pedicels variable in length 1.5-2.5 cm. long, all green, mostly glabrous except in the forms approaching the densifolia types. Few flowers per umbel. Calyx lobes overlapping at base, apex narrowly rounded almost mucronate, glabrous except for very sparse hairs on outer surface, curved downward (reflex), ligule at base of each lobe. Corolla lobes triangular acute and reflexed, outside glabrous, inside papillose. Corona raised in the center, anther appendages exceeding inner lobe which is extended forward. Scales slightly concave above, outer lobe narrowly ovate. Lobes with side protrusions and channeled below. Stigma is capitate and well developed. Pollinia 0.06 cm. long x 0.02 cm. wide, keeled and with adjacent vacuole, outer lobe rounded. Translators sort and narrow. Retinacula long,

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narrow 0.03 cm. x 0.01 cm. with divided outer lobe. Luzon (Benguet, Bontoc, Bataan, Rizal, Camarines), Mindoro, Ticao, Palawan, Mayon Volcano, Albay Prov, Luzon et. al. References: Decaisne, DC Prodramus. Vol. 8 p.636 (1844). Fleur des Serres, Vol. 7, p. 176 (1851-1852). Fleur des Serres, Vol. 23, pp. 121-122 (1852-1853). DeVries Tuinbouw Flora, Vol. 1, p. 71 (1855). Zollinger & Miquel, Fl. Ind. Bat., Vol. 2 p. 518 (1856). Icones Addendae Annales Bot. Sys. Vol. 5 p. 506. Curtis's Bot. Mag. tab. 5148, (1859). Revue Horticole de Paris, p. 502 (1860). Drawing & detail FDS, Vol. 23, p. 120 (1880). F. Vill. Novis App. p. 135 (1880). The Garden p. 55 Jan. 20 (1883). Vidal Phan. Cuming Philip. p. 127 (1885). Geo. Nicholson Dict. of Gdng. (1885). Rev. Pl. Vasc. Filip. p. 189 (1886). Jour. of Bot. Vol. 36 p. 416 (1896). Philip. Jour. Sci. Vol. 1. Supp. 119, 5 (1911). Philip. Jour. Sci. Vol. 5, p. 379, Nov. (1910). Enum. Philip. Pl. Vol. 3, pp. 351-352 (1923). Sunyatsenia Vol. 3 #2/3, p. 176 (1936). Leaflets of Philip. Bot. Vol. 10, Art. 131 p. 3581 (1938). Dict. RHS.. Paxton's Fl. Gdn. Vol. 2, p. 114 n 382. Paxton's Mag. of Bot. xiii, p. 263. Watercolor by Daniel Girard Elliot, ornithologist at New York Bot. Gdn. Library. The Hoyan Vol. 8 #3 pp. 40-41 1987. Herbarium Sheets: Bartlett #14356 1935 (A); Celestino #4401 1948 (PNH,A); C.M.Z. #16074 ; Cuming #1480 ; Elmer #6419 1909; Hernandez #19059 1966 (CAHUP); Kienholz #134 1924 (UC); Loher #14291 1913 (UC); Clemens #5888 1925 (UC); Merrill #6352, #9435 (A), #11562 ; Mcgregor #8803 ; Ramos #26221 1916, #3256, #22214 ; Ramos & Edano #75716 ; Rimando #18681 ; Sijor #27431 1919 (UC); Sinclair #9757 1958 (PNH, SING).Lindl #382; Witford #311 1906. In The Hoya handbook (1992) 69. D. Kloppenburg & A. Wayman. Hoya cumingiana Decaisne. In spite of forest destruction and habitat reduction, this is one species that I feel will survive. The plant pictured here came to me from Professor Juan V. Pancho, who collected it in 1981 from in Northern Luzon in the Philippines. It was taken from a road cut through a limestone area, attesting to the fact that this species does not need a primary forest to survive and propagate. It readily sets seed pods and has a high percent of viable seed which grow vigorously. In this respect it is somewhat unusual among hoya species. This is not to say that there are not other hoya species that produce pods, but most are not as prolific in this respect as Hoya cumingiana is. Its ability to reseed itself even in disturbed areas seems to assure its permanence as a species. This species is not a vine as most hoyas are, but rather has a bushy type growth. As its branches become long they bow over and become pendant unless it finds support from a nearby plant or other surface. As pictured this plant has a lot of closely formed leaves, close also to the stem. The flower cluster hangs down from the arching stems and are very waxy and showy with their contrasting colors. Since this species like many others prefers coral and other limestone areas, it is advisable to include a source of calcium in the potting mix. Some crushed coral, broken up limestone, even gypsum board from construction can act as a calcium source. In Asklepios 58 (1993) 17-19. P. I. Forster & D. J. Liddle. Asclepiad Profile. 3: Hoya cumingiana Decaisne. Hoya cumingiana Decne. in DC., Prodr.8: 636 (1844); Hook., Bot. Mag. 8S: I. 5148 (1859); J. D. Hook., Fl. Brit. India 3:692 (1882); Merrill, Philipp. J. Sci. 1, suppl.

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119 (1911); Merrill, Enum. Philipp. Flow. Pl.3: 351-352 (1923); Tsiang, Sunyatsenia 3: 176 (1936); Elmer, Leafl. Philipp. Bot. 10: 3581 (1938); C.M. Burton, Hoyan 8: 40-41 (1987); Kloppenburg, Philipp. noes 42- 160 43(1991); C. M. Burton, Hoyan l3(3), part2: 17(1992). Type: Philippines Prov. Batanes, Luzon, *1811, H. Cuming 1480 (iso: K [photo at BRI!). Hoya densifolia Turcz., Bull. Soc. Nat. Mosc. 21: 261 (1848). Type: Java. 16 fan 1845, Zollinger 2620 (iso: BO!). Hoya golamcoiana Kloppenburg, Fraterna, 3rd quarter 1991: 2 (1991). Type: "sheet # 41930 El Nido Rest Area Palawan, Philippines found by Professor Juan V. Pancho 15 August 1988. Holotypus: CAHUP." [n.v.] Epiphytic subshrub, latex white. Stems cylindrical, with sparse indumentum; internodes up to 22 mm long and 4 mm diameter. Leaves petiolate; lamina elliptic to orbicular, 30-55 mm long, 20-30 mm wide, discolorous, glabrous; upper surface mid-green, venation obscure; lower surface pale-green, secondary veins 9- 10 per side of midrib and visible, tertiary venation just visible; tip acute; base cordate; petiole 2-3 mm long, c. 1 mm diameter, with sparse indumentum. Cyme umbelliform to racemiform, up to 20 mm long; peduncle 12-17 mm long, c. 1 mm diameter, with sparse indumentum; bracts lanceolate, 0.5-1 mm long, O.4-0.5 mm wide, with sparse indumentum. Flowers 15-20 in umbel, c. 9 mm long and 10 mm diameter; pedicels 17-22 mm long, O.5-.6mm diameter, glabrous. Sepals lanceolate, 2-2.3 mm long, 0.9-1.1 mm wide, with scattered indumentum. Corolla reflexed, upper surface with short dense indumentum, lower surface glabrous; tube c.3 mm long and 3 mm diameter; lobes lanceolate to triangular, 4~.3 mm long, 2.5-3.8 mm wide. Staminal corona 2.5-3 mm long, 6-7 mm diameter, inserted on staminal column well above surface of corolla; each lobe 2.5-3.7 mm long, 1 -1.7 mm broad, 2.8-3.1 mm high, inner edge spathulate and held erect above style-head, outer edge upturned and held slightly lower than inner. Staminal column 3-3.5 mm long, c. 2 mm diameter; alar fissure 1-1.2 mm long; anther appendages triangular, c. 1 mm long and 1 mm wide. Pollinaria 0.5-0.55 mm long, 0.5-0.64 mm wide; pollinia oblong, 0.55-0.58 mm long, 0.23-0.26 mm long, with pellucid germination mouth on outer edge; corpusculum ovate-oblong, e.0.27 mm long, O.13-0.14 mm wide; caudicles c. 0.09 mm long and 0.02 mm wide, unwinged. Follicles and seed not seen. Fig. 1. Specimens Examined: Philippines: Aguja Point, Burias, viii.1922, Merrill 11562 (BRI). Java: G. Muriah above Djorang, xii.1951, Kern 8622 (BO); G. Goentoer Gover Garech, xi.1912, Backer 5222 (BO); N. Kaambanean, Koorders 39688 (BO); [unlocalised] 1915, Backer 21006 (BO); ii.l914, Lorzing 1018 (BO); Feb 1907, Backer [BO111021] (BO); Nov 1919, van Sloolen 226 (BO), iii.l931, Docters van Leeuwen 13878 (BO). Cultivated (Australia): Strathpine, Brisbane, ii.l989, Forster 4965 (BRI); Emerald Creek, Mareeba, vi. l991, Liddle IML168 (BRI). (Germany): Botanischen Garten Berlin-Dahlen, vii. 1990, Cebu 27101 (B). Distribution and Habitat: H. cumingiana occurs in eastern Malasia in the Philippines and in Java and could be expected to exist in other islands in the region. Plants grow as epiphytes in rainforest trees or lithophytically on limestone rock outcrops (D. M. Cumming, personal communication) from sea-level to 1600m. (Merrill 1923). Notes: H. cumingiana is a distinctive species with its non-twining habit. Although this species is widely cultivated, few of the clones have any locality data and attempts should

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be made to reintroduce material from the wild to allow an assessment of the variation in this plant. The species H. densifolia Turcz. was based on a collection from Java and although the name has not been applied to material in cultivation, specimens at Herbarium Bogoriense (BO) were identified as this species. Burton (1992) correctly concluded that H. densifolia was conspecific with H. cumingiana, placing the former in the synonymy of latter earlier name. Burton also placed H. golamcoiana Kloppenburg (1991) in synonymy with H. cumingiana and although we have not seen the type specimen, it is probable that this poorly described species is also synonymous. Kloppenburg (1991) states: "This species differs from other known Hoya species. Its [sic] foliage blades are similar to H. cumingiana Decaisne but it is petiolate and on a dangling plant. "The illustration accompanying Kloppenburg's article shows a plant of H. cumingiana, although no voucher details for the plate are given. Cultivation: This species is easily grown either as a pot plans or in a hanging basket and should tee treated in a manner suitable for most epiphytic Hoyas. Plants are easily propagated by rooting cuttings of older stems. Acknowledgements: Dr. P.S. Short (MEL), while Australian Botanical Liaison Officer at Kew, England located and photographed type material of H. cumingiana. The Directors/Curators of Herbarium Bogoriense (BO) and Singapore Botanic Gardens (SING) allowed access to the herbarium collections in their care. References: Burton, C. M. (19921 Hoya densifolia Bl.. Hoyan 13(3) part 2:17. Kloppenburg, D. (1991). 4 new Hoya species from the Philippine Islands Fraterna 3 rd quarter 1991:1 -4 + unpaged plates. Merrill, E. D. (1923). An Enumeration of Philippine Flowering Plants. Vol. 3:351-352. (Bureau of Printing Manila). * All the literature shows Sheet # 1480 as the Type specimen. In Addition this was collected from the Island of Mindoro. See The Type description above in DeCandolle’s Prodr. 1844.

Hoya Curtisii King & Gamble In Journal of the Royal Asiatic Society, Bengal Branch 74/2 (1908). “Flora of the Malayan Peninsula” King & 563 Gamble. 3. Hoya Curtisii, King & Gamble, n. sp.. A twining small epiphytic undershrub, rooting at almost all the nodes on the bark of trees or on rocks; branches slender pubescent. Leaves sessile, very thick; orbicular, abruptly triangular-acuminate at apex, rounded or sometimes orbicular very slightly attenuate at the base; both surfaces scarious-puncticulate when dry; margine thickened ; .4 to .7 in. long, .4 to .6 in broad; midrib when dry faint, other nerves invisible; petiole none. Umbels sessile. terminal and axillary, 10 to 16 flowered, on a .25 in long tubercular rachis; pedicels slender, pubescent, .75 to 1 in. long; flowers rather large, Calyx with very short tube or almost none; lobes oblong, ciliate, villous without, membranaceous, .1 in. long, much recurved; scales minute. Corolla also much recurved; tube very short, with an annular spreading ring (or corolline corona ?) a little above the base; lobes spathulate, acute at the tip, minutely scurfy-puberulose, .25 in long. Corona: corolline (see above) , annular; staminal, of 5 cuculate processes nearly globose when closed, but formed of 3 lobes, the two side ones rounded, the upper ovate, the three enclosing the cavity, and the

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whole corona much overtopping the staminal column. Staminal-column short and shortly stipitate, attached at the base of the corolla-tube; anther cells much divergent; appendages ovate, membranaceous, dentate; pollen-masses oblong-obtuse, about .03 in. long, attached by oblong fleshy thick caudicles to the conical pollen-carriers. Style-apex shortly 5-angled, somewhat depressed; tip bifid, very short. Fruit not known. Perak: at Waterloo, 2000 ft., Curtis 2894. Corona-processes with lower lobe globose, hollow; upper shorter-curved; corolla-tube with a broad annular ring (corolline corona ?) (Sect. III Kloiophora 3. H. Curtisii In Flora of the Malaya Peninsula 4 (1923) 394. H. Ridley. (15). H. Curtisii King & Gamble l.c. 563. Slender epiphyte creeping and rooting at nodes. Leaves orbicular, abruptly acute, base round sessile. .4 to .7 in. long about as wide. Umbels terminal and axillary, 10 to 16 flowered on a sessile rachis, .25 in. long, pubescent . Calyx-lobes oblong villose. Corolla .5 in across, lobes deflexed, spathulate, acute. An annular corona, surrounding a staminal corona of 5 hooded processes of 3 lobes, side ones round, upper ovate, enclosing a cavity and overtopping the staminal column. Hab. Perak Waterloo at 2000 ft. altitude (Curtis) Very rare. In Malayan Nature Journal 30 (1978) 505 Rintz. 15) Hoya curtisii King & Gamble. J.A.S. Beng. IV (1903) 563. Type: Malaysia, Perak, Waterloo Estate, Curtis 2894 (Sing). - Fig. 19. Distinguishing Features: Stems thin hirsute. Leaves fleshy, oval, apically cuspidate, surface rugulose; c. 1.5 cm. long and wide. Peduncle horizontal, rigid, c, 5 mm long. Umbel negatively-geotropic, convex with rigid, uniform pedicels c. 2.5 cm long; pedicles hirsute; 1-30 flowers. Corolla lobes reflexed, pubescent inside except glabrous at the tips of the lobes; c. 1 cm long by 7 mm diam. Corona subtended by an inflexed annulus; lower lobe inflated, globose. Corpuscle small. Ecology: Endemic to Malaysia and known only from a single collection near Taiping, Perak at c. 700 m. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 8687 “The Genus Hoya in Thailand” O. Thaithong. 3. H. curtisii King & Gamble, J. As. Soc. Beng., 4(1903) 563; FMP 2 (1923) 394-395; Malay. Nat. J. 30 (1978) 505, Fig. 19. Occurrence: (PEN): Chumphon. Notes: Leaves usually variegated, flower 1.8-2 cm dia.: corolla coloured, outside sometimes pinkish purple, reflexed in full bloom; inner end of coronal scales purple, not covering the yellow anther appendages.

Hoya cystiantha Schlechter In Flora of Indiae Batavae 2 (1856) 516. H. Zollinger & F. A. W. Miquel. 2. Cystidianthus laurifolius Bl. Mus. bot. I. p. 57. Folia e basi vulgo acuta obverse oblonga vel lanceolata acuminata, coriacea venosa, 3—4 ½ poll. longa; Flores mediocres, iis praeced. simillimi, sed ½ minores, limbi laciniis erectis triangulari-ovatis acutis. Sumatra (Praetorius, Korthals). Translation: Leaves at bases generally acute, obverse oblong or lanceolate acuminate, leathery veined, 3-4 ½ inches long; flowers middle-sized, it is very similar to the preceding (H. campanulata) but ½ smaller, lobes erect triangular-ovate, acute.

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In Annales Botanices Systematicae 3 (1848-1859) 63. W. G. Walpers. 2. PH.. (Cystidianthus) Laurifolium Blume, Mus. bot. Lugd. Batav. 57. no, 141. t —Foliis oblongis vel lanceolatis utrinque subacutis, coriaceis venosis, floribus mediocribus.— Habitat in sylvis insulae Sumatrae. Translation: Leaves oblong or lanceolate, both ends somewhat acute, leathery, veined; flowers medium sized. Habitat: in trees on the island of Sumatra. In Museum Botanicum Lugduno-Batavum 1 (1849) 57. C. L. Blume. 141. Cystidianthus laurifolius. Bl.: foliis oblongis v. lanceolatis utrinque subacutis coriaceis venosis; floribus mediocribus. Habitu plane cum C. campanulato conveniens; ramis teretibus tenacissimis, sub epidermide fibras copiosas subtiles sericeas, fortasse usibus technicis utiles, continentibus, varie tortis, glabriusculis v. in nodis nonnihil tumidis, subpubescentibus. Petioli 4—5 lin., compresso-teretes, supra canaliculati, pube. rare appressa. Folia Lauri nobilis simillima, 3—4 poll., 1 1/3—2 poll. lata, utrinque subacuta v. apice in acumen breve obtusiusculum angustata, supra petiolum glandula obtusa aucta, coriacea, glabra, venosa, siccando reticulato-venosa, nervo medio supra canaliculato, subtus, perinde ac venae arcuatae, prominulo. Umbellae interpetiolares et terminales, solitariae, simplices, brevissime pedunculatae. Pedunculus 3—4 1in., apice incrassatis et bracteolis parvis latoovatis ciliatis obsessus. Pedicelli 10—14 lin., graciles, glabri. Calycis parvi laciniae ovato-oblongae, obtusae, membranaceae, ciliatae. Corolla qualis C. campanulati, sed dimidio brevior, limbi laciniis erectis triangulari-ovatis acutis.—In sylvis Sumatrae. Translation: leaves oblong or lanceolate both ends acute leathery veined; flowers medium size. Appearance distinctly agreeing with C. campanulata, branches round very tenacious, below epidermal fibers somewhat delicately sericeous, very strong beneficial used techniques, held together, very twisted, slightly glabrous and at the nodes not at all thickened somewhat pubescent. Petiole 4 to 5 lines, compressed-round, grooved above downy rarely appressed. Leaves like Lauris nobilis, 3 to 4 inches, 1 1/3 to 2 inches wide, both ends somewhat acute and apex at acumen short nearly obtuse narrow, above on the petiole a obtuse acute gland, leathery, glabrous, veined, secondary reticulate veins, central nerve channeled above, below, in a like manner curved and veined prominently. Umbels between the petioles and terminal, solitary, simple, shortly pedunculate. Peduncles 3 to 4 lines, apex thickened and bracteated, small broad ovate ciliate obtuse. Pedicels 10 to 14 lines, narrow, glabrous. Calyx small leaflets ovate-oblong, obtuse, membranous, ciliate. Corolla like C. campanulatum, but smaller by half, limbs of the flaps erect triangular-oval acute. It lives in the forests of Sumatra. In Kew Bulletin (1925) 85 H. N. Ridley. Physostelma laurifolia Ridley. Only some sprays of flowers were sent of this specimen, which I take to be the little-known Cystidianthus laurifolius Bl. which is hardly fully described and of which I have seen no specimens. The flowers are described as half the size of those of P. campanualtum, and the sepals are oblong, blunt and ciliate, which agrees with Blume’s description, but tile corolla-lobes can hardly be described as acute. The flowers of Brook’s specimen are crowded in a head about 30, on slender pedicels 3 cm. long; they are described as very

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pale brown. A specimen of Wray's collected in Perak at Relau Tujor, 4014, very closely resembles these specimens, but Gamble has passed it as P. campanulatum. Lubok Tanudai, at 9000 ft., flowers very pale brown, no number. In Botanische Jahrbücher 50 (1913) 127. R. Schlechter. H. campanulata Decne Kommt auf der malayischen Halbinsel und Java vor und H. cystiantha Schltr. (Cystianthus laurifolius Bl.) ist sumatranisch. Translation: H. campanulata exists on the Malayan peninsula and on Java, and H. cystiantha Schlechter (Cystianthus laurifolius Blume) is from Sumatra. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. cystiantha Schlechter (Type of Cystianthus laurifolius Blume) – 15.

Hoya dalrympleana F. Mueller In Encyclopedia of Australian Plants suitable for cultivation (1990) 386. W. Roger Elliot & David L. Jones. Hoya dalrympleana F. Muell. = H. australis R. Br. ex Trail.

Hoya darwinii Loher In Gardeners Chronicle 47 (1910) 66. Loher. Hoya Darwinii, n. sp., sect Euhoya. — Ramosa, ramulis glabris; folia dimorpha, altera normalia elliptica oblonga (15-20 cm. longa, 5-6 lata) glabra, coriacea nervis non conspicuis, breviter petiolata; altera in aci abbreviata per 2-3 paria recurvata et demum capsulam globosam (marsupium), 4-6 localarem. bene clausam, formicis inhabitatam formantia; capsula longe persistens, indurescens, demum fuscam; umbellae multiflorae; flores magni (2 cm. diam.) glaberrimi nitidi, corolla profunde lobata tubos perbrevi; lobi cordata-ovata, reflexi purpureo-rosei; coronae folia erecta, magna triquetro-conica, albida. subtus usque ad basin sulcata; antherae margine papyraceo denticulato stigmati multico incumbentes; pollinia subcompressa in glandula rhomboidea fusca fere sessilia. Habit. montes Luzoniae. Floresc. Mart.-April. At the top of page 66: Hoya Darwinii. This plant is one of the most interesting Hoyas known to me, for it has dimorphous leaves. Flowers are pink coloured and very large for the genus. I take special pleasure in naming it in honor of Charles Darwin in the centenary year. H. Loher. Translation: Much branched, branches glabrous, leaves dimorphic, the normal ones elliptic oblong (15-20 cm. long, 5-6 cm. wide) glabrous, leathery nerves non conspicuous, briefly petiolate, the second type with edges shortened all over either curved backwards or recurved 2-3 times and at last forming a globose capsule, (pouch) with 4-6 cavities, well closed, inhabited by ants, capsule long persistent, becoming at length, dark in color; umbels many flowered, flowers large (2 cm. in diameter) very glabrous, shinny, corolla deeply lobed; the tube very short; the lobes cordate-ovate, reflexed., purple pink; leaflets of the corona erect, large triangular conic, white. on the lower surface grooved near the base; margines of the anthers papery toothed stigmas blunt, incumbent; pollinia somewhat compressed with glands rhomboid dark colored nearly sessile. Habitat: Mountains of Luzon. Flowering in March to April.

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In Repertorium Specierum Novarum 11 (1912) 96. Leveille. Hoya (sect. Euhoya) Darwinii Loher, l. c., p. 66. — Ramosa, ramulis glabris; folia dimorpha, altera normalia elliptica oblonga (15-20 cm longa, 5-6 cm lata) glabra, coriacea nervis non conspicuis, breviter petiolata; altera in aci abbreviata per 2-3 paria recurvata et demum capsulam globosam (marsupium), 4-6 localarem. bene clausam, formicis inhabitatam formantia; capsula longe persistens, indurescens, demum fuscam; umbellae multiflorae; flores magni (2 cm diam.) glaberrimi nitidi, corolla profunde lobata tubos perbrevi; lobi cordata-ovata, reflexi purpureo-rosei; coronae folia erecta, magna triquetro-conica, albida. subtus usque ad basin sulcata; antherae margine papyraceo denticulato stigmati multico incumbentes; pollinia subcompressa in glandula rhomboidea fusca fere sessilia. Montes Luzoniae. Floresc. Mart.-April. Translation: Same as Type description above. In An Enumeration of Philippine Flowering Plants. (also as An Enum. Philippine Plants) 3 (1923) 352. E. D. Merrill. Hoya darwinii Loher in Gard. Churn. III 47 (1910) 66, Fedde Repert. 11 (1912) 96. Luzon (Rizal, Tayabas), Panay, Whitford 790, B. S. 35522 Ramos & Edano. In forests at low or medium altitudes. Endemic. Local name: Kilitkit (Tag.). In Hortus (1976) 574. Hoya darwinii Loher. Glabrous, climbing vine; lvs. obovate to oblanceolate, to 3 ½ in. long, acuminate; umbels loosely few-fld.; corona lobes strongly reflexed, to ½ in. long, whitish with purple base, corona lobes to ¼ in. long, whitish. Philippine Is. In The new York Botanical Garden Illustrated Encyclopedia of Horticulture 5 (1981) 1726. H. darwinii, an endemic of the Philippine Islands, is a hairless vine with oblong to obovate, prominently-veined leaves up to 3.5 inches long. Its upward-facing flowers in loose umbels of rather few, have ½ -inch-long, backward pointing, whitish to rose pink petals with purple bases. In Tropica 4 (1992) 1020. A. B. Graff. Hoya darwinii (Philippines); climbing vine with thin wiry stems, and distant internodes bearing oblong-ovate recurved leaves; stunning cupped 3 cm flowers with purple center. Occasionally with pouched leaves as in Dischidia. In Philippine Hoya Species 3rd Edition (1996) 63-64. R. D. Kloppenburg. Hoya darwinii Loher in Gardener's Chronicle 47 (1910) 66 (3rd Series) Type: 14574, Loher (UC). Mountains of Luzon flowering in March-April 1909. Section Rudimantalia Kloppenburg. Key #35 A much branched, glabrous vine with dimorphic leaves. Normal leaves elliptic oblong, 15-20 cm. long, 5-6 cm. wide, glabrous, leathery, with inconspicuous nerves, briefly petiolate; other leaves bullate, a globose pouch of 4-6 cavities, well closed. (ants habitat). Pedicel glabrous 4.0 cm. long 0.2 cm. in diameter. Calyx broadly ovate 0.24 cm. tall x 0.26 at broadest just below middle , edges ciliate ragged and hyaline, ligules present on a ring inside calyx proper. Corolla deeply lobed, salmon pink, lobes cordate ovate, 2

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cm. diameter, 3.4 cm. flat, reflexed, glabrous, shiny, collar very short. Corona lobes erect, large triangular, conical, below sulcate nearly to the base, column and underside of scale to channel covered with silky matted hairs, inner angles touching in center and covering the stigma head. Anther wings swollen. Stigma blunt, incumbent. Pollinia compressed, broad, keeled only a short distance, almost sessile, broadly rounded at apex, narrowly rounded at base, retinaculum plainly visible, broad rounded head, broader at wings, sharp bifid apex, hyaline. Luzon (Rizal, Tayabas) Panay. Tagalog name "Kalitkit". References: Fedde Repertorium 11, #418 (1912) 96; Enumeration of Philippine Plants 3 (1923) 352; US Nat. Mus., Plants of the Philippine Islands (5 pictures) Photos from Munich Bot. Gdn. by T. H. Haas; The N.Y. Bot. Gdn. Ill. Encyclopedia of Hort. 5 (1981) 1726; The Hoyan 1 #3 (1979) 71. Herbarium Sheets: Whitford #790, Ramos and Edano #35522 1910; Loher #14990 1909 (UC), #14574 1909 (UC), Damo without # 1926 (UC). Peele #815 1926. Meaning: darwinii — Named in honor of Charles Darwin (who was not the collector as the ii ending indicates.

Hoya dasyantha Tsiang In Sunyatsenia 3 (1936) 174-175. Tsiang. Hoya dasyantha Tsiang, sp. nov. Suffrutex epiphyticus scandens praeter corolla intus farinosa glaber, ramulis cinereis 1 mm. diametro, internodiis 6 ad 13 cm. Longis. Folia chartacea late ovata 4 ad 7.5 cm. longa 3 ad 4.8 cm. lata breviter acuminata, basi rotundata, nervis utraque leviter elevatis lateralibus 4 ad 5 jugis, venis transversis conspicuis, supra petiolum 4-ad 6-glandulifera, petiolo 5 mm. longo. Cymae umbelliforme ad 22-florae, cicatricibus pedicellorum paucis, foliis breviorae, pedunculo 2..5 ad. 5 cm. longo 1 mm. diametro, pedicello 1.2 cm. Longo. Flores alba 8 mm. diametro, calice glabro, corolla cinaracteres sectionis Euhoyae, intus farinosa, angulo postero coronae acuto. Gynostegium gynaeciumque characteres generis. Folliculi non visi. Epiphytic climbing subshrub glabrous except the corolla which is mealy inside; branchlets grayish, 1 mm. in diameter; internodes 6-13 cm. long: Leaves chartaceous, broad ovate, 4-7.5 cm. Long, 3-4.8 cm. wide, shortly acuminate, rounded at the base; nerves slightly elevated on both surfaces, lateral nerves 4-5 pairs, veinlets transverse, conspicuous; glands 4 to 6 in a group above the petiole; petiole 5 mm. long. Cymes umbelliform, to 22- flowered, shorter than the leaves, scars of fallen pedicels not many; peduncle 2.5-3 cm. long, 1 mm. in diameter; pedicel 1.2 cm. long. Flowers white, 8 mm. in diameter, calyx glabrous, corolla of the characters of the section Euhoya, mealy inside; posterior angle of coronal-lobes acute. Gynostegium and gynoecium of the generic characters. Hainan: Man-ning, alt. 1100 m., in woods climbing on rock, F. C. How 72103 (Type), April 12, 1935; Boting, F. C. How 72558, May 23, 1935. Allied to H. Fungii Merrill, differing from it in smaller leaves, densely flowered inflorescence, smaller flowers and glabrous calyx. In Flora Republicae Popularis Siamicae, Toms 63 (1977) 480-481. Tsiang Hoya dasyantha Tsiang in Sunyatsenia 3: 174. 1936; followed by script.

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Hoya davidcummingii Kloppenburg In Fraterna 2nd. Quarter (1995) 10-11. R. D. Kloppenburg. Hoya davidcummingii Kloppenburg. Type: #9 (BISH); isotype (UC), discovered by David Cumming 1.5 Km. from Lake Bulusan turn off towards the lake at the forest edge, when coming from Irosin. Elevation ca. 500’. Section Acanthostemma (Blume) Kloppenburg. This specimen he labeled 3232. Epiphytic, gracilis, scandens, ramosa; ramis ramulisque filiformibus, flexuosis, glabris vel sparsis, lax foliatis; radicantibus; foliis petalis breviter petiolatis, lanceolatoellipticus acutis vel obtusiusculis basi cuneatis, glabris, apice recurvis, textura crasse coriaceis ca. 5-7.5 cm. longis 2-2.5 cm. latis; petiolo 0.5-1.2 cm. longo, glabro; cymis umbelliformibus, 10-30 floris; pedunculo glabro 3-6 cm. longo; pedicellus filiformibus ca. 0.85 cm. longo, quam ligulae presentia; corolla subrotata recurvula ca. 1.30 cm. diametiente, extus glabra punctata, intus dense pubescentia; coronae foliola dorsa erecta, subtus in ligulas 2 oblongas foliolum superantes extensa; corona ca. 0.60 cm. diametiente. Stems: twining branched and rebranched, brown becoming woody, flexible and mostly glabrous but with occasional hair cells especially when young, frequently rooting on the basal side of the nodes. Milky white sap present. Internodes: ca. ± 8 cm. long, somewhat enlarged. Petiole: twisted (with slight curve) round, not grooved, with a small gland at upper blade attachment. 1 cm. long by 0.18 cm. in diameter. Leaf Blade: spreading lanceolate-elliptic, upper surface deep green even olivaceous, glabrous with fine granulation’s, edges rounded and slightly thickened, lighter green below: apex mostly acute turning down, base cuneate, overall thickened and coriaceous, veins not visible but pinnate, 5-7.5 cm. long, widest about the middle 2-2.5 cm. Peduncle: straight tapering slightly larger toward both ends 1 cm. long 0.21 cm. in diameter (0.30 at rachis base) Rachis 0.07 cm. long x 0.30 cm. diameter. Pedicel: round with a slight curve, filiform, yellowish, mostly glabrous 0.85 cm. long. Calyx: glabrous shinny inside, ligules present but difficult to see, outside glabrous and granular especially near the rachis. Lobes triangular 0.10 cm. long x 0.09 cm at base with ¼ or less overlap. Ovaries: in pairs, bottle shaped 0.17 cm. tall with the paired base 0.11 cm. wide. Corolla: tends to be patent soon becoming revolute, outside glabrous punctate with scattered yellow spots, base rose; with acute apical lobes, ends revolute and glabrous inner surface with very heavy white pubescent cell structure, otherwise rose to deep rose. Sinus -sinus 0.35 cm.; center - sinus 0.35 cm.; center to apex (flattened) 0.65 cm. making the flower diameter 1.30 cm.; sinus- apex 0.40 cm.; with the widest portion of the corolla lobe 0.35 cm. just outward from the sinus. Corona: fairly upright, inner apex of lobes spatulate and upon drying turn outward at a 90° angle to the central axis, anther apices triangular, membranaceous extending will above the stylar head, prominent and visible, outer apex broadly obtuse; scale saddle shaped in cross section, broad on top with a central ridge, yellow centrally rose; two long broad bilobes from side and under scale incurving at apical ends especially near maturity and on drying. Apes-apex 0.18 cm; apex-end 0.26 cm.; center to apex 0.22 cm.; widest part of scale top 0.12 cm.; anther wing apex - anther wing apex 0.17 cm; retinaculum

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(inner apex) - retinaculum 0.07 cm.. Pollinarium: very distinctive and different. Pollinia broad, short and coarse textured within, 0.035 cm. long, 0.017 cm. wide, with long vacuole adjacent to the sterile edge, this edge extends from ½ of the top down the outer edge to one sixth from the base where it curves under becoming very narrow. Caudicles are bulbous and translucent about 0.01 cm. in diameter at pollinia attachment. Translators are broadly wedge shaped with a wide flat external edge about 0.01 cm wide; narrowing toward the retinaculum which is long and somewhat linear, with no definite head, waist, or hip protrusions. Inner apex narrow and slightly rounded, outer apex more so with slender bifid extensions to the stigma and covered with a clear membrane. Inner apex - outer apex 0.02 cm. long; inner apex - bifid ends about 0.03 cm. long, widest portion 0.009 cm. This new species has been thought to be a variety or subspecies of Hoya gracilis Schlechter. The hoya species in the Section Acanthostemma (Bl.) Kloppenburg can be divided into two broad groups based on the conformation of the inner coronal lobe. One group including this new species have a spatulate inner lobe. The lobe is horizontally much wider than it is thick. Hoya bilobata Schlechter and Hoya loheri Kloppenburg also belong in this division. The other more prevalent form for the inner lobe has been refereed to as “dentate”. Here the inner lobe is laterally compressed, thus it is thicker than broad occupying a vertical position in relation to the scale plane. Hoya gracilis Schlechter along with Hoya wayetii Kloppenburg and Hoya micrantha Hemsley among others exhibit this type of inner coronal lobe. The inner coronal lobes of Hoya gracilis Schlechter well overlaps (cover up) the anther appendages at the flower center whereas with this new species the anther appendages are longer than the inner coronal lobe and are readily visible in the flower center. Another marked difference between the two species is the shape of the stylar head. In Hoya gracilis Schlechter it is conic whereas in Hoya davidcummingii Kloppenburg the apex is expanded into a bulbous head atop the penta columnar base. A major difference exists in the pollinaria of the two species. Hoya gracilis Schlechter has translators “almost lacking” with “very small” retinaculum and relatively narrow finely grained pollinia. The pollinarium of this new species exhibit distinctive very truncated, short, wide pollinia, enlarged prominent translator arms and a relatively long narrow retinaculum. The pollinarium among other differences set this species apart from and other Philippine hoya species and all other Section Acanthostemma species. This species is named in honor of David Cumming who is an avid plantsman. David has collected extensively in the Philippines and elsewhere and contributed many clones to living hoya inventories. He is a conservative collector, taking cuttings only for propogation, leaving the major portion of wild hoya vines to continue their growth in the native habitat. 5 March, 1995 This hoya has twining branched and rebranched stems, becoming brown and woody with age, flexible and mostly glabrous but with occasional hairs especially when young. It frequently roots on the basal side of the stem nodes. A milky sap is present. Internodes are ca. 8 cm long. The petiole has a slight curve is round, not grooved, 1 cm. long by 0.18 cm. in diameter. The leaf blades are spreading lanceolate-elliptic, upper surface deep green, glabrous with fine granulations, edges rounded and slightly thickened, lighter green below; apex mostly acute, turning down, base cuneate, overall thickened and coriaceous, nerves obsolete (but pinnate) 5-7.5 cm. long, widest in the

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middle 2-2.5 cm. Peduncle strict, tapering toward both ends 1 cm. long x 0.21 cm in diameter. Pedicel round with a slight curve, filiform, yellowish, mostly glabrous 0.85 cm. long. Calyx glabrous with ligules present, outside granular, lobes triangular 0.10 cm. long x 0.09 cm. at base with 1/4 or less overlap. Ovaries in pairs, bottle shaped 0.17 cm. tall with paired base 0.11 cm. wide. Corolla revolute, outside glabrous punctate, rose in color with acute apical lobes, ends revolute and glabrous, inner surface with heavy white pubescence, otherwise deep rose. flower diameter flat is 1.30 cm.. Corona fairly upright, inner apex lobes spatulate and on drying turn out at a 90° angle to the central axis; anther apices triangular, membranaceous extending well above the central stylar head. Scale saddle shaped, outer apex obtuse broad on top with a central ridge, yellow centrally rose, two long broad bilobes from the side and under the scale, 0.18 long widest 0.12 cm. Pollinaria very distinctive with broad short pollinia, long wide retinaculum prominent translators and medium caudicle, a well developed structure. References: Philippine Hoya Species 3rd. Ed. 65-66. Translation of Latin above: Epiphytic, slender, with trailing branches , branches rebranched, threadlike flexible, glabrous or sparsely so, loosely leaved, rooting, leaves flat shortly petiolate, texture thick leathery about 5 to 7.5 cm long and 2 to 2.5 cm wide, petiole 0.5 to 1.2 cm long glabrous, cymes umbelate10 to 30 flowers, peduncle glabrous 3 to 6 cm long. Pedicels threadlike about 0.85 cm long with a ligule present; corolla somewhat rotate recurved about 1.3 cm in diameter, outside glabrous punctate, inside densely pubescent, leaflets of the corona back erect, below with 2 ligules oblong extended beyond and rising above the scale; corona about 0.60 cm in diameter. In Philippine Hoya Species 3 rd Ed. (1996) 65-66. R. D. Kloppenburg. Hoya davidcummingii Kloppenburg in Fraterna 2nd. Quarter 1995.Type: #9 (BISH); isotype (UC), discovered by David Cumming 1.5 Km. from Lake Bulusan turn off towards the lake at the forest edge, when coming from Irosin. Elevation ca. 500’. Section Acanthostemma (Blume) Kloppenburg. Key #43 This hoya has twining branched and rebranched stems, becoming brown and woody with age, flexible and mostly glabrous but with occasional hairs especially when young. It frequently roots on the basal side of the stem nodes. A milky sap is present. Internodes are ca. 8 cm long. The petiole has a slight curve is round, not grooved, 1 cm. long by 0.18 cm. in diameter. The leaf blades are spreading lanceolate-elliptic, upper surface deep green, glabrous with fine granulations, edges rounded and slightly thickened, lighter green below; apex mostly acute, turning down, base cuneate, overall thickened and coriaceous, nerves obsolete (but pinnate) 5-7.5 cm. long, widest in the middle 2-2.5 cm. Peduncle strict, tapering toward both ends 1 cm. long x 0.21 cm in diameter. Pedicel round with a slight curve, filiform, yellowish, mostly glabrous 0.85 cm. long. Calyx glabrous with ligules present, outside granular, lobes triangular 0.10 cm. long x 0.09 cm. at base with 1/4 or less overlap. Ovaries in pairs, bottle shaped 0.17 cm. tall with paired base 0.11 cm. wide. Corolla revolute, outside glabrous punctate, rose in color with acute apical lobes, ends revolute and glabrous, inner surface with heavy white pubescence, otherwise deep rose. flower diameter flat is 1.30 cm.. Corona fairly upright, inner apex lobes spatulate and on drying turn out at a 90° angle to the central axis; anther apices triangular, membranaceous extending well above the central stylar head. Scale

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saddle shaped, outer apex obtuse broad on top with a central ridge, yellow centrally rose, two long broad bilobes from the side and under the scale, 0.18 long widest 0.12 cm. Pollinaria very distinctive with broad short pollinia, long wide retinaculum prominent translators and medium caudicle, a well developed structure. Meaning: named for the collector David Cumming, from Australia and now S. Africa.

Hoya dennisii P. I. Forster et D. J. Liddle In Austrobalia 4(1) (1993) 51-52. P. I. Forster & D. Liddle. Hoya dennisii P. I. Forster et D. J. Liddle, sp. nov. affinis H. chloranthae Rech. a qua venis secondariis laminae folia 3 vel 4 vix manifestis, floribus minoribus (16-17 mm in diametro), lobis coronae longioribus (circa 7 mm longis), et lobis coronae staminalis majoribus margins exteriore corollam excedentibus et apice lobis obtuso producto differt. Typus: Solomon Islands, Guadalcanal Province: Kwalo Range, Mt. Gallego, eastern ridge, 7 July 1965, T.C. Whitmore R.S.S. 6073 (halo: L!; iso: A!, BSIP!, K!, LAE!). Epiphytic coriaceous Liane to several meters; latex white. Stems cylindrical, wiry, with sparse trichomes, becoming glabrous with age, internodes up to 100 mm long and 1 mm diameter. Leaves petiolate; lamina lanceolate-ovate, coriaceous, up to 72 mm long and 30 mm wide, discolorous, glabrous, venation obscure; upper surface dark green lower surface pale green; tip acuminate; base rounded to cordate petiole 5-7 mm long and c. 0.8 mm diameter, with scattered trichomes colleters 2, at lamina base. Cyme racemiform, up to 100 mm long peduncle up to 85 mm long and 0.5 mm diameter, glabrous or with scattered trichomes, bracts ovate 0.5-0.6 mm long, 0.4-0.5 mm wide glabrous. Flowers 2-3 mm long, 16-17 mm diameter; pedicels 30-35 mm long, c. 0.6 mm diameter, glabrous Sepals lanceolate, c. 1.2 mm long and 0.6 mm wide, glabrous. Corolla flattened campanulate, crimson-pink externally glabrous, internally with short dense white trichomes, tube c. 5 mm long and 10 mm diameter, lobes triangular, c. 7 mm long and 7 mm wide. Staminal corona purple, c. 1.8 m long and 4.5 mm diameter inserted on column ± flush with corolla; lobes c. 3.2 mm long, 1.8 mm wide and 1.8 mm high, outer edge rounded and somewhat drawn out into lip not higher than the style-head inner edge acute. Staminal column c. mm long and 2.5 mm diameter; anther appendages lanceolate, c. 0.8 n long and 0.5 mm wide; alar fissure 3-1.1 mm long. Style-head pressed-globose, c. 0.8 mm meter. Pollinaria c. 0.55 mm long and 0.46 mm wide; pollinia oblong, c. 6 mm long and 0.18 mm wide, with pellucid germination mouth on outer edge; corpusculum oblong-ovate, c. 14 mm long and 0.09 mm wide. Fruit and seed not seen. Specimens examined: Solomon Islands. Guadalcanal Province: near Gold Ridge Village, 9°35'S, 160°07'E, Jun 1991, Forster 8714 & Liddle (BRI); Summit Mt. Gallego, NW Guadalcanal Rep 1966, Dennis BSIP4630 (BSIP), Popomanasia, Oct 1965, Corner 156 (K). Distribution and habitat: H. dennisii has been collected only on Guadalcanal where it grows in montane and submontane situations from 650 to 1200 m altitude, usually as an epiphyte in bryophyte mats in rainforest. Notes: H. dennisii belongs to a group of small, wiry stemmed, coriaceous to herbaceous leaved hoyas that occur in upper montane areas in Papuasia and Melanesia. The group includes H. venusta Schltr. in New Guinea and H. chlorantha Rech. in Samoa. H. dennisii is probably most closely related to H. chlorantha but differs from that species in

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the just visible 3 or 4 secondary veins in the leaf lamina, the smaller flowers 16-17 mm in diameter with longer corolla lobes c. 7 mm long and the larger staminal corona lobes in which the outer edge; raised above the corolla and drawn ut into a blunt lip. Entomology: It gives us much pleasure to name this new species for Geoff Dennis of Honiara, Guadalcanal former Government Botanist and long-time resident in the Solomon Lands, a plantsman of high repute with a garden of tropical delights. Geoff has been responsible for introducing many of the Hoyas from the Solomon Islands into general cultivation. Translation: with an affinity to Hoya chlorantha Rechinger, 3 or 4 veins are barely visible from the secondary veins of the leaf blade, flowers small (16 to 17 mm in diameter) and the lobes of the staminal corona are large with the apex of the lobes exceeding the outside margines of the corolla and the apex of the lobes obtuse and different.

Hoya densifolia Turczaninow In Bulletin de la Societe Imperiale des Naturalistes de Muscou 21, pt.2 (1848) 261. N. Turczaninow. Hoya densifolia Turczaninow. Erectiuscula: foliis subsessilibus membranaceis dense imbricatis ellipticis, sinu brevi cordatis, acutiusculis penninerviis reticulatis glabris; umbellis lateralibus multifloris; pedicellisque cano-pubescentibus; corollae laciniis triangularibus glabris; coronae stamineae gynostegio adnatae parse infera foliolorum horizontal) compressa, subtus profunde sulcata. - Foliorum dispositiore nec forma et consistentia ad H. imbricatam approximatur, in ordine a Decne, institute prope H. Iaurifoliam ponenda, foliorum tamen consistentia ad Watahakas vergit. Habitat in Insula Java. Translation: Somewhat erect, leaves somewhat sessile membranaceous densely imbricate elliptic, with short cordate sinus, slightly acute penninerved reticulate glabrous; umbels lateral many-flowered; peduncles and pedicels grayish-white pubescent; with the lobes of the corolla glabrous triangular; with the corona stamina horizontally compressed, part beneath the leaflet adnate to the gynostegium, below definitely sulcate. Leaflets approaching H. imbricata not only in arrangement but also in consistency based in the order of Decaisne arrangement near H. laurifolia, leaf blades tending toward or near to Watahaka. - Habitat in the Island of Java. In Bulletin de la Societe Imperiale des Natualistes de Moscou 21 (1848) 261. N. Turczaninow. Hoya densifolia Turcz. Turcz. Bull. Mosc. XXII, 2, 261. — Miquel, Flora Ind. Bat, II, 525 — Z. M. 2620! In Annales Botanices Systematicae 3 (1852) 66. W. G. Walpers. 5. H. densifolia Turcz. 1. cit. pag. 261, — Erectiuscula: foliis subsessilibus membranaceis dense imbricatis ellipticis, sinu brevi cordatis, acutisculis penninerviis reticulatis glabris; umbellis lateralibus multifloris; pedunculis pedicellique cano-pubescentibus; corollae laciniis triangularibus glabris; coronae stamineae gynostegio adnatae parte infera foliolorum horizontali compressa, subtus profunde sulcata. — Foliorum dispositione nec forma et consistentia ad H. imbricatam approximatur, in ordine a Decn. instituto prope H.

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laurifoliam ponenda, foliorum tamen consistentia ad Watahakas vergit, — Habitat in Insula Java. Translation: Very erect: leaves somewhat sessile membranaceous densely imbricate elliptic, at the recess briefly cordate; slightly acute reticulately penninerved glabrous; umbels lateral many flowered; both peduncles and pedicles grayish white pubescent, with the leaflets of the corolla triangular glabrous; with the staminal corona adnate to the Gynostegium interior leaflet compressed horizontally, below definitely grooved. — Foliage disposition and form approximately consistent with that of H. imbricata, after the arrangement order of Decaisne. near Hoya laurifolium, nevertheless the foliage tending towards and consistent with Watahakas. Habitat: in the Island of Java. In Flora van Nederlandsch Indie Bataviae 2 (1856) 525-526. F. A. W. Miquel & Zollinger. 33. Hoya densifolia Turcz. Ramuli juniores puberi, folia brevissime petiolata dense saepe imbricata, e basi rotundata cordato-emarginata elliptico-ovata acutiuscula vel brevissime acuta, carnosulo-subcoriacea nitida, adulta glabra, nascentia praesentim secus margines parce pilifera, obsolete (in sicco) venulosa, 1 ¼ — 1 ¾ poll. longa, pedunculi vulgo pollice breviores, receptaculo brevi ellipsoideo, bracteis hirtellis, pedicelli hirtelli pollicares, sepala anguste elliptica dorso patula subhirtella, coronae staminae phylla dorso conico-obtuso-producta, basi denticulis 2 obtusiusculis. Hoya densifolia Turcz. in Bull Mosc. XXII. 2. 0. 261. Java (Horsf. Zoll., n. 2620 ). Translation: Juvenile stems puberulus, leaves with short petioles, dense often imbricate, with the base rounded cordate-emarginate, elliptic-ovate very acute or shortly acute, fleshy somewhat leathery shiny, adults glabrous, to begin with chiefly with sparse hairs along the margins (when dry) no veins, 1 ¼ to 1 ¾ inches long, peduncles generally less than 1 inch long, receptacle shortly ellipsoidal, bracts hairy, pedicels hairy 1 inch long, sepals narrowly elliptic back spread out somewhat hairy, leaflets of the staminal corona back conic-obtuse, base 2 tooth-like ends, somewhat obtuse. In Icones Bogorensis (1909) 157-158. C. A. Backer. Hoya densifolia Turcz. Turcz. Bull. Mosc. XXII, 2, 261. — Miquel. Flora Ind. Bat. II, 525. — Z. M. 2620 !. 1. Ramulus florens 1/1 — Fig. 2. Flos a latere visus 2/1. — Fig. 3. Flos ab apice visus 7/2 — Fig. 4. Flos verticaliter dissectus 2/1. — Fig. 5. Pollinia 10/1. Hoya caule subscandenti, foliis densis., internodiis plus quam duplo longioribus, breviter petiolatis, pro genere tenuibus, subcarnosis, e basi cordata subobovatis acutiusculis, floribus in apice pedunculi brevis subumbellatis, corollae mediocris cerinae intus papilloso-hirtellae segmentis reflexis, coronae phyllis margine recurvatis, calcar breve bicostulatum extrorsum spectans efformantibus, stigmata obtusiuscule rostrato. Flores umbellato-racemosi, racemis in axillis superioribus pedunculatis, plurifloris, sub-nutantibus. Pedunculus communis teres vel subcompressus, viridis, glaber, 10—20 mm. longus, ± 1¼ mm. crassus, apice incrassato. Receptaculum breve. Bracteae ovato-ellipticae, obtusae, in dorso hirtellae, vix ¾ mm. longae. Pedicelli teretes virides glabri vel subglabri 17—25 mm. longi, apicem versus sensim incrassati. Calycis fere ad basin 5-partiti diametrus ± 6 mm., segmenta corollae appressa, anguste ovato-elliptica, obtusiuscula, hyaline marginata, ± 2 ½ mm. longa, in margine dorsoque subhirtella. Corolla subcarnosa cerina nitidula extus glabra intus tenuiter papilloso-hirtellae, in fauce pilis longioribus retrorsis instructa, a basi tubi usque ad

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apicem segmentorum 8 mm. longa, segmenta reflexa late triangulari-ovata acuta, marginibus recurvatis, hinc specie anguste ovato-elliptica, ± 5 mm. longa. Phylla coronae stamineae stellatae ± 7 mm. diametrum metientis, pallide flava marginibus recurvatis sibimet arcte appressis hinc calcar breve obtusum concavum extrorsum spectans juxta marginem inferiorem utrinque costa longitudinali horizontali prominula instructum, efformantibus, in centro depressa angulo interiore in dentum oblique erectum membranum antherarum subaequantem protracto. Gynostegium breviter stipitatum, stipes ± 1 mm., antherarum membranum latiusculum, acutiusculum stigma breviter superans. Corpuscula minuta brunnea, pollinia sessilia deltoidea, apice oblique truncata cerina margine postico pellucida ± ¾ mm. longa, apice ± 1/5 mm. lata. Ovaria 2, sibimet appressa, ovoidea viridia glabra, ± 1 1/3 mm. longa styli liberi, ± 2/3 mm. longi cylindrici virides glabri, stigmate communi obtusiuscule rostrato coronati. Gynoecium totum 3 ½ mm. longum. Folliculi adhuc ignoti. Frutex subscandens, succo 1acteo albo glutinoso repletus, rami teretes glabri, cortice griseo. Folia opposita densa internodiis plus quam duplo longiora, hinc in sicco imbricata. Stipulae minutae, triangulares, acutae, caducae. Petioli breves suberecti, 2—3 mm. longi, supra planiusculi, subtus convexi. Folia pro genere tenuia, subcarnosa e basi cordata elliptico-subobovata acutiuscula integerrima lucidula 40—60 mm. longa, 20—30 mm. lata, in medio ± ¾ mm. crassa nascent in margine costaque parce pilifera, mox glabra laete viridia utrinque minutissime albido-punctata, distincte penninervia, costa media basi prominula, nervis lateralibus utrinque 7-10, patulis, pellucidis, haud longe a margine ramos arcuatum confluentes emittentibus, in vivo immersis in sicco vix prominulis. Duae formae exstant nonnisi in anthesi distinguendae: 1. concolor. Phyla coronae stamineae ommino flavae. 2. bicolor. Phylla coronae stamineae in medio purpureo-maculata. Legerunt: 1. Zollinger, ad pedem montis Lamongan. 16, 1, 1845 [N°. 2620!] 2. Collector indigenus prope Tamdjong Priok [forma bicolor, specimen quod nunc in Horto Bogoriensi colitur’]. 3. Collector indigenus in moute Goentoer [forma concolor, specimen nunc in Horto Bogoriensi' cultum ']. This species which seems to be rare in Java has been wrongly placed by Miquel in the section Cathetostemma, it is a true Plocostemma. The Index Kewensis unites it, perhaps rightly, with H. Cumingiana Decaisne, a native of the Philippine Islands. From the brief diagnosis of this species in the Prodromus of De Candolle it does not appear that the two plants are quite identical. I have not been able to procure the original description of Turcz. It is not impossible that this specimen is wrongly labeled and does not come from there. Translation: Hoya stems somewhat creeping, leaves dense, internodes more than twice longer, petioles short, narrow, for the genus, somewhat fleshy, cordate at the base, somewhat umbellate, segments of the medium sized corolla waxy inside papilose-hairy reflexed, leaflets of the corona margins recurved spur short and additionally 2 ribbed outside laying toward the stigma somewhat obtusely beaked. Flower in umbellate racemes, racemes in the axils above the peduncles, many flowered somewhat nodding. Peduncles commonly 3 or somewhat compressed. Green many flowered 10 to 20 mm long, plus or minus 1 ¼ mm thick, apex thickened.

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Receptacle short. Bracts ovate-elliptic obtuse, on the back hairy barely ¾ mm long. Pedicles round green glabrous or somewhat glabrous 17 to 25 mm long, with the apex gradually thickened. Calyx 5 parted nearly to the base, diameter plus or minus 6 mm, segments of the corolla appressed, narrowly ovate-elliptic, obtuse, with a hyaline margin plus or minus 2 ½ mm long, on the back margins somewhat hairy. Corolla somewhat fleshy waxy shiny, outside glabrous, inside slightly papilose-hairy, in the throat are long pointed hairs, at the base of the tube all the way to the apex segments 8 mm long, segments reflexed broadly triangular-ovate acute with recurved margins from this place narrowly-ovate elliptic plus or minus 7 mm in diameter, pale yellow, margins recurved or closely appressed from a short obtuse spur, outside concave laying towards or near the lower margins, both sides of the midrib longitudinally horizontal prominently formed, depressed in the center, interior angle tooth-like, oblique erect somewhat equal to the anther membrane. Gynostegium shortly stalked, stalk plus or minus 1 mm, anther membrane a little broad a little acute, stigma shortly upward. Corpuscle minute brown, pollinia sessile deltoid, apex oblique truncate waxy margins outside pellucid plus or minus ¾ mm long, apex plus or minus 1/5 mm wide, 2 ovaries, or appressed, ovoid green glabrous plus or minus 1 ½ mm long, free from the style plus or minus 2/3 mm long cylindrical green glabrous. Stigma very common with a beaked obtuse corona. Gynoecium all of 3 ½ mm long. Follicles not seen. Twining somewhat climbing, full of milky-white gelatinous sap, stems round glabrous bark gray. Leaves opposite dense, internodes at least 2 times longer, at least when dry imbricate. Stipules minute triangular, acute falling. Petioles stout somewhat erect 2 to 3 mm long, above a little flat below convex. Leaves narrow for the genus, somewhat fleshy cordate at the base elliptic ovate a little acute absolutely entire 40 to 60 mm long 20 to 30 mm wide, in the middle plus or minus ¾ mm thick, arising from the margins and the midrib a few fine hairs, soon glabrous bright green on both sides whitedots, distinctly penninerved, midrib prominent in the center base, lateral nerves on both sides 7 to 10, spreading, pellucid, not at all long, branched near the margins meeting in an arch, when alive imbedded, when dry scarcely prominent. When flowering two distinct forms can be distinguished: 1. one colored. Leaflets of the staminal corona yellow all over 2. bicolored. Leaflets of the staminal corona in the center with purple spots. Legerunt: 1. Zollinger, from the foot of Mt. Lamongan 16, 1, 1845 (#2620!) 2. collected indigenous near Tamdjong Priok (forma bicolor, specimen each at present in culture at the Bogor Hortium. 3. Collector indigenous on Mt. Goentoer (forma concolor, specimen not at all in culture at Bogor Hortium. In Flora of Java 5 (1965) 266. Backer. (Key). 2 b. Corolla-segments glabrous inside or covered with very short patent hairs; top stigma shortly conical; corona inserted distinctly above the base of the corolla. Leaves without trichomes ............................................................................................................................ 3 3a. Leaf-pairs 1.5 — 4 cm distant; petiole 2 — 4 mm long; leaves oblong or ellipticobovate, from a cordate, subamplexicaul base, obtuse or very shortly acuminate, thinly fleshy, thinly coriaceous when dry, on either side of midrib with several erecto-patent thin nerves, glabrous or, when young, sparingly hairy on midrib and along margins, 4 —

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6.5 cm by 2 — 3.5 cm. Peduncle 1 — 2.5 cm, 10 — 15 flowered; pedicels thinly patently pubescent, 1¾ — 2 ½ cm; calyx on the inside with 5 short, linear subulate glands; segments narrowly ovate-elliptic, rather obtuse, patently pubescent outside, 2 — 3 mm long; corolla shiningly waxy-yellow, glabrous outside, densely and very shortly patently pubescent inside, in the throat with a whorl of rather long hairs; from the base of the tube to the top of the segments measuring c. 8 mm; segments ovate-triangular with recurved margins; corona inserted 1½ mm above the base of the staminal tube, c. 7 mm diam., entirely yellow or violet-blotched in the centre, its scales horizontal, at the inner angle ending in an erecto-patent mucro which is appressed against the stigma; pollinia narrowly deltoid, narrowly pellucid-margined, c. ¾ mm long. 0.40 — 0.60; 1 — XI; W. C. E.; 1 — 100; sunny or slightly shaded localities, open forest boarders, on tree-trunks, often also on rocks ............................................................................................... H. densifolia Turcz. In Systematisches Verzeichnis Family 1 (1911) 6. S. H. Koorders & Ann Schumacher. Hoya densifolia Turcz. — Windender Strauch, bis 3 m. hoch. Mittel-Java: Banjumas: Nusakambangan: Region I: Kds. 30588  (5 * 24. IV. 1902). Translation: Twining shrub, up to 3 meters high. From Central-Java. etc. In Exkersionsflora von Java 3 (1912) 97. “Flora of Java” S. H. Koorders & Ann Schumacher. (Key) 1b. Coronazipfel sternförmig spreizend. Gynostegium ganz oder fast ganz sitzend. Korolle glocken-odor radförmig, mit brieten, bisweilen zurkück-gebogenen Zipfeln. (4. Sektion. Euhoya Miq.) ..................................................................................................... 2 2a. Blätter nicht dachziegelig deckend ............................................................... 3 2b. Blätter sehr gedrängt stehend, stets ± dachziegelig deckend, sehr kurtz-gestielt, elliptisch-eiförmig, am Grunde herzförmig ausgerandet, kahl, in der Jugend unterseits spärlich behaart, fleschig-lederig, bis 5 cm lang. Pedunculi ± bis 2 ½ cm. Pedicelli 2 ½ cm, kurz steifhaarig. Narbenkopf in der Mitte gespitzt. Windenden Strauch, bis 3 m hoch. Miq l. c. c. 526; Backer in Icon, Bogor. tab. 262. Ganz Java: Im unteren Gebirge und in der Ebene auf nackten Felsen und im lichten Regenwald sehr zerstreut. (Kds. n. 39688 , Zollinger n. 2620.) ..... H. densifolia Turcz.* Translation: 1b. Corona scales star-shaped spreading. Gynostegium quite or almost completely sedentary. Corolla bell-shaped otherwise wheel-shaped, with breadth, from time to time recurved margin corners. (4.section. Euhoya Miq.) .....……............ 2 2a. Leaf not imbricate covered .....................................…….................. 3 2b. Leaves very much crowded upright, constant ± imbricate covered, very much shortpetioled, elliptic-oval, at the base heart-shaped upright, glabrous, in the youth on the undersides sparsely hairy, fleshy-leathery, up to 5 cm long. Peduncle ± until 2 ½ cm. Pedicel 2 ½ cm, shortly stiff-hairy. rachis-head sharpened in the middle. Winding shrub, up to 3 m high. Miq L. C. C. 526; Backer in Icon, Bogor. tab. 262. Completely Java: In the low mountain and in the level on bare rocks and in the light rainforest, very much scattered.

Hoya dickasoniana P. T. Li

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In Journal of the South China Agricultural University 15/2 (1994) 74. “Three New Species of Hoya (Asclepiadaceae) Form Myanmar Li Bingtao. Hoya dickasoniana P. T. Li, sp. nov. TYPE: Myanmar, Mogok, 1800 m, April 1934, F. G. Dickason 3 032 (holotype, A). Frutex epiphyticus; ramulis pubescentibus; folia opposite etiam 3 - vet 4— verticillata, elliptica vet ovate, 9-19mm x 5—10mm, basi late cuneata vet rotundata, apice obtuse, glabra; petiolo 1mm longo; umbellae terminates, 3 - 6—florae; pedicelli 1 cm long), pubescent); segmentis calycinis oblongis, 2 — 2.5 mm x 1 mm, pubescentibus; corolla alba, cat 1.7cm diam., tuba 4mm longo, extus glabro, intus basi pubescent), lobis ovato-deltatis, cat 5mm x 5mm; corona 5—lobata, carnosa, ovato-oblonga, cat 2.5 mm longa, marginibus revolutis; polliniis oblongis, cat 1 mm longis, erectis, margine pellucido. Epiphytic shrubs. Branches pubescent, internodes shorter than leaves. Leaves approximate, overlapping, opposite and in whorls of 3 or 4, elliptic or ovate, 9—19 mm x 5 — 10 mm, fleshy, glabrous, base broadly cuneate to rounded, apex obtuse; mid and lateral veins obscure; petiole 1 mm long. Umbels terminal, 3-6—flowered; peduncles 5 mm long, pubescent; pedicels 1 cm long, pubescent. Calyx with 5 basal glands inside; lobes oblong, 2—2.5mm x 1mm, pubescent outside, glabrous inside. Corolla white, 1.7cm in diem., fleshy; tube 4 mm long, glabrous outside, pubescent basally inside; lobes ovate-deltate, ca. 5mm x 5mm, glabrous. Corona scales 5, fleshy, ovate-oblong, cat 2.5mm long, margin revolute, outer angle obtuse, inner angle acuminate and as high as the apical membrane of anthers. Pollinia oblong, cat 1 mm long, erect, outer margin pellucid. Ovaries glabrous; stigma apex restate. Follicles not seen. Hoya dickasoniana, which is so far known only from the type collection, is most closely related to H. imbricata Callery ex A. DC. It is readily distinguished by its opposite and whorled, elliptic or ovate leaves and pubescent branches, peduncles, and pedicels.

Hoya dictyoneura K. Schumann In Nachtrage zur Flora Deutschen Schutzgebiete in der Südsee (1905) 362-363. K. Schumann & K. Lauterbach. Hoya dictyoneura K. Sch. n. sp. Frutex scandens ramis florentibus gracilibus novellis ipsis complanatis glabris; foliis breviter petiolatis oblongis vel ellipticis vel suborbicularibus acutis basi rotundatis quintuplinervis utrinque glabris; racemis breviter pedunculatis congestis primum subumbellatis demum rhachide crassa elongata; floribus pedicellatis pluribus; sepalis ovatis glabris parvis; corolla alto in lobos ovatos acutos divisa intus prope basin papillosa; coronae lobis radicantibus acutis subtus subtilissime oblique striatis. Der etwa meterlange blühende Zweig ist am Grunde 3 mm. dick, holzig und mit dünner zimmtfabiger Rinde bekleidet, Der sehr kräftige, oben abgeflachte Blattstiel ist höchstens 1 cm lang; die Spreite hat eine Lange von 5 - 8.5 cm und eine Breite von 3.5 - 6 cm das oberste Paar der Grundnerven verläust bis zur Spitze, stärkere Seitennerven sind nicht entwickelt, dafür springt aber das Nervennetz an beiden Seiten Kräftig, wenigtens an der getrockenten Pflanze vor. Der Blütenstand wird von einem 1 cm langen, sehr kräftigen Steil getragen; die Rhachis miszt 5 mm; die Blütenstielchen sind 1.5 cm. lang. Der Kelch miszt reichlich 1 mm; die weisz Blumenkrone ist 6 mm long. Die Staminodien sind 4 mm lang. Kaiser-Wilhemsland:

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Sattelberg, 850 m u. M. (Nyman n. 720, bl. Juli 1899). die Form die Blätter und die Ziemlich kleinen weiszen Blüten kennzeichnen die Art. Translation: Climbing shrub branches flowering slender young stems themselves flattened and glabrous; leaves briefly petiolate oblong or elliptic or somewhat orbicular acute with the base rounded five nerved both sides glabrous; racemes shortly peduncled crowded together somewhat umbellate at first later the rachis thick elongate; flowers from multiple peduncles; sepals ovate glabris small; corolla high with lobes ovate acute divided inside near the base papillose; lobes of the corona spreading acute below somewhat very little obliquely striate. The nearly meter long blooming branch is near the base 3 mm. thick, woody and clothed with cinnamon-colored bark. The very strong petiole is somewhat flat and measures 1 cm. long. The leaves are 5 - 8.5 cm. long and 3.5 - 6 cm. wide. The principle pair of thick basal nerves reach almost to the leaf tips; side nerves are not well developed, instead there is more a network of strong veins on both sides, at least on the dried plants. The inflorescence is held on a very strong 1 cm. long peduncle carrying a 5 mm. long rachis. The peduncles are 1.5 cm. long. The calyx measures as much as 1 mm; the white corolla is 6 mm. long. The corona scales are 4 mm. long. New Guinea: Sattelberg, 850 meters above sea level (Nyman #720, blooming in July 1899). The form of the leaves and the handsome small white flowers characterize this species. In Botanische Jahrbücher 50 (1913) 117. “Die Asclepiadaceen von Deutsch Neu Guinea” R. Schlechter. 19 ? H. Dictyoneura K. Schum. in K. Schum. u, Lauterb., Nachtr. (1905) p. 362. Nordöstl. Neu-Guinea: auf dem Sattelberg, bei 850 m u. M. (E. O. Nyman n. 720. — Blühend in Juli 1899). Das Original dieser Art im Berliner Herbar besteht aus einen sehr dürstigen Exemplare mit einer recht schlechten Blüte. Soweit sich danach überhaupt etwas erkennen läszt. möchte ich glauben. dasz die Art in diese Verwandtschaft gehört, denn sie hat die dicken Zweige und bereiten Blätter der beiden oben behandelten Arten. An eine identität mit einer der breits beschriebenen möchte ich nicht glauben, da die Standortsverhältnisse zu verschieden sind. Auszerdem hat H. dictyoneura K. Schum. auf der Unterseite fünf stark hervortretende Nerven, welche bei den übrigen fehlen. Beeseres Material der Art ware jedenfalls sehr erwünscht. Nach Schumann sind die Blüten weisz. Translation: Northeastern New Guinea on Saddle Mountain, at 850 meters altitude (E. O. Nyman #720 — blooming in July 1899). The original of this species is in the Berlin Herbarium, but is a very poor example with rather a bad bloom. As far as can be determined, I would like to believe that the species belongs in this group, for it has the thick branches and wide leaves of both of the above discussed species. I would not like to think it is identical with one of those already described, since the local conditions are different. Moreover, H. dictyoneura K. Schum. has five very prominent veins on the underside that are lacking in the others. In any case better samples of the species are very much desired. According to Schumann, the blooms are white. In Hoyas of Northeastern New Guinea (1992) 42. R. D. Kloppenburg (Translation). Hoya dictyoneura K. Schumann in K. Schumann and Lauterbach. Nactr. (1905) p. 362.

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Northeast New guinea: in the saddle mountains, at 850 m. altitude (E. O. Nyman #720 Blooming in July 1899). The original of this species in the Berlin Herbarium is a very poor example, with a rather worthless bloom. As far as generally determined, I would like to believe basically that the species belongs in this group, for it has the thick branches and wide leaves of both above discussed species. I would not like to think that it is identical with one of those already described, since the conditions are so different,. Moreover, H. dictyoneura K. Schumann has five prominent veins (nerves) on the underside, which are lacking in the others. In any case, however, better material of the species is very desirable. According to Schumann the blooms are white. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. dictoneura Schumann – 33.

Hoya dimorpha F. M. Bailey In Queensland Agriculture Journal 3 (1898) 156. F. M. Bailey. H. dimorpha (n. sp.) A stout twiner, the extremities often slender with much smaller leaves. Leaves thickcoriaceous, oblong-acuminate, slightly cordate at the base; those on the thick portion of the stem exceeding 5 in. in length and 2 in. in breadth; those on the slender ends often cordate and under 2 in. long, glabrous, undersides pale, the gland on the face indicating the attachment of the petiole, beneath often a broad ciliate disk at times mote or less umbonate. Petiole stout, usually less then ½ in. long. Peduncles slender, 2 to 3 in. long, bearing an umbel of many — 27 or more — small flowers. Pedicels about 1 in. long. purplish; flower-buds 5 angled, about 3 lines in diameter. Calyx-segments triangular, ½ line long. Corolla nearly white. 5-lobed, expanding to about 5 lines diameter, silky-hairy on the back, more prominently so on the margins, and also forming a ring around the orifice of the corolla tube, nearly glabrous on the face. Corona white glossy, segments about 1 line long, concave above, the sides produced into thick wing-like ribs beneath, Hab. Twining over shrubs and trees growing on the margin of the small bays at the foot of Mount Trafalgar, New Guinea.

Hoya diptera Seemann In Flora Vitiensis (1866) 163. 2. H. diptera, (sp. nov.) Seem. in Bonplandia vol. ix. p.257; scandens, glabra; ramulis 4-angulatis; foliis ovato-ellipticus acuminatis basi ovatis carnosis penninerviis; pedunculis compressis subulatis, petiolo longioribus; floribus flavis. —Viti Levu and Taviuni, on trees (Seemann ! n. 320) also collected by U.S. Expl. Exped. Perhaps a specimen, without flower, collected in the Isle of Pines (M'Gillivray), which has leaves twice as large as those of my specimens, may belong to this species, of which there are also imperfect specimens only. Translation: climbing, glabrous stems 4 angled; leaves ovate-elliptic, acuminate with the base ovate, fleshy, penninerved; pedicels flattened laterally somewhat winged, the petioles longer; flowers yellow.

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In Journal of the Linnean Society of London 43 (1915) 33. “To the Flora of Fiji” L. S. Gibbs & W. B. Turrill. Hoya diptera, Seemann in Bonplandia, vol. ix (1861) p. 257, et Fl. Vit. p. 163. Suva, on Ivy trees in Government House paddock, in flower April 7th, 1905, im Thurn, F 7. Distr. Fiji (Viti Levu). Flowers yellow, i. e. “naiskun” coloured. In Sargentia 2 (1942) 113. "Fijian Pant Studies" A. C. Smith. 5. Hoya diptera Seem. Ex A. Gray in Proc. Am. Acad. 5: 336, nomen. 1862; A. Gray in Bonplandia 10” 37, nomen. 1862; Seem. Fl. Vit. 163. 1866; Turill in Jour. Linn. Soc. Bot. 43: 33. 1915. Vine with slender glabrous or distally puberulent branchlets, sometimes the branchlets short, lateral, densely foliaceous; petioles rugulose, 5-15 mm. long, pale puberulent or glabrous; leaf-blades subcarnose, elliptic or ovate-elliptic or narrowly oblong. 3.5-8 cm. long, (1-) 2-3.2 cm. broad, obtuse base cuspidate or obtusely short acuminate at apex, pinnately-nerved. the costa slightly impressed or plane above, subprominent beneath, the secondary nerves 3 or 4 per side, ascending, oriented from costa toward base, prominent on both surfaces or submersed, the veinlet-reticulation immersed; inflorescences axillary, umbellate, the peduncle 10-45 mm. long, glabrous or obscurely puberulent, ellipsoid-capitulate at the apex, sometimes verrucose for the distal 5 mm., the bracts inconspicuous; flowers 5-10 per umbel at anthesis, the pedicels slender, 8-20 mm. long, glabrous or sparsely pale puberulent; calyx-lobes membranaceous, inconspicuous, deltoid, 0.7-1.1 mm. long and broad, glabrous except for the ciliate margin; corolla submembranaceous, 11-16 mm. in diameter, yellow, copiously and conspicuously puberulent within, the lobes deltoid or ovate-deltoid, 4-6 mm. long and broad, often recurved at margin, the sinuses acute; lobes of the corona thick, oblong, 34.2 mm. long, 1.6-1.8 mm. broad, obtuse at apex, acuminate at base, flattened above, rounded beneath; pollinia 0.5-0.6 mm. long; carpels glabrous. Herbarium sheets: Viti Levu: Tholo North; Nauwanga, near Nandarivatu, alt. about 750 m., Degener 14333 (A) (vine, in open forest; corolla yellow, reddish toward center), Degener 14755 (A) (liana, in forest; corolla yellow); Naitasiri: Vicinity of Nasinu, alt. 150 m., Gillespie 3556 (BISH, UC) (liana with pendent inflorescences, in woods. Vanua Levu: Thakaundrove: Savuthuru Mt. near Velethi, alt. 90 m., Degener & Ordonez 13832 (A) (Vine in open forest; corolla yellow); Vatunivuamonde Mt. alt. 240 m., Degener & Ordonez 14014 (A) (vine in forest: corolla yellow). Without definite locality: Seemann 320 (Type coll., GH) (Viti Levu and Taveuni); U. S. Expl. Exped. (US). As represented by the cited specimens, this species is characterized by inconspicuous calyx-lobes, a comparatively thin corolla which is yellow and perhaps reddish tinged at base within and by the ascending secondary nerves of its leaf-blades. The cited type duplicate does not show the flattened sbualate pedicels mentioned by Seemann, and I am inclined to believe that this character, apparently the source of the specific name, was due merely to the degree of pressing of the actual type. In foliage the type duplicate is an excellent match for the other cited specimens. Corolla submembranaceous, yellow, the sinuses acute; secondary nerves 3 to 4 per side, ascending, oriented from costa toward base ………………………5. Hoya diptera. Editors editions: Additional herbarium sheets: Seemann Type #320 (A,K); Gillespie #4009, 3556; Degener #14333, 14755; Degener & Ordonez #13832, 14014; Horne #66; Pahan #497;

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DA #11212, 15301, 16141, A. C. Smith #9680 (UC), #7202, 6171, 6775, 6586; 62110, 6775 (US); Tothill #608; Thurn F-7. Additional references: Gibbs & Turrill in Journal of the Linnean Soc. of London 43 #288 (1915-1917) 33. Parham in Flora of the Fiji Is. (1972). Symonds in Hoyan 2 (1980) #2. A. C. Smith 4 (1988) 123 in Flora Vitiensis Nova. Comments: This species along with Hoya eitapensis Schlechter, Hoya nummularioides Constantine, and Hoya flavida Forster et Liddle belongs to a Subsection of Section Acanthostemma (Bl.) Kloppenburg; (Subsection Angusticarinatae Kloppenburg). These species have very narrow shelves on the sides of the coronal scales which protrude only slightly beyond or are equal in length to the outer apex of the scale. This represents a transitional stage between species without side modifications of the scale and those species with well developed side extensions termed bilobed since they extend beyond the outer coronal apex to varying degrees. All coronal scales are fully sulcate below as are all species in this section. In Flora of the Fiji Islands (1972). J. W. Parham. Hoya diptera Seem, ex A. Gray Bulibuli Sewaro; Draubibi; Wabi, Climbing with 5 to 10 yellow flowers per umbel. Moderately common. — 22, 29, 35, 480, 488, 770. (Suva) In Flora Vitiensis Nova 4 (1988) 123. “A New Flora of Fiji” Albert C. Smith. 5. Hoya diptera Seem. in Bonplandia 9: 257* (Editors note: not on this page), nom. nud. 1861; A. Gray in Proc. Amer. Acad. Arts 5: 336, nom. nud. 1862, in Bonplandia 10:37, nom. nud. 1862; Seem. Viti, 439. nom. nud. 1862, Fl. Vit. 163. 1866; Drake, Ill. Fl. Ins. Mar. Pac. 236. 1892;Turrill in J. Linn. Soc. Bot. 43:33.1915; A. C. Sm. in Sargentia l:113. 1942; J. W. Parham, Pl. Fiji Isl. 186. 1964, ed. 2. 263. 1972………………. Figure 50. An often high-climbing vine with white latex, found at elevations from near sea level to about 900 m. in dense, dry, or open forest. The corolla is yellow (infrequently noted as cream-colored), sometimes faintly reddish-tinged at base on both surfaces and the corona lobes are pale yellow to white, often dull red or pink toward base. As far as collections are dated. flowers have been obtained between October and January and in June: no fruiting specimens have been noted. Typification: The type is Seemann 320 (K Holotype; Isotype at BM, GH); the holotype bears too specimens mounted on one sheet, one noted as from Taveuni and dated June. 1860. and the other from Viti Levu. It is not now possible to separate these and to indicate one as the Lectotype; they obviously represent the same species, and it seems best to treat them as jointly constituting the holotype. The binomial was mentioned several times before its description by Seemann in 1866. Distribution: Presumably endemic to Fiji, thus far known from three of the high islands. However, the Samoan species should be critically, examined in this connection; Hoya filiformis Rechinger (1908) and H. attenuate Christophersen (1935) appear scarcely separable from H. diptera, although both are recorded as having white corollas. Local Names: The recorded names are perhaps more strictly applicable to the better known Hoya australis: they are wa tambua, mbulimbuli sewaro, ndraumbimbi, and wa mbi. Available Collections: Viti Levu: Mba: Vicinity of Nandarivatu. Gillespie 4009; Nauwanga, south of Nandarivatu. Degener 14333. 14735. Serua: Namboutini, H B. R.

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Parham 497; flat coastal strip in vicinity of Ngaloa. Smith 9680. Namosi: Nakavu. on Navua River, Horne (K); track to Mt. Vakarongasiu, DA 16141. Naitasiri: Between Naomi and Nasonggo. DA 15301; Waimbau, Sawani. DA 11212; Central road. Tothill 60'8, vicinity of Nasinu. Gillespie 3556. Rewa: Government House paddock. Im Thurn 7. Vaniua Levu: Mathuta: Southern base of Mathuata Range. north of Natua, Smith 6775: southern slopes of Mt. Numbuiloa, east of Lambasa. Smith 6586. Thakaundrove: Savuthuru Mt., near Valethi. Degener & Orderez: 13832; Vatunivuamonde Mt.. Savusavu Bay region. Degener & Ordonez 14014. Fiji without further locality. U. S. Expl.. Exped., Horne 66. In Hoya Section Acanthostemma (Blume) R. D. Kloppenburg (1994) 31-34. R. D. Kloppenburg. Hoya diptera Seemann nomen in Flora Vitiensis (1866) 163; Bonplandia 9 (1861) 257. Scandens, glabra; ramulis 4-angulatis; foliis ovato-ellipticus acuminatis basi ovatis carnosis penninerviis; pedunculis compressis subulatis, petiolo longioribus; floribus flavis. —Viti Levu and Taviuni, on trees (Seemann ! n. 320) also collected by U.S. Expl. Exped. Perhaps a specimen, without flower, collected in the Isle of Pines (McGillivray), which has leaves twice as large as those of my specimens, may belong to this species, of which there are also imperfect specimens only. Translation: Climbing, glabrous stems 4 angled; leaves ovate-elliptic, acuminate with the base ovate, fleshy, penninerved; pedicels flattened laterally somewhat winged, the petioles longer; flowers yellow. A. C. Smith in Sargentia 2 (1942) 113 "Fijian Pant Studies" gives a lengthy description: Vines with slender glabrous or distally puberulent branchlets, sometimes the branchlets short, lateral, densely foliaceous; petioles rugulose, 5-15 mm. long, pale puberulent or glabrous; leaf-blades subcarnose, elliptic or ovate-elliptic or narrowly oblong. 3.5-8 cm. long, (1-) 2-3.2 cm. broad, obtuse base cuspidate or obtusely short acuminate at apex, pinnately-nerved. the costa slightly impressed or plane above, subprominent beneath, the secondary nerves 3 or 4 per side, ascending, oriented from costa toward base, prominent on both surfaces or submersed, the veinlet-reticulation immersed; inflorescences axillary, umbellate, the peduncle 10-45 mm. long, glabrous or obscurely puberulent, ellipsoid-capitulate at the apex, sometimes verrucose for the distal 5 mm., the bracts inconspicuous; flowers 5-10 per umbel at anthesis, the pedicels slender, 8-20 mm. long, glabrous or sparsely pale puberulent; calyx-lobes membranaceous, inconspicuous, deltoid, 0.7-1.1 mm. long and broad, glabrous except for the ciliate margin; corolla submembranaceous, 11-16 mm. in diameter, yellow, copiously and conspicuously puberulent within, the lobes deltoid or ovate-deltoid, 4-6 mm. long and broad, often recurved at margin, the sinuses acute; lobes of the corona thick, oblong, 34.2 mm. long, 1.6-1.8 mm. broad, obtuse at apex, acuminate at base, flattened above, rounded beneath; pollinia 0.5-0.6 mm. long; carpels glabrous. Viti Levu: Tholo North; Nauwanga, near Nandarivatu, alt. about 750 m., Degener 14333 (A) (vine, in open forest; corolla yellow, reddish toward center), Degener 14755 (A) (liana, in forest; corolla yellow); Naitasiri: Vicinity of Nasinu, alt. 150 m., Gillespie 3556 (BISH, UC) (liana with pendent inflorescences, in woods. Vanua Levu:

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Thakaundrove: Savuthuru Mt. near Velethi, alt. 90 m., Degener & Ordonez 13832 (A) (Vine in open forest; corolla yellow); Vatunivuamonde Mt. alt. 240 m., Degener & Ordonez 14014 (A) (vine in forest: corolla yellow). Without definite locality: Seemann 320 (Type coll., GH) (Viti Levu and Taveuni); U. S. Expl. Exped. (US). As represented by the cited specimens, this species is characterized by inconspicuous calyx-lobes, a comparatively thin corolla which is yellow and perhaps reddish tinged at base within and by the ascending secondary nerves of its leaf-blades. The cited type duplicate does not show the flattened sbualate pedicels mentioned by Seemann, and I am inclined to believe that this character, apparently the source of the specific name, was due merely to the degree of pressing of the actual type. In foliage the type duplicate is an excellent match for the other cited specimens. Additional herbarium sheets: Seemann Type #320 (A,K); Gillespie #4009, 3556; Degener #14333, 14755; Degener & Ordonez #13832, 14014; Horne #66; Pahan #497; DA #11212, 15301, 16141, A. C. Smith #9680 (UC), #7202, 6171, 6775, 6586; 62110, 6775 (US); Tothill #608; Thurn F-7. Additional references: Gibbs & Turrill in Journal of the Linnean Soc. of London 43 #288 (1915-1917) 33. Parham in Flora of the Fiji Is. (1972). Symonds in Hoyan 2 (1980) #2. A. C. Smith 4 (1988) 123 in Flora Vitiensis Nova. Comments: This species along with Hoya eitapensis Schlechter, Hoya nummularioides Constantine, and Hoya flavida Forster et Liddle belongs to a subsection of section Acanthostemma (Bl.) Kloppenburg; (Subsection Angusticarinatae Kloppenburg). These species have very narrow shelves on the sides of the coronal scales which protrude only slightly beyond or are equal in length to the outer apex of the scale. This represents a transitional stage between species without side modifications of the scale and those species with well developed side extensions termed bilobed since they extend beyond the outer coronal apex to varying degrees. All coronal scales are fully sulcate below as are all species in this section.

Hoya dischorensis Schlechter In Botanische Jahrbücher 50 (1913) 116. R. Schlechter. 15. H. dischorensis Schltr. n. sp. — Suffrutex epiphyticus parum ramosus scandens, Rami filiformes, flexuosi, laxe foliati, teretes, glabri. Folia patentia vel patula, breviter petiolata, oblonga, obtusiuscule acuminata, basi obtusa carnosula, utrinque glabra, petiolo brevi, superne leviter sulcato. Inflorescentiae umbelliformes 10 — 15 florae, breviter pedunculatae pedunculo pedicellisque glabris. Flores in sectione inter mediocres, illis H. Montanae Schltr. similes. Calycis foliola ovata, obtusiuscula, glabra quam corolla multi breviora. Corolla rotata, usque infra medium 5-fida extus glabra, lobis ovatis, acutis, margines versus dense villosulis, medium versus sensim papilloso-puberulis, basin versus glabratis. Coronae foliola patentia, subhorizontalia, superne subplana, oblonga, antice apiculata, extus obtusa paulo producta, lateribus incrassato-rotundata, apice, antice antheris paulo breviora. Pollinia oblique oblongoideo, basin versus sensim paulo attenuata, translatoribus brevissimis, retinaculo rhomboideo. Ein zierlicher, weinig verzweigter Schlinger mit schnurartigen, locker beblättern Zweigen. Blätter 6.5 — 11.5 cm lang, in der Mitte 2.8 — 5 cm breit, Blattstiele 0.5 — 1 cm lang, fleschig. Blütenstände doldig, 10 — 15 blütig, auf 1 — 1.5 cm langen Stielen

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Blütenstiele sehr fein, kahl, gegen 2 cm lang. Kelchzipfel klein, 2 mm lang, kahl korolla 1.6 cm im Durchmesser. Koronaschuppen von der Spitz bis zum äuszern Rande etwa 3.5 mm lang. Nordöstl. Neu-Guinea: auf Bäumen in der Waldern des Dischore-Geberges oberhalb Dschischungari im Waria-Gebiet, ca. 900 m u. M. (Schlechter n. 19834. — Blühend in Juni 1909). Die Art ist mit der oben beschriebenen H. reticulata Schltr. nahe verwandt, doch hat sie anders geformte Blätter oben die charakteristische Nervatur der H. reticulata Schltr. sowie eine andere Behaarung der Blüten und viel breitere Koronaschuppen Die Blüten sind kremegelb gefärbt. Translation: Same as the following entry. In Hoyas of New Guinea (1992) 42. R. D. Kloppenburg. (Translation): 15. Hoya dischorensis Schlechter n. sp. Epiphytic half shrub, few branches, climbing. Branches threadlike, flexible, loosely leaved, circular. glabrous. Leaves spreading or outspread, shortly petioled, oblong, somewhat obtuse, acuminate, base obtuse, a little fleshy, both sides glabrous, petiole short, slightly grooved above. Inflorescences umbellate. 10-15 flowered, slightly pedunculate, peduncle and pedicels glabrous. Flowers mid sized in the section, definitely similar to H. montana (montanae) Schlechter. Calyx segments ovate, somewhat obtuse, glabrous, much smaller than the corolla. Corolla rotate, 5 parted to just below the middle, lobes ovate, acute, toward the margins densely villose, toward the center gradually papillose-puberulous, base becoming glabrous, Scales of the corona spreading, almost horizontal, almost flat above, oblong, anterior apiculate, outside obtuse, a little elongated, sides thickly rounded, apex much shorter than anthers. Pollinia obliquely oblong, toward the base gradually a little attenuate, translators very short, retinaculum small, rhomboid. An ornamental, a cord like little branched vine, branches with few leaves. Leaves 6.5-11.5 cm. long, 2.8-5 cm. wide in the middle. Petiole 0.5-1 cm. long, fleshy. Inflorescence umbellate, 10-15 flowers, on peduncles 1-1.5 cm. long. Petiole very fine, bare to 2 cm. long. Calyx lobes small, 2mm. long, bare. Corolla 1.6 cm. in diameter. Corona scales, from the apex to the outer end about 3.5 mm. long. Northeast New Guinea: on trees in the forest of the Dischore Mountains above Dschischungari in the Wari region, about 900 m. altitude (Schlechter #19834 - blooming in June 1909). This species is closely allied to the above mentioned H. reticulata Schlechter but it has different leaf form and characteristic venation of H. reticulata Schlechter as well as different pubescence of the blooms and much broader corona scales. The flowers is creamy-yellow in color. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. dischorensis Schlechter (Schlechter 19834) – 15. In Dr. Schlechter’s Hoya Species (1993) 52-53. R. D. Kloppenburg. Hoya dischorensis Schlechter. Epiphytic half shrub, few branches, climbing. Branches threadlike, flexible, loosely leaved, circular. glabrous. Leaves spreading or outspread, shortly petioled, oblong, somewhat obtuse, acuminate, base obtuse, a little fleshy, both sides glabrous,

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petiole short, slightly grooved above. Inflorescences umbellate. 10-15 flowered, slightly pedunculate, pedicels of the peduncle glabrous. Flowers mid sized in the section, definitely similar to H. montana (montanae) Schlechter. Segments of the calyx ovate, somewhat obtuse, glabrous, much smaller than from the corolla. Corolla rotate, 5 parted to just below the middle, outside glabrous, lobes ovate, acute, toward the margins densely villose, toward the center gradually papillose-puberulous, base becoming glabrous, Scales of the corona spreading, almost horizontal, almost flat above, oblong, anterior apiculate, outside obtuse, a little elongated, sides thickly rounded, apex much shorter than anthers. Pollinia obliquely oblong, toward the base gradually a little narrow, with the translators very short, retinaculum small, rhomboid. An ornamental clinging cord like little branched vine, branches with few leaves. Leaves 6.5-11.5 cm. long, 2.8-5 cm. wide in the middle. Petioles 0.5-1 cm. long, fleshy. Inflorescences umbellate, 10-15 flowered, on peduncles 1-1.5 cm. long. Petioles very fine, bare to 2 cm. long. Calyx lobes small, 2mm. long, bare. Corolla 1.6 cm. in diameter. Corona scales, from the apex to the outer end about 3.5 mm. long. Northeastern New Guinea: on trees in the forest of the Dischore Mountains above Dschischungari in the Wari region, about 900 m. altitude (Schlechter #19834 - blooming in June 1909). This species is closely allied to the above mentioned H. reticulata Schlechter but it has different leaf form and characteristic venation of H. reticulata Schlechter as well as different pubescence of the blooms and much broader corona scales. The flowers are creamy-yellow in color. Publisher by Dr. R. Schlechter in: 1913 Botanische Jahrbücher 50:116, "Die Asclepiadaceen von Deutsch Neu Guinea".

Hoya diversifolia Blume In Bijdragen tot de Flora van Nederlandsche Inde (1826) 1064. C. L. Blume. Hoya diversifolia Bl. H. foliis aveniis ovalibus acutiusculis orbiculatisve carnosis glabris, corollis intus sericcis (Rumph. Herb. Amb. V. 175, fig. 2,). Crescit: in calcaris Kuripan et circa Bataviam. Floret: omni tempore. ** Corolla quinquefida, reflexa: corona staminae basi subulata; coronae foliolis angulo exteriore et incumbent longlier productis integerrimis. Translation: Leaves veinless oval acute orbicular fleshy glabrous, corolla incised silky (Rumphius Herbarium Amboinensis V. 175 figure 2) Growing: in calcareous Kuripan and around Batavia. Floret: entirely temporary. ** Corolla 5 parted, reflexed; corona staminea tubular at the base; lobes of the corona exterior angle and interior long produced absolutely entire. In Allgem Med-Pharm. Fl. 3 (1834) 1084. Kosteletsky. H. diversiflora, Blum. (Rumph. 5. t. 175. f. 2.) wächst ebenfalls blosz auf Bäumen, hat eben so wurzelnde Stengel, herab hängende Aeste, ovate, spitzliche ober fast kreisrunde, fleischige, aberlose, kahle, balaszgrüne. 1” lange Blätter, vielblüthige, kurze Dolden mit weissen Blüthen und fingerlange, bünne Balgkapseln. — Sie wächst auf Java und den Molukken, blüht das ganze Jahr und dient zu gleichem Zwecke, wie die schon erwähnten Arten.

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Translation: It thrives likewise naked on trees, with the main stem rooting, stems hanging down, ovate, with the apex pointed or slenderly rounded, fleshy, somewhat loose, glabrous, pale-green. Leaves 1 inch long, many flowered, short umbel with white flowers and a finger-long, round follicle. - They grow on Java and the Molukken, blooming the whole year and from the same sprig, like the already mentioned beautiful species. In General System of Gardening and Botany 4 (1838) 127. G. Don. 26. H. diversifolia (Blume. bijdr. p. 1064.) leaves veinless, oval acute, or orbicular, fleshy, glabrous; corollas silky inside. A woody shrub. Native of Java, in calcareous soils, about Kuripan and Batavia. Divers-leaved Hoya. Fl. year. Shrub. tw. In De Candolle Prodromus Syst. Veg. 8 (1844) 637. Decaisne. 16. H. diversifolia (Bl. bijdr. p. 1064), volubilis, foliis ovalibus v. oblongis rarius orbiculatis obsolete acuminatis v. acutisculis aveniis utrinque laevibus carnosis, umbellis breviter pedunculatis, corollis introrsum sericcis vix dimindium transversalem digitum latis albicantibus, laciniis crassis reflexis (woody shrub) in calcariis Kuripan et circa Bataviam (Blume). Sussuela esculenta Rumph. Amb. 5, p. 467. t. 175, fig. 2. (v. s. sin. fl. h. Mus. par.) Translation: twining, leaves oval or oblong rarely orbicular obsoletely acuminate or slightly acute both sides smoothly fleshy, umbels shortly pedunculate, corolla inside silky, divided transversely through the middle one half as wide, a dull white, leaflets thick reflexed. woody shrub, in calcareous Kuripan and near Batavia. (Blume). Sussuela esculenta of Rumphius in Herbarium Amboinensis 5 (1747) 467 table 175 figure 2. In Flora Indiae Bataviae 2 (1856) 518. Miquel. 6. Hoya diversifolia Bl. Volubilis, folia in iisdem ramis ovalia vel oblonga vel obversa, raro orbiculata, subapiculata vel acutiuscula, avenia, utrinque laevia, crasse carnosa, 4 — 2 poll. longa, umbella breviter pedunculatae (receptaculo florifero subellipsoideo), corollae introrsum sericeae, vix ½ digitum transversum latae, albicantis laciniae crassae reflexae. Hoya diversifolia Bl. Bijdr. p. 1064. Decaisn. l. c. p. 636. — Sussuela esculenta Rumphia. Herb. Amb. V. p. 467, tab. 175, fig. 2. Java, op de rotsen bij Koeripan, Batavia enz. (Bl.) — Molukken, fig. 2. Translation: Twinning, leaves far apart on branches, oval or oblong or obverse, rarely orbicular, somewhat apiculate or slightly acute, both sides smooth, thickly fleshy, 4 to 2 inches long, umbels shortly pedunculate (receptacle of the flower somewhat ellipsoidal), Inside of the corolla silky, barely ½ a finger wide, dull white, lobes thick reflexed.. etc. In Flora of British India 4 (1883) 61. J. D. Hooker. 36. H. diversifolia, Blume Bijdr. 1064; quite glabrous, leaves 2-3 in. broadly obovate or elliptic obtuse or apiculate very thick margines recurved, nerves indistinct, peduncles and short pedicels stout; corolla puberulous within, follicles very thick. Miquel Fl. Ind. Bat. ii, 518; Decne. in DC. Prodr. viii. 637. H. orbiculata, wall. in Wight Contrib. 36; Wall Cat. 8151; Decne. l. c. — Sussuela esculenta, Rumph. Herb. Amb. v. 467, t. 175, f. 2. Burma: at Prome, Wallich. Malacca, Maingay (Kew Distrib. 1130). — Distrib. Java, Amboina.

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A stout climber. Leaves 1.5- 2.5 in diam., base acute or rounded, nerves arched; petiole ¼ - ½ in. Peduncle ¼ - ½ in., very thick; pedicels about ½ in. Sepals small, elliptic, obtuse. glabrous (ciliate, Maingay). Corolla 1/3 in. diam,; cream-colored; lobes broad. Coronal-processes pale pink, short thick, ovate, with the broad end outwards concave above, with a boss near the center, inner angle produced into a short, erect spur shorter then the anther-tip. Follicles 6 in. long, ¾ in. dia,., falcate, sublinear; pericarp thick, apparently ¼ in. or more. Seeds about ¼ in. long. In Transactions of the Linnean Society 3 (1888) 321. H. Ridley. H. diversifolia, Blume. Kwala Pahang. Flowers rose-colour. In Revisio Genera Plantarum 2 (1891). O. Kuntze. H diversifolia, Bl. Java: Batavia. In Journal of the Royal Asiatic Society, Bengal Branch 2 (1903) 570. “Flora of the Malaya Peninsula” King & Gabble. 12. Hoya diversifolia, Blume Bijdr. 1064 (1826). A climbing glabrous shrub; branches flexuous, smooth, brown; branches paler; the bark very thin. Leaves fleshy, very thick; broadly elliptic or obovate, obtuse or apiculate at apex, rounded at base; both surfaces glabrous, dull; margins recurved; 2 to 4 in. long, 1.5 to 2.5 in. broad; nerves only visible when dry, midrib broad; main nerves about 4 to 6 pairs, rather irregular, at about 60° with the midrib, soon branching; reticulations very indistinct; petiole .25 to .5 in. long tubercular rachises at the ends of very thick .5 to 2.5 in. long peduncles, which are lateral one to each pair of leaves; pedicels slender, .5 in. long; buds flattened with 5 rounded angles; flowers .25 to .35 in. broad, white or rosecoloured with pink corona (yellowish-white (Maingay)). Calyx membranaceous; lobes elliptic-obtuse, usually ciliate at the tips. 0.75 in. long; scales apparently none. Corolla glabrous without , puberulous within; lobes apparently none. Corolla glabrous without. puberulous within; lobes cordate-acuminate. Corona of 5 processes attached to the bases both of the staminal-column and of the corolla-tube; lower lobe of each process orbicular or ovate, concave above and with a small central boss. 2-wingewd below, the wings connivent; upper lobe erect, triangular-acute. shorter than the anther-appendages. Staminal-column very short; anther-cells parallel above, divergent and empty below; appendages scarious. acute or acuminate; pollen-masses oblong, truncate at top, thin on the outer margine, attached by very short thick caudicles to the rather long orbicular pollen-carriers. Style-apex 5 angled, depressed, with short conical apiculus. Follicles sublinear, falcate. 6 in. long, .75 in. broad; pericarp thick, apparently .25 in. or more. Seeds oblong, .25 in. long. Decne in DC Prodr. VIII. 636; Miq. Fl. Ind. Bat. II. 518; Hook. f. Fl. Br. Ind. IV. 61. H. orbiculata. Wall. in Wight Contrib. 36; Wall. Cat. 8151: Decne. l.c. 635; Miq. l.c. 519. “Sussuela esculenta” Rumph. Herb. Amb. V. 467, t. 175 f. z. Penang: at Pulau Boetong, Curtis 928. Perak: Wray 2718; Scortechini 118. Malacca: Griffith 3787; Maingay (K.D.) 1130. Pahang: at Pekan, Ridley 1619. Singapore: Ridley 1626, 2733 — Distrib. Burma (at Prome, Wallich 8151), Java, Amboina. Leaves moderate-sized, under 4 in. long; elliptic or ovate; main nerves indistinct, at about 60° with the midrib; climbing ........................ 12. H. diversifolia.

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In Fragmenta Florae Philippinae 1 (1904) 129. R. Schlechter & O. Warburg. 2. Hoya diversifolia Blume ? Culion Isl. (Merrill no. 585). In Systematisches Verzeichnis Family 1 (1911) 6. S. H. Koorders & Ann Schumacher. Hoya diversifolia ? Bl. — Windender Strauch. West-Java: Batavia: Tjiampea: Region I: Kds 30721  (936 * 8 VII. 1898. Nur mit Blättern. Bestimmung sehr zweifelhaft). Translation: A twining shrub. In Exkersionsflora von Java 3 (1912). S. H. Koorders & Ann Schumann. (Key). 9c. Blätter 2 ½ — 7 ½ x 2 ½ — 5 cm, vielgestaltig, von ± rundlich bis breit ovat odor elliptisch, oben stumpf oder apiculat, sehr dick, völlig kahl. Blattstiel ½ — 1 1/3 cm. Pedunculi bis 1 1/3 cm, sehr dick. Pedicelli ± 1 1/3 cm. Korolle ± ¾ cm breit, gelblichweisz. Coronazipfel blasz purpurn. Follikel 15 x 2 cm, sichelförmig. Winderder Strauch. Hook. l. c. 61; Miq. l. c. 518; H. orbiculata Wall. Java: Ohne näheren Standort (Blume in Herb, Leiden). Im Buitenzorger Herbar fand ich kein einwandfreies Material von dieser Art, während ich im Hortus Bogor. lebende (vielleicht authentische) Exemplare sah. Vielleicht gehört hierber ein steriles Spezimen (Kds. n. 30712 ) aus Tjampea bei Buitenzorg. ....................................................................H. diversifolia Bl. * Translation: 9c. Leaves 2 ½—7 ½S x 2 ½—5 cm, multiform, of ± roundish until wide ovat or elliptical, above dully or apiculate, very thick, entirely glabrous. petiole ½ - 1 1/3 cm. Peduncle up to 1 1/3 cm, very thick. Pedicel ± 1 1/3 cm. Corolla ± ¾ cm wide, whitishyellow. Corona apex pale purple. Follicles 15 x 2 cm, sickle shaped. Winding shrub. Hook. l. c. 61; Miq. l. c. c. 518; Hoya orbiculata Wall. Java: Without nearer location (flower in sharp, ailments). I found no perfect material of this type in the Buitenzorger Herbar, while I was in the Hortus Bogor living (maybe authentic) copies I saw. Maybe a sterile Spezimen belongs here (Kds. N. 30712) from Tjampea with Buitenzorg. In Flore Generale de l'Indo-Chine 4 (1912) 131-132. J. Constantine. 5. H. diversifolia Blume Bijdr., p. 1064; Miq. Fl. Ind. Bat., II, p. 518; Dcne in DC Prodr., VIII, p. 637; H. orbiculata Wall. ex Wight Contrib., p. 36; Boerl. Hand. Fl. neder. Indie, II, p. 439; Sussuela esculenta Rumph. Herb. Amb., V. p. 467, tab. 175. Epiphyte. Tige vigoureuse, pouvant devenir assez epaisse, volubile, tres finement pubescente, puis glabre. Feuilles tres charnues, orbiculaires ou largement ovales, arrondies, parfois un peu aigues a la base, arrondies ou obtuses au sommet, ou avec un large mucron, legerement pubescentes vers le milieu et le base; nervures rarement visibles; limbe long de 1.5 - 10 cm. sur 4.5- 6 cm.; pedoncule pubescent, parfois assez epais, long de 4 cm.; pedicelles pubescents, longs de 1.5 cm.; fleurs larges de 8 mm. avant l’epanouissement. — Calice a sepales pubescents. Corolle: lobes triangulaires, pubescents en dedans, se rabattant entre les pieces de la coronule, Coronule a 5 pieces assez epaisses, peu aigues, ou obtuses, largement ovales, faiblement concaves en dessus, avec une cote au milieu; bec posterieur accuse, elegant les membranes connectivales des antheres. Etamines: pollinies ordinaires, aplaties, arrondies a l’extremite, avec une bordure inclore; anthere a membrane connectivale dressee sur le style. Fruit ... — Fig. 18, p. 131.

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Laos: bassin du Se-moun, forets (Harmand). — Cochinchine: mont Kaki (Harmand). Cambodge: (Harmand). — Java, Malacca. ** Feuilles mucronees a l’extremite, non en coin a la base. * Feuilles tres epaisses; nervures secondaires non visibiles ......... H. diversifolia. Translation: Epiphyte. Vigorous stem, can become thick, twining, very finely pubescente, then glabrous. Very fleshy leaves, orbicular or extensively oval, rounded leaves, a few leaves sharp at the basis, either rounded at the top or obtuse, or with a large mucron, slightly pubescentes toward the middle and the bases; rarely visible nerves; long limb of 1.5 - 10 cm. on 4.5 - 6 cm.; stalked pubescent, sometimes very thick, as long as 4 cm.; pedicelles pubescents, as long as 1.5 cm.; large flowers of 8 mm. before blossoming. Calyx has pubescents sepals. Corolla: triangular lobes, pubescents inside, moving back between pieces of the corona, Corona has 5 pieces thick pieces, a little sharp, or obtuse, extensively oval, weakly concave on top, with a umbo near the middle; posterior beak acute, elegant membranes connected to the anthers. Stamens: pollen ordinary, flattened, rounded at the extremity, with a solid colored border; anther is membranaceous trained connected on the style. Fruit... - Fig. 18, p. 131. Laos: at the base of the mountain, in forests around (Harmand). - Cochinchine: Mount Khaki (Harmand). Cambodia: (Harmand). - Java, Malacca. * * Leaves mucronate at the extremity, no in corner has the basis. * Leaves very sparse; secondary veins not visibiles......... H. diversifolia In Florae Siamensis Enumeratio (1951) 35. W. G. Craib & A. F. G. Kerr. Hoya diversifolia Blume Bijdr. 1064 (1826); F.B.I. iv 61; Mat. F.M.P., no. 19, 570 (780); F.M.P. ii. 396; F.I.C. iv. fig. 18. Ridl. i 137. Ubon. Valley of the River Mun, dry forest, Harmand (ex F.I.C.) Surat. Chumpawan, Haniff et Nur 4387! Puket, Lang-kawi (ex Ridl.). Nakawn Sritamarat. Songkla, Ban Pien, under 50 m., in scrub, Kerr 14816! Distr. Burma! Cambodia (ex F.I.C.) , Pen. Mal.! Java! (type) Borneo! Local name: Lin Kwai (.......) , Siamese, Songkal. In Flora of the Malay Peninsula 2 (1923) 396. H. Ridley.(4) H. diversifolia Bl. Bijdr. 1064; King l.c. 579. Long climbing. Leaves fleshy very thick broad elliptic or blunt ovate, base round; nerves 4 to 6 pairs (visible only when dry); 2 to 4 in. long, 1.5 to 2.5 in wide; petioles .25 to .75 in. long. Peduncles thick; pedicels .5 in long. Flowers greenish white with corona pink or all pink, .25 to .35 in. wide. Corolla velvety inside, lobes ovate blunt. Corona lower lobes oval blunt. Pollinies sub linar.6 in. long, .75 in. roundish. Herb common often draping trees especially near the sea. Singapore, Jahor, Serangoon, Kranji, Jahor, Sendai River,; Kukub. Malacca, Panjong Kling. Pahang, Perak, Pulau, Pekan; Sungei Jelai (Machado) Negri Semindan, Bukit Sulu (Cantley). Perak, Kwala Kangsa Province Welleslev, Bukit Kawan, Penang, Pulan Betong. Waterfall Curtis' Kedah, Alor Stat Gumong Geriang river bank. Distrib. Burma, Java. Native names: Akar Sesudu Bukit, Akar Chapang Kra.

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In Illustrated Guide to Tropical Plants (1960) 661. Hoya diversifolia Bl. Large climber, white latex. Leaf 5 — 14 cm long, fleshy, thick. Flower 9 — 14 mm wide, pink to reddish violet. Pods 13 cm long. In Flora of Java 4 (1965) 269. (Key). 12 a. Leaves with a very obtuse-broadly rounded emarginate, rarely shortly narrow top, nearly always with minute broadly triangular apical cusp, broadly oval or obovate-oblong, with distinct midrib and very obscure nerves, glabrous, with 1 — 2 very thick trichomes, fleshy, coriaceous when dry, 5 — 14 cm, by 3 — 5 cm; petiole thick, glabrous ¾ — 1 ½ cm; stem glabrous. Peduncle thick, glabrous, 2 ½ — 4 cm long; rachis producing at intervals groups of flowers; pedicels glabrous 1 ½ — 2 cm, much thickened under the fruit; adult flower-buds flat; calyx segments oval, obtuse, ciliate, 2 — 2 ½ mm long; corolla 9 —14 mm diam., shallowly lobed; lobes acute, or seemingly obtuse because of the recurved top, outside pale, glabrous inside tinged with violet, and densely appressed white-hairy especially along the margins; corona-angle, concave and with a subcentral knob above; apical membrane of anthers exceeding the conical top of the stigma; pollinia oblong, pellucid-margined; follicles lanceolate, rather obtuse, glabrous. c. 13 cm long; seed c. ½ cm long; coma c. 3 cm. Not rarely terrestrial and rather high-climbing, mostly however epiphytic. 1.00 — 4.00; I —XII; W.C.E., Bawean. Mad.; 1 — 400; Campong and road side trees, open forest (? H. crassipes Turcz., — H. zollingeriana Miq.) .............................. H. diversifolia Bl. In Journal of the Royal Horticulture Society (1973). A. J. Wills. H. diversifolia. A strong grower. In Malayan Wildflowers (1974) 298. M. R. Henderson. Leaves not hairy, flowers smaller. Leaves very thick, usually rounded at tip and base, petals usually blunt, the processes from the stamen column rounded .................... 4. diversifolia 4. H. diversifolia (oval-leaved Hoya). Fig. 282. Stem smooth, leaves very thick, usually elliptic in outline, sometimes oval, tip rounded, or with a broad, short point, base rounded, from about 2 in. x 1 in.. or sometimes smaller, to about 1 ½ to 2 ½ in. leafstalks from very short to nearly 1 in. long; inflorescence stalks thick, reaching about 2 in. long, the thickened , rough end from which the flowers arise lengthening to about 1 ½ in., flowers rather small, 1/3 -2/5 in. wide, on stalks about ½ in. long, corolla velvety hairy inside, pinkish, sometimes greenish, the flat, blunt, spoon shaped processes from the stamen column deeper pink, or purplish; fruit pods long and slender, up to 5-6 in long, plume of seeds 1 ½ - 2 in long. Common on trees near the sea, sometimes on inland river banks. In Malayan Nature Journal 30 (1978) 518-519. Rintz. 240 Hoya diversifolia Blume, Bijdr. (1826) 1064. Type: Java, Blume — Fig. 29. Distinguishing features: Stem stout. Leaves fleshy, elliptical-ovate with cuneate basis, obtuse apically; c. 13 cm long by 5 cm wide; veins not visible. Peduncle horizontal, rigid, 3 — 5 cm long. Umbel negativegeotropic, convex with rigid, uniform pedicels c. 2 cm long; 1 — 20 flowers. Corolla spreading, densely pubescent inside; c. 1.3 cm diam; pink. Corona lower lobe flat; both lobes pink. Follicles c. 14 cm long by 6 mm diam. Ecology: throughout the peninsula in

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lowland forests; common on limestone hills along the coasts, occasionally mixed and easily confused with H. parasitica when not in bloom. Distribution: Thailand, Indo China, Sumatra, Borneo, Java. In Asklepios 33 (1985) 62-63. L. Wood. Hoya diversifolia. Described as a beginners plant, one may find it easy to grow but there is a lesson to be learned from this plant as there is with all other Hoyas. I was excited at the sight of three large umbels appearing on Hoya diversifolia, one day in June of 1903. I began a constant watch so that I could get a photograph of the plant when it eventually flowered. On my visits to the greenhouse I noticed the buds getting larger all the time. One particular dinnertime on returning home from work, expecting to see my plant in flower, I suddenly became horrified when I noticed that at least one third of the buds had dropped on to the floor. Looking at the hygrometer (humidity meter) I saw that the humidity had dropped to thirty per cent. Realizing what had happened I immediately filled a large tray with water and a little gravel, and stood the pot on top of the gravel. Then I filled two large containers with water and stood these near the plant, after this I damped the floor with water and managed to get the humidity back up to fifty per cent. Twenty four hours after, I was rewarded; the flowers had opened and filled the greenhouse with one of the sweetest scents imaginable. Click click went the camera and the job was completed. Hoya diversifolia - largest leaf on my plant is approximately four and a half inches long and two to two and a half inches wide. Leaf very fleshy, mid green on the top of the leaf and light green below it. Various leaf shapes appear on the plant at the same time. The umber was some two and a half inches in diameter and consisted of thirty one flowers. The corolla is approximately a half inch across and is slightly off white in colour, the corona is about a quarter of an inch across and described as rose coloured by one grower, and drips nectar which stains the corolla and causes different colours to show, from tawny yellow to pink. In conclusion may I say that although Hoya diversifolia may be a beginner plant, it is certainly not one to be missed. Keep the humidity up above fifty per cent and your Hoyas will grow and flower well. Have you caught the Hoya bug yet? In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. diversifolia Blume – 32. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 86 “The Genus Hoya in Thailand” O. Thaithong. H. diversifolia Blume, Bijdr. (1826) 1064. FBI 4 (1883) 61; FMP 2 (1923) 396; Malay. Wild Fl. Dicot. (1959) 298, Fig. 282; Malay. Nat. J. 30 (1978) 517, Fig. 29. Occurrence: (PEN) Surat Thani, Chuphon, Nakhon Si Thammarat, Puket, Songkhla, Ranong (SW) Pachuab Kiri Khan (SE) Chon Buri.

Hoya dodecatheiflora Fosberg

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In Lloydia 3 (1940) 118-119. Fosberg. 42. Hoya dodecatheiflora Fosberg, sp. nov. Scandens; folia elliptico-lanceolata, crassa, inaequalia:, maxime 8 cm. longa,. umbella axillaris, multiflora, pedicellis gracilibus, 1 cm. longis; flores 5 mm. longi; calyx minutus; corolla reflexa, valde lobata., intus pubescens, squamis coronae lanceolatis, erectis, sulcatis, 3 mm. longis; pollinia 0.5 mm. longa; carpella 1.2 mm. longa. Slender vine, glabrous (except corolla), rooting at nodes, internodes 3.5-6.S cm. long; leaves opposite, elliptic-lanceolate, apex acute to acuminate, base acute, or smaller leaves obtuse, blade fleshy, unequal in size, even at same node, up to 8 cm. long and 2 cm. wide, secondary veins obscure, margins slightly resolute, petiole fleshy, up to 9 cm. long; peduncle axillary, one at a node, up to 9 cm. long, the lower 4-5 mm. fused with petiole of leaf, ending in a simple or bifid or fleshy pulvinus, this being the center of a many-flowered umbel of small flowers, pedicels slender, about 1 cm. long; calyx divided almost to base, lobes unequal, ovate or triangular, 0.7 mm long, apex blunt; corolla strongly reflexed, divided more than two thirds to base, lobes sharply triangular slightly acuminate, 2.5 mm. long, tube 1 mm. long, corolla glabrous outside; lobes strongly pubescent inside, tube puberulent inside; corona scales lanceolate, erect, apices prolonged, connivent, bases broader, rounded, deeply notched, surface shining but striate, longitudinally sulcate with two deep folds in the dower portion leading to notch at base, three sharp ridges above, the whole 3 mm. long, pollinia 0.5 mm. long, pale yellow, waxy, the connecting portion dark brown, anthers fused with stigma, carpels 1.2 mm. long, free except the stigmas; whole flower from tip of corona scales and anthers to tips of reflexed corolla lobes about 5 mm. long; fault unknown. Santa Crux Islands, Vanikoro, Tevia Bay, on trees, Bray 6, 1933, Stewart (type). The flowers of this species resemble superficially those of certain Solanum species, or even more so those of the primulaceous genus Dodecatheon, hence the name. Not closely related to any species that I have seen. Translation: Climbing, leaves elliptic-lanceolate, thick, unequal, up to 8 cm. long, umbels axillary, many flowered, pedicles slender, 1 cm. long; flowers 5 mm. long; calyx minute; corolla reflexed, deeply lobed, inside pubescent, scales of the corona lanceolate, erect, grooved, 3 mm. long; pollinia 0.5 mm. long; carpels 1.2 mm. long.

Hoya dolichosparte Schlechter In Beihefte zum Botanischen Centralblatt 35/2 (1916) 13-14. "Neu Asclepiadaceen von Sumatra und Celebes" R. Schlechter. Hoya dolichosparte Schltr. n. sp. Epiphytica, ramosa; ramis filiformibus, elongatis, laxe foliatis, glabris, lenticellis numerosis verruculosis; foliis patulis, petiolatis, late ovalibus, obtusis , nervis primariis 5 supra prominulis, utrinque glabris, 9 —13.5 cm longis; medio fere 6.5—9 cm latis, petiolo carnoso 1.5—2 cm longo; cymis umbelliformibus multifloris, pedunculo pedicellisque glabris. c. 2 cm longis; floribus in genere mediocribus; calycis foliolis ovatis, obtusis, glabris, extus sparsim verruculosis, 2 mm longis; corolla subrotata, c. 1.3 cm diametiente, usque supra medium fida, extus glabra intus minutissime papilloso-puberula, lobis late ovatis, breviter acuminatis; coronae foliolis dorso leviter adscendentibus, superne ovatoellipticis apice breviter et obtuse acuminatis, extus subacutis, lateribus incrassatorotundatis, antheris paulo brevioribus; polliniis oblique oblongoides, erectis,

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translatoribus perbrevibus triangulis, retinaculo rhomboideo polliniis fere 4-plo breviore. Celebes: Auf groszen Bäumen bie Toli-Toli, c. 10 m u. d. M. (R. Schltr. Nr. 20642 — blühend im Januar 1910). Eine Art aus der Verwandtschaft der Hoya bandaensis Schltr., aber mit weiszlichen Blüten mit weiszer. in der Mitte roter Korona. Translation: See below. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. dolichosparte Schlechter (Schlechter 20642) – 47. In Dr. Schlechter’s Hoya Species (1993) 54. R. D. Kloppenburg. (Translation of German text). Hoya dolichosparte Schlechter. Epiphytic, branched, branches threadlike, elongate, loosely leaved, glabrous with numerous wart like lenticels; leaves outspread, petiolate, broadly oval, obtuse, with 5 prominent primary nerves above, both sides glabrous 9-13.5 cm. long, almost 6.5-9 cm. wide at the middle, petiole fleshy 1.5-2 cm. long; cymes shaped like umbels, many flowered, peduncle and pedicels glabrous about 2 cm. long; flowers in general mid sized; with the lobes of the calyx ovate, obtuse, glabrous, outside sparsely wart like, 2mm. long; corolla somewhat rotate, about 1.3 cm. in diameter, 5 lobed all the way to above the middle, outside glabrous, inside minutely papillose-puberulous, lobes widely ovate, briefly acuminate; with the scales of the corona at the outside slightly ascending, above ovate-elliptic, tip short and obtuse, acuminate, outer somewhat acute, lateral thickly rounded, anthers small short; pollinia obliquely oblong, erect, translators very small triangular, retinaculum rhomboid, nearly 4 times smaller then the pollinia. Celebes: on large trees near Toli-Toli, about 10 meters altitude (R. Schlechter #20642-- blooming in January 1910). A species from the Hoya bandaensis Schltr. Group but with strikingly long, very high climbing branches and whitish flowers with a white corona with a red center. Published by Dr. R. Schlechter in: 1916 Beihefte zum Botanischen Centralblatt V.35 #2 p.13-14 "Neu Asclepiadaceen von Sumatra und Celebes".

Hoya edanoi C. M. Burton In The Hoyan (1990) 2/I . Hoya edanoi C. M. Burton, nov. sp. Acanthostemma Section. Ramulis crassis 3 mm. in diametro, glabris, petioli 0.5-1.5 cm. longis; foliis crasse coriaceis, rotundatis vel oblongis, cordatis vel fere peltatis* (word corrected), apicis rotundatis vel mucronatis, 3-7 cm. longis x 3-5 cm. latis. Pedunculatis 3-12 cm. longis; pedicelis filiformibus 2-3 cm. longis. Sepalis ovatis vel rotundatis, obtusis 1 ½ mm. latis et longis. Corollis 10-12 mm. in diametra, intus puberulis, albidis; coronae lobis 3 mm. longis, apicem interiore versus acutis, apicem exteriore rotundatis, auriculis 2 supra apicem productis. Typus: G. Edano, Bureau of Science #11655 (A) Lake Balinsasayao, Tanjay, Negros Oriental, The Philippines, 21 May 1949; M. Ramos and G Edano, Bureau of Science #44121 (US), (B), Tawitawi, Sulu Province, The Philippines July-August 1924 . The species differs from others in the section in the following ways: 1). The corolla lobes are not reflexed on the specimens examined. This may be due to age of the flowers as many of those that are normally reflexed open flat, reflex flatten and then close

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before falling. I have never seen another of the Acanthostemmas do that. 2). The entire inner side of the corolla is puberulous, including the outer tips (usually bare in this section). 3). The corona is only slightly elevated in the center (Others of the section are so elevated as to appear nearly upright). The flatness of the corona could be due to pressing. 4). The flowers of this species are much larger than any other known flowers of this section. Translation: See below. In Hoya Section Acanthostemma (Blume) Kloppenburg (1994) 35-37. R. D. Kloppenburg. Hoya edanoi Burton ex Kloppenburg. Hoya edanoi Burton in Hoyan 13 #1 (1991) pt. 2:3. Ramulis crassis 3 mm. in diametro, glabris, petioli 0.5-1.5 cm. longis; foliis crasse coriaceis, rotundatis vel oblongis, cordatis vel fere peltatis* (word corrected), apicis rotundatis vel mucronatus, 3-7 cm. longis. Sepalis ovatis vel rotundatis, obtusis 1.5 mm. latis et longis. Corollis 10-12 mm. in diametra, intus puberulis, albidis; coronae lobis 3 mm. longis, apicem interiore versus acutis, apicem exteriore rotundatis, auriclis 2 supra apicem productis. Typus: G. Edano, Bureau of Science #11655 (A) Lake Balinsasayao, Tanjay, Negros Oriental, Philippines, 21 May 1949; M. Ramos & Edano Bur. Sci. #44121 (UC,US,B), Tawitawi, Sulu Province, Philippines. J.-A. 1924. Translation of Latin: Branches thick 3 mm. in diameter, glabrous, with the petiole 0.5-1.5 cm. long. Leaves thick leathery, round or oblong, cordate or nearly (no such Latin word) peltate ? (shield shaped), with the tip round or mucronate 3-7 cm. long x 3-5 cm. wide, with the peduncles 3-12 cm. long; with the pedicels threadlike 2-3 cm. long with the sepals ovate or round, obtuse 1.5 mm. wide and long. With the corollas 10-12 mm. in diameter, inside puberulous dull white; with the lobes of the corona 3 mm. long; with the interior tip towards acute, with the exterior tip round, with 2 auricles produced above the apex (tip). The stems, leaves, petioles, peduncles, pedicels and calyx are all glabrous in this species (not mentioned in the original description). On herbarium sheet #44121 at (UC) the corolla is reflexed and the diamond shaped apex area of the corolla lobes inside are glabrous as is always the case in this section. The bilobed extensions from the side of the outer coronal lobe arise from deep on the side, extend beyond the outer apex and touch at their tips. Additional notes: Calyx 0.36 cm. in diameter, lobes 0.12 cm. long, with apex rounded, membranaceous, with overlap at the base 1/4, 5 veined, with tiny occasional cilia, ligules present. Ovaries relatively long and narrow. Corolla reflexed, inside puberulous except for the glabrous apex area, outside glabrous diameter 1.14 mm. in diameter flattened. Sinus to Apex 0.37 cm.; widest portion of lobe 0.30 cm.; sinus to center 0.27 cm. On the (UC) specimen the flowers are red and the umbel has 23 flowers. References: Kloppenburg in Philippine Hoya Species 2nd Ed. (1993) 55. In Philippine Hoya Species 3rd Edition (1996) 67-68. R. D. Kloppenburg. Hoya edanoi Burton in Hoyan 12 #2 pt.2 (1990).Type: #11655 (A) collected by G. Edano at Lake Balinsasayao Tanjay, Negros Oriental, Philippines. Section Acanthostemma (Blume) Kloppenburg. Key: not in key.

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A branching epiphytic glabrous vine with rather fleshy, broadly ovate foliage with cordate bases and mucronulate apices. Leaves pinnately veined 3-7 cm. long and 35 cm. wide. Peduncles 3-12 cm. long and glabrous as are the threadlike 2-3 cm. long pedicels. Sepals ovate to round 1.5 mm x 1.0 mm tall, membranaceous, overlapping 1/4 at the base and with ligules at the inside bases. Corolla revolute, inside puberulous, except for the apical area which like other Acanthostemma Sectional species is glabrous. Flowers in flat umbels, 1-1.2 cm. in diameter flattened. Corona rather flattened but sloping upward to the inner apex. The bilobed ligule-like side extensions are rather broad, flattened on the upper surface and ornately linearly lined (sulcate). The extend considerable beyond the outer apex and the outer apex are curved upward. The ovaries are long and narrow like those of H. merrillii Schlechter. Meaning: edanoi — Named in honor of the collector, G. Edano.

Hoya edeni King ex Hooker In Flora of British India 4 (1883) 53. J. D. Hooker. 3. H. Edeni, King in Herb Hort. Calc. Centrostemma sp., Herb. Ind. Or. H. f. 7 T. Sikkim Himalaya, alt. 3-6000 ft., Hreb. Griffith, J. D. H., & c. Stem rather slender, climbing and rooting. Leaves 3-4 by 1- 1 ¾ in., elliptic or obovate-lanceolate, obtusely caudate-acuminate, base acute, thinly coriaceous, puberulous or glabrate beneath; petiole 1/12 in. pubescent. Umbels terminal; peduncle ¼ - ½ in.. pedicels 1 in. Sepals small, ovate, obtuse. Corolla pubescent within; lobes ½ in. long, subacute. Coronal-processes horny, deeply grooved down the back, much exceeding the anthers. Follicles 4 in. long, slender, straight. Seeds 1/8 in. long. In Die Naturlichen Pflanzenfamilien 4(2) (1895). K. Schumann. Sect. II. Ancistrostemma Hook. fl. Blkr. Zurückgebrochen, mit schmalen Zipseln; Gynostegium sitzend; Coronzipsel am Grunde gebruckelt, oben in 2 ausrechte Spitzen vorgezogen, von denen jede in einen 2 spaltigen, eigekrümmten Haken ansgeht, H. Edenii King vom Sikkimhimalaya. Translation: Bent backwards, with narrow ends; Gynostegium sedentary; Corona points at the base bent-back, above with 2 groves on tops, from which each one splits into 2 special curved hooks at the end, H. Edenii King from the Sikkim Himalaya. In Indian Botanical Survey Record 4 (1903) 222. 595. Hoya Edeni King. Tong, 5000 ft., No. 891. In Flora of the Eastern Himalaya (1966) 261. H. Hara. Hoya Edeni King ex Hook. f., Fl. Brit. Ind. 4: 53 (1883). Mul Pokhari Nissim, 1600-19000m. (Oct. 30, 1963, ft.); Chyangthaphu-Birwa, 1300- 2400m (Nov. 27, 1963, fr.); Birwa-Yektin. 2900-1600 m (nov. 28, 1963). Distr. E. Himalaya (Nepal, Sikkim), and Manipur.

Hoya eitapensis Schlechter In Botanische Jahrbücher 50 (1913) 109. “Die Asclepiadaceen von Deutsch Neu Guinea” R. Schlechter. 4. H. eitapensis Schltr. n. sp. Suffrutex epiphyticus, ramosus scandens. Rami filiformes, flexuosi, laxe foliati, teretes, glabri. Folia erecto-patentia

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elliptica, obtusiuscula, basi cuneata, carnosa, glabra, brevissime petiolata vel subsessilia. Inflorescentiae umbelliformibus pedunculatae, folia superantes, 20-35 florae, pedicellis gracilibus, inaequilongis. Flores in sectione inter minores. Calycis segmenta ovatatriangula obtusiuscula, minutissime et sparsissime puberula et ciliolata, corolla multo breviora. Corolla rotata apicibus reflexis, usque ad tertiam partem basilarem 5-fida, lobis ovatis acutis, extus glabris, intus minutissime papilloso-puberulis. Coronae foliola horizontalia oblonga, antice leviter adscendentia, acuminata, extus obtusa, lateribus rotundato carinata, antheris vix longiora. Pollinia oblique oblongoideo, apice subfalcata, translatoribus quam maxime abbreviatis, retinaculo rhomboideo, minuto. Ein zierlicher Schlinger mit fadenförmigen, locker beblätterten Zweigen. Blätter 3 5 cm lang, in der Mitte 1.2-2 cm breit. Blattstiele kaum 2 mm überschreitend. Blütenstande doldig auf etwa 5 cm langen Stielen. Blütenstiele sehr schlank, bis 2 cm lang, nach der Spitze zu sehr kurz und spärlich behaart. Kelchzipfel ca. 1.75 mm lang, sehr kurz und spärlich behaart. Korolla 0.8 cm Durchmesser, innen sehr kurz papillös. Koronaschuppen von der Spitz bis zum äuszeren Rande etwa 0.2 cm lang. Nordöstl. Neu-Guinea: auf Bäumen in der Waldern in der Umgebung von Eitape, ca. 20 m. u. M. (Schlechter n. 19964. Blühend im August 1909). Diese und die beiden folgenden Arten habe ich an den Anfang der Sektion gestellt, da sie sowohl im Habitus wie der geringen Grösze ihrer Blüten den Arten der Sektion Otostemma am nächsten stehen, Die vorliegende unterscheidet sich von H. microstemma Schltr. durch die gröszeren Blätter und die kürzer behaarte Korolla sowie durch schmälere Koronaschuppen. Ihre Blüten sind gelblich-weisz. Translation: See next below. In Hoyas of Northeastern New Guinea (1992) 19-20. R. D. Kloppenburg. (English Translation). 4. Hoya eitipensis Schlechter n. sp. A twining perennial branched plant, woody at the base, epiphytic. Branches round, thin, flexible and glabrous, loosely leaved. Leaves erect, spreading, elliptic, somewhat obtuse, with wedge shaped base, fleshy, glabrous with very short petioles, almost sessile. Inflorescences umbellate, with the peduncle overtopping the leaves, 20-35 flowered, pedicels slender, unequal in length. Flowers small for the section. Calyx segments ovate triangular, somewhat obtuse, minutely and sparingly puberulous and ciliate, much smaller than the corolla. Corolla rotate, apices reflexed, 5 parted up to 1/3 from the base, lobes ovate acute, outside glabrous, inside of minute soft hairs. Corona scales horizontal, oblong, interior slightly elevated, acuminate, exterior obtuse, sides rounded and keeled, anthers barely longer. Pollinia obliquely oblong, apex somewhat curved, translators much shorter, retinaculum rhomboid, minute. A twining ornamental with threadlike loosely leaved branches. Leaves 3-5 cm. long, in the middle 1.2-3 cm. wide. Petiole hardly 2 mm. overall. Inflorescence umbellate on about 5 cm. long peduncles. Peduncles very slender, up to 2 cm. long, toward the apex it is very shortly and sparsely pubescent. Calyx lobes about 1.75 mm. long very shortly and sparsely pubescent. Corolla 0.8 cm. in diameter, on the inside very shortly papillose. Corona scales from the tip to the outer end about 0.2 cm. long. Northeast New Guinea: on trees in the forest in the vicinity of Eitape, about 20 meters in altitude (Schlechter #19964. -- Blooming in August 1909).

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In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. eitapensis Schlechter (Schlechter 19964) – 15. In Dr. Schlechter’s Hoya Species (1993) 56. R. D. Kloppenburg. Hoya eitapensis Schlechter. A twining perennial branched plant, woody at the base, epiphytic. Branches round, thin, flexible and glabrous, loosely leaved. Leaves erect, spreading, elliptic, rather obtuse, with wedge shaped base, fleshy, glabrous with very short petioles, almost sessile. Inflorescences shaped like an umbel, with the peduncle longer than the leaves, 20-35 flowered, with the pedicels rather slender, unequal in length. Flowers small for the section. Segments of the calyx ovate triangular, somewhat obtuse, very minutely and very sparingly puberulous and ciliate, much smaller than the of the corolla. Corolla rotate, apex reflexed, 5 parted up to 1/3 from the base, lobes ovate acute, outside glabrous, inside very minutely pimply and puberulent. Scales of the corona horizontal, oblong, interior slightly elevated, acuminate, exterior obtuse, sides rounded and keeled, anthers barely longer. Pollinia obliquely oblong, apex somewhat curved, with the translators much shorter, retinaculum rhomboid, minute. A pretty clinging vine with threadlike loosely leaved branches. Leaves 3-5 cm. long, in the middle 1.2-3 cm. wide. Petiole hardly 2 mm. overall. Inflorescences like an umbel on about 5 cm. long peduncles. Pedicels very slender, up to 2 cm. long, toward the apex they are very shortly and sparsely pubescent. Calyx lobes about 1.75 mm. long very shortly and sparsely pubescent. Corolla 0.8 cm. in diameter, on the inside very shortly papillose. Corona scales from the tip to the outer end about 0.2 cm. long. Northeastern New Guinea: on trees in the forest in the vicinity of Eitape, about 20 meters in altitude (Schlechter #19964. -- Blooming in August 1909). This and the following two species I have placed at the beginning of the section, since their habitat and small flower size, are closest to the section Otostemma. This one differs from H. microstemma Schlechter by the larger leaves and with shorter corolla pubescence as well as by the narrower corona scales. Its blooms are yellowish-white. Editors Note: This species does not belong in the Otostemma Section. Published by Dr. R. Schlechter in: 1913 Botanische Jahrbücher v.50 p.109, "Die Asclepiadaceen von Deutsch Neu Guinea". In Hoya Section Acanthostemma (Blume) Kloppenburg (1994) 38-42. R. D. Kloppenburg. Hoya eitapensis Schlechter n. sp. - Suffrutex epiphyticus, ramosus scandens. Rami filiformes, flexuosi, laxe foliati, teretes, glabri. Folia erecto-patentia elliptica, obtusiuscula, basi cuneata, carnosa, glabra, brevissime petiolata vel subsessilia. Inflorescentiae umbelliformes pedunculatae, folia superantes, 20- 35 florae, pedicellis gracilibus, inaequilongis. Flores in sectione inter minores. Calycis segmenta ovatatriangula obtusiuscula, minutissime et sparsissime puberula et ciliolata, corolla multi breviora. Corolla rotata apicibus reflexis, usque ad tertiam partem basilarem 5-fida, lobis ovatis, extus glabris, intus minutissime papilloso-puberulis. Coronae foliola horizontalia, oblonga, antice leviter adscendentia, acuminata, extus obtusa, lateribus rotundato carinata, antheris vix longiora. Pollinia oblique oblongoideo, apice subfalcata, translatoribus quam maxime abbreviatis, retinaculo rhomboideo, minuto.

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Ein zierlicher Schlinger mit fadenförmigen, locker beblätterten Zweigen. Blätter 3-5 c, lang, in der Mitte 1.2-2 cm. breit, Blattstiele kaum 2 mm überschreitend. Blütenstande doldig auf etwa 5 cm langen Stielen. Blütenstiele sehr schlank, bis 2 cm lang, nach der Spitze zu sehr kurz und spärlich behaart. Kelchzipfel ca. 1.75 mm. lang, sehr kurz und spärlich behaart. Korolla 0.8 cm im Durchmesser, innen sehr kurz papillös. Koronaschuppen von der Spitze bis zum äuszern Rande etwa 0.2 cm lang. Nordöstl. Neu-Guinea: auf Baümen in der Wäldern in der Umgebung von Eitape, ca. 20 m u. M. (Schlechter n. 19964. - Blühend im August 1909). Diese und die beiden folgenden Arten habe ich an den Anfang der Sektion gestellt, da sie sowohl im Habitus wie in der geringen Grösze ihrer Blüten den Arten der Sektion Otostemma am nächsten stehen. Die vorliegende unterscheidet sich von H. microstemma Schltr. durch die gröszeren Blätter und die kürzer behaarte Korolla sowie durch schmälere Koronaschuppen. Ihre Blüten sind gelblich-weisz. Translation of the Latin and German: Hoya eitapensis Schlechter in Engler's Botanische Jahrbücher 50 (1913) 109. "Die Asclepiadaceen von Deutsche Neu-Guinea". Hoya eitapensis Schlechter n. sp. - A twining perennial branched plant, woody at the base, epiphytic. Branches round, thin, flexible and glabrous, loosely leaved. Leaves erect, spreading, elliptic, rather obtuse, with wedge shaped base, fleshy, glabrous with very short petioles, almost sessile. Inflorescence shaped like an umbel, with the peduncle overtopping the leaves, 20-35 flowered, with the pedicels rather slender, unequal in length. Flowers small for the section. Segments of the calyx ovate triangular, somewhat obtuse, very minutely and very sparingly puberulous and ciliate, much smaller than those of the corolla. Corolla rotate, apices reflexed, 5 parted up to 1/3 from the base, lobes ovate acute, outside glabrous, inside very minutely pimply and puberulent. Leaflets (scales) of the corona horizontal, oblong, interior slightly elevated, acuminate, exterior obtuse, sides rounded and keeled, anthers barely longer. Pollinia obliquely oblong, apex somewhat curved, with the translators much shorter, retinaculum rhomboid, minute. A pretty clinging vine with threadlike loosely leaved branches. Leaves 3-5 cm. long, in the middle 1.2-3 cm. wide. Petiole hardly 2 mm. overall. Inflorescences like an umbel on about 5 cm. long peduncles. Pedicels very slender, up to 2 cm. long, toward the apex they are very shortly and sparsely pubescent. Calyx lobes about 1.75 mm. long very shortly and sparsely pubescent. Corolla 0.8 cm. in diameter, on the inside very shortly papillose. Corona scales from the tip to the outer end about 0.2 cm. long. Northeastern New Guinea: on trees in the forest in the vicinity of Eitape, about 20 meters in altitude (Schlechter #19964. -- Blooming in August 1909). Housed at (B). This and the following two species I have placed at the beginning of the section, since their habit and small flower size are closest to the section Otostemma. This one differs from H. microstemma Schlechter by the larger leaves and with shorter corolla pubescence as well as by the narrower corona scales. Its blooms are yellowish-white. References: Kloppenburg in Hoyas of Northeast New Guinea (1992) 10-20. A. Nicholas in The Asclepiadaceous Works of Schlechter (1992) 30. Herbarium Sheet: Hartley #12633 1964 (BRI).

Hoya elegans Kosteletsky

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In Rumphius’ Herbarium Amboinensis (1917) 439. “Interpretations of Rumphius’ Herb. Amb.” E. D. Merrill. Hoya elegans Kostel. Allgem. Med. Pharm. Fl. 3 (1834) 1084 (type!) Nummularia lactea major III (albo-purpurea) Rmph. Herb. Amb. 5:471. Amboina, Soja and Hitoc messen, Robinson Pl. Rumph. Amb. 83, 604. August and October, 1913, in light forests, altitude 175 to 300 meters, locally known as hunga nasi. Hasskarl, Neu Schlüssel (1866) 148, thought that this Rumphian form might be a synonym of Hoya macrophylla Blume, a species originally described from Javan material. Hoya elegans Kostel. Was based wholly on the Rumphian description, and this name is here retained.

Hoya elliptica Hooker In The Flora of British India 4 (1883) 58. J. D. Hooker. 27. H. elliptica, Hook. f.; glabrous, leaves 1 ½ - 2 ½ in. elliptic obtuse et both ends thin in texture nerves very prominent, peduncle very short, pedicels long slender, sepals very small, corolla glabrous without puberulous within. Malacca, Maingay (Kew Distrib. 1137). Stem rather slender, twining. Leaves 1- 1 ½ in. intramarginal nerve very distinct; petiole 1/6- ¼ in. Peduncles ½ in., pedicels 1-1 ½ in. Sepals ovate, obtuse puberulous. “Corolla white.” Maingay; lobes obcordate, apiculate, inflexed (when dry). Coronal-processes longer than the corolla-tube, suberect, laterally subcompressed, ovate-oblong, upper (inner) surface grooved, inner angle produced into a short subulate erect point. In The Journal of the Royal Asiatic Society, Bengal Branch 74 (1908). King & Gamble. 6. Hoya elliptica, Hook. f. in Fl. Br. Ind. IV. 58 (1833). A lender twining undershrub; branchlets pale, terete, glabrous. leaves fleshy (?) when fresh, chartaceous: when dry; elliptic, obtuse both at apex and base; glabrous on both surfaces and pale, especially beneath; margin hardened, nerve-like, slightly recurved ; .15 to 2.5 in. long, 1 to -1.5 in. broad; nerves prominent when dry; midrib slender, main nerves nearly at right angles to the midrib, irregular, 8 to 10 pairs, straight to join a prominent intramarginal looped nerve at 1 to 15 in. from the edge, the said intramarginal nerve connected by many cross-nerves with the hardened marginal nerve; reticulations netted; petiole .15 to .25 in. in. long, flattened. Umbels racemose, about 8- to 10-flowered, on a short tubercular rachis at the end of a thickened pink peduncle reaching 1 in. in length; pedicels slender, 1 to 1.5 in. long, puberulous; buds 25 in. long, 5-angled, the incurved-valvate corolla-lobes forming a central con-cavity; flowers “white" (Maingay), nearly black when dry. Calyx puberulous, pale pink; tube very short; lobes ovate-oblong, obtuse, .05 in. long; scales minute. Corolla 5 in. broad when open, glabrous without, sparingly pilose within; lobes obcordate, apiculate, inflexed (when dry). Corona of 5 processes; the lower and large lobes suberect, laterally compressed, bladder-like (the interior of the cavity delicately hairy), ovate, slightly apiculate; the upper short, flattened, slightly recurved, oppressed to and covering the anther. Staminal-column attached to the base of the corolla-tube; anthers short, much inflexed so as to lie nearly horizontal; cells parallel above, divergent and except below; appendages scarious, acute; pollen-masses oblong-falcate, pellucid on the outer margin, nearly horizontal, attached by thickened caudicles to the rather large elongate pollen-carriers which are obtuse above and 2 winged below. Style-apex

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5-angled, fattened, with; a short central umbo having 5 divergent rays to the angles. Fruit not known. Perak: Scortechini. Malacca; Maingay (KD.) 1137. Lower lobes beaked; upper lobes short, acute, stems slender; leaves thin, under 4 in. long. .........................................................................6. H. elliptica. In Flora of the Malaya Peninsula 2 (1901) 394. H Ridley. (18) H. elliptica Hook. fil. F.B.I. iv. 58; King, l.c. 565. Slender twiner. Leaves chartaceous when dry, elliptic blunt both ends; nerves 8 to 10 pairs, horizontal joining and intramarginal nerve; 1.5 to 2.5 in. long, 1 to 1.5 in. wide; petioles .15 to 1.25 in. long. Peduncles pink, thick, 1 in. long, rachis short; pedicels slender, 1 to 1.5 in. long. Calyx puberulous. pink, lobes ovateoblong, blunt. Corolla .5 on. across, pilose inside white,, lobes obcordate apiculate. Corona, lower lobes sub-erect, bladder-like ovate; upper lobes short, covering anthers. Hab. Malacca (Maingay). Perak (Scortechini). A rare and very little known plant. Flowers small; corona lobes beaked; leaves elliptic with horizontal nerves meeting and intramarginal. ......................................(18) H. elliptica In Malayan Nature Journal 30 (1978) 505-508. Rintz. 70 Hoya elliptica Hooker, F.B.I. IV (1883) 58. Type: Malaysia, Melaka, Maingay 3286 (K). – Fig. 21. Distinguishing Features: Leaves fleshy, elliptic and very flat with the veins at nearly right angles to the midrib; up to 10cm long by 1.4cm wide. Peduncle horizontal, rigid, 4 — 5cm long; pink. Umbel negative-geotropic, convex with rigid, uniform, pink-colored pedicels 3—4cm long; 1—30 flowers, open 4 days and strongly scented at night. Corolla spreading, pubescent inside; 1.5—2 cm diam.; white. Corona lower lobe inflated and thin horizontally with a narrow ridge at the apex and a large gland inside; entirely white. Pollinia with angled wings and a very large corpuscle. Follicle 15—20cm long by 5mm diam.; often pale pink in color. Ecology: Endemic to Malaysia: common along rivers in Selangor and also known from Melaka, Perak and Kranji, Singapore. The roots are often associated with ant nests on the surface of trees. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 87 “The Genus Hoya in Thailand” O. Thaithong. 5. H. elliptica Hook. f., FBI 4 (1883) 5*, FMP 2 (1923) 399; Malay. Nat. J. 30 (1978), 505, 508, Fig. 21. Occurrence: (PEN): Narathiwat. Note: page citation incorrect should be 58-59.

Hoya elmeri Merrill In University of California Publications in Botany 15 (1929) 258. E. D. Merrill. Hoya elmeri Sp. nov. § Eu-hoya. Scandens, floribus exceptis glabra, caulibus teretibus; 5 ad 6 mm diametro, ramis paucis, levibus, circiter 3mm diametro, saepe radicantibus, internodiis usque ad 12 cm longis: foliis oppositis, carnosis. in siccitate coriaceis. penninerviis, subellipticis, 11 ad 15 cm longis, 5 ad 6.5 cm latis. in siccitate pallidis, utrinque concoloribus. vix verruculosis, utrinque subaequaliter angustatis, basi acutis, apice acutis ad breviter obscureque acuminatis, nervis primariis utrinque circiter 7, tenuibus, distantibus. distinctis vel interdum subindistinctis, arcuato-anastomosantibus,

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reticulis laxis; petiolo crasso, circiter 1 cm longo; umbellis pendulis, pedunculo saltem 3.5 cm longo, circiter 1.5 mm crasso, glabro, pedicellis circiter 1.5 cm longis; floribus rubidis, 10 ad 12 mm diametro, rotatis, sepalis late ovatis, rotundatis, tenuibus, glabris, circiter 1.3 mm longis; corolla extus glabra intus villosa, lobis patulis vel recurvatis, latissimis, circiter 3mm longis et 5 mm latis, nee profunde 3-lobatis, lobis lateralibus late rotundatis, aliis acutis; corona circiter 6.5 mm diametro, processibus corneis, patulis vel leviter adscendentibus, acutis vel subacutis, compressis, circiter 3.5 mm longis, margine superiore subcarinato, haud plano. Elmer 20652, near Tawao. Scandent in dense thickets among large boulders in densely forested ravines, the flowers very dark red, easily falling. I have not been able to refer this to any of the described species of this rather difficult genus. It is characterized by its rather small, dark red flowers, and its distinctly pinnately nerved, subelliptic leaves which are subequally narrowed to both the acute base and apex. Translation: Climbing, glabrous except for the flowers, stalk round, 5 to 6 mm. in diameter, branches few, smooth, about 3 mm in diameter, often rooting, internodes up to 12 cm. long; leaves opposite, fleshy, when dry leathery, penninerved, somewhat elliptic, 11 to 15 cm long, 5 to 6.5 cm wide, when dry pale, both sides the same color, sparsely covered with small wort-like growths, both ends equally narrow , base acute, apex acute to briefly and obscurely acuminate, primary nerves on both sides of the costa 7, slender, distinct or sometimes somewhat indistinct, curved anastomosing, loosely reticulated ; petiole thick, about 1 cm long; fleshy, glabrous 1.3 mm. long’ corolla outside glabrous inside silky, lobes spreading or recurved, very broad, about 3 mm. long and 5 mm. wide, not deeply 3 lobed, sides of lobes broadly rounded, others acute; corona about 6.5 mm in diameter, processes horny, spreading or slightly ascending, acute or somewhat acute, compressed, about 3.5 mm long, margines above somewhat keeled, not all flat. In Sunyatsenia 3. 2/3 (1934) 175. Tsiang. Hoya Elmeri Merrill in Univ. Calif. Publ. Bot. 15: 258. (Pl. Elmer. Born.) 1929. Borneo: Near Tawao, A. D. E. Elmer 20652, Oct., 1922 - March, 1923. Endemic. Of this, we have a sterile twig and there detached flowers which enable me to place it here. The most interesting thing in this species is that the coronallobes are very long, surpassing the sinus of the corolla-lobes. Otherwise, it closely resembles H. carnosa.

Hoya elmeri Schlechter In Dr. Schlechter’s Hoya Species. (1993) 58. R. D. Kloppenburg. Hoya elmeri Schlechter (unpublished). This species is not the same species as Hoya elmeri Merrill, which was collected near Sandakan in North Borneo. This species was never published so there is no Latin description. The sheet is in the Berlin Herbarium, Germany.

Hoya el-nidicus Kloppenburg In Fraterna 3rd Quarter (1991) I. R. D. Kloppenburg. Hoya el-nidicus Kloppenburg n. sp. Type #41931 El Nido Rest Area, Palawan (northern) Philippines, vine on limestone ledge, collected by Professor Juan Pancho 15 August 1988. Holotype (CAHUP).

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Suffrutex parum ramosus, alte scandens. Rami filiformes, flexuous, laxe foliati, tereste, glabri. Folia patentia vel patula, petiolata, ovata vel obovata, acuminata (apiculata, basi breviter cordata, textura coriacea, utrinque glaberrima. Inflorescentiae umbelliformes c. 6-12 florae, pedunculatae, pedunculo pedicellisque glabris. Calycis segmenta ovalia obtusissima, margine breviter ciliata, quam corolla multo minora. Corolla rotata, usque infra medium 5-fida, extus glabra punctata, lobis ovatis, subacutis vel obtusis, intus dense et minute papillosa. Coronae foliata patentia, subhorizontalia, superne sub convexa, ovata, antice apiculata extus obtusa vel brevi apiculata lateribus incrassata-rotundata. Pollinia oblique, oblongoidae, translatoribus brevissimis, retinaculo minutissimo rhomboideo. The accompanying drawing is by R. D. Madena of the University of the Philippines of Los Banos, Laguna, Luzon. It was drawn from live material in 1988. An epiphytic ornamental vine, with cordlike, loosely leaved branches, with long internodes c. 15 cm. long. Leaves outspread, 8.5-10.3 cm. long, at the widest above the middle 5.1-6.8 cm wide, with the petiole round, glabrous, 1.7 cm. long, curved. Inflorescences pendulous, umbel like, thick 3-4 cm. long, bare; pedicels c. 1.1 cm. long greenish white to reddish-pink with age, bare. Calyx lobes small, with ciliate margins, 2.7 mm. long 2.5 mm. wide below the middle, papillose outside, glabrous, membranaceous, no ligules seen. Corolla rotate, white, the margins reflexed, patently white, villose along the edges, c. 1 cm. in diameter. Corona scales 5 spreading, pinkish, 3.5 mm. long, apex extending somewhat upward, anthers greatly exceeding the apex outer end abruptly blunt, turned down, top sides broadly rounded, linearly lined, center above raised, under side shortly channeled. Ovaries stubby. Among the rotate Philippine species with pubescent inner corona surfaces and ciliate calyx lobes, the flowers are delineated from H. pubicalyx Merrill by being smaller and with the corona scales much broader and shorter. Also the calyx surface on this species is glabrous whereas H. pubicalyx Merrill is pubescent. as is that with H. halconensis Kloppenburg. The foliage is larger and different shape than with H. cagayanensis Burton and with a much shorter peduncle, like this species the calyx lobes are glabrous with ciliate margins but this new species has broadly round lobes, not somewhat linear as in H. cagayanensis Burton, the flower size is larger in this species. It is distinctive and unique among Philippine Hoya species. This species is named after this international island resort area, El Nido, where it was discovered. A very rugged rocky area, and very isolated, into which Dr. E. Merrill did not send his collectors. It is an extremely difficult area in which to collect. Other Measurements: Sinus to Sinus 4.0 mm. Sinus to Collar 3.3 mm. Sinus to Center 3.6 mm. Sinus to apex 4.0 mm. Retinaculum 0.3 mm. tall, 0.1 mm. wide. Pollinia 0.7 mm. long, 0.3 mm. wide at apex. Translation: Perennial a little branched, high climbing. Stems threadlike, flexible loosely foliaged, round, glabrous. Leaves spreading or outspread, petioled, ovate or obovate, acuminate, apiculate, base somewhat cordate, texture leathery, both sides glabrous. Inflorescence like an umbel of about 6 to 12 flowers, pedunculate, peduncles and pedicels

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glabrous. Calyx segments ovate, very obtuse, margins shortly ciliate, much smaller then the corolla. Corolla rotate 5 lobed all the way to the middle, outside glabrous punctate, lobes ovate, somewhat acute or obtuse, inside densely and minutely papillose. Leaflets of the corona spreading, somewhat horizontal, above somewhat convex, ovate, outside apiculate, inside obtuse or shortly aciculate, laterally thickened rounded. Pollinia oblique, somewhat oblong, translators short, retinaculum minute rhomboid. In Philippine Hoya Species 3rd Edition (1996) 69-70. R. D. Kloppenburg. Hoya elnidicus Kloppenburg in Fraterna 3rd Quarter 1991 p.l. Type #41931 El Nido Rest Area, Palawan (northern) Philippines, vine on limestone ledge, collected by Professor Juan Pancho 15 August 1988. Holotype (CAHUP). The accompanying drawing is by R. D. Madena of the University of the Philippines of Los Banos, Laguna, Luzon. It was drawn from live material in 1988. An epiphytic ornamental vine, with cord like, loosely leaved branches, with long internodes c. 15 cm. long. Leaves outspread, 8.5-10.3 cm. long, at the widest above the middle 5.1-6.8 cm wide, with the petiole round, glabrous, 1.7 cm. long, curved. Inflorescences pendulous, umbel like, thick 3-4 cm. long, bare; pedicels c. 1.1 cm. long greenish white to reddishpink with age, bare. Calyx lobes small, with ciliate margins, 2.7 mm. long 2.5 mm. wide below the middle, papillose outside, glabrous, membranaceous, no ligules seen. Corolla rotate, white, the margins reflexed, patently white, villose along the edges, c. 1 cm. in diameter. Corona scales 5 spreading, pinkish, 3.5 mm. long, apex extending somewhat upward, anthers greatly exceeding the apex, outer end abruptly blunt, turned down, top sides broadly rounded, linearly lined, center above raised, under side shortly channeled. Ovaries stubby. Among the rotate Philippine species with pubescent inner corona surfaces and ciliate calyx lobes, the flowers are delineated from H. pubicalyx Merrill by being smaller and with the corona scales much broader and shorter. Also the calyx surface on this species is glabrous whereas H. pubicalyx Merrill is pubescent. as is that with H. halconensis Kloppenburg. The foliage is larger and different shape than with H. cagayanensis Burton and with a much shorter peduncle, like this species the calyx lobes are glabrous with ciliate margins but this new species has broadly round lobes, not somewhat linear as In H. cagayanensis Burton, the flower size is larger in this species. It is distinctive and unique among Philippine Hoya species. This species is named after this international island resort area, El Nido, where it was discovered. A very rugged rocky area, and very isolated, into which Dr. E. Merrill did not send his collectors. It is an extremely difficult area in which to collect. Other Measurements: Sinus to Sinus 4.0 mm. Sinus to Collar 3.3 mm. Sinus to Center 3.6 mm. Sinus to apex 4.0 mm. Retinaculum 0.3 mm. tall, 0.1 mm. wide. Pollinia 0.7 mm. long, 0.3 mm. wide at apex.

Hoya endauensis Kiew

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In The Malayan Nature Journal 42 (1989) 262-265. R. Kiew. Hoya enduensis sp. nov. Hoya endauensis is presently known only from tributaries of the Ulu Endau, where it grows on trees on river banks. This new species adds to the growing list of endemics recorded from the Ulu Endau (Kiew et al., 1987), which is in the process of being gazetted as a state park. Foliis glabris ovatis-cordatis apicibus acutis 2 ad 2.5 cm longis et 2 ad 2.5 cm latis a congeneribus Peninsulae Malaysiae divisa. Typus: R. Kiew RK2746 (UPM) Sg, Jasin, Jahore. Glabrous epiphyte climbing with a thin stem (1.5 mm thick in dried state (with internodes 6-9 cm long. Stem with climbing roots at the nodes and a more expansive root system developing below the leaves, Leaf ± sessile. Lamina ovate-cordate, humped with margines appressed to bark of supporting tree, 2.2 cm -2.5 by 2.5 cm, apex acute, in life succulent (not tough), plane above and venation obscure in life, 3 pairs of veins just visible in dried state, in life pale yellow-green above and below, upper surface sometimes spotted orange (by algae), margin not thickened. Inflorescence a pendant, axillary, sparsely-flowered corymb. Peduncle 23-45 mm long and 1 mm thick in dried state, distal part (2-7 mm long) covered by fruit scars, Pedicels very slender, 5-8 mm long. Flower buds pendant, 4 mm across, pale green suffused purple-red towards the base. Calyx pale green, glabrous, lobes oblong, 1 by 1 mm, apex rounded, pustulate on outer surface. Corolla glabrous on inner and outer surfaces, 4 mm across; lobes triangular, 2.5 by 2.5 mm. apex acute. Corona 3 mm across apex of upper lobes upturned and acute, lower lobes sloping down from and projected beyond upper lobes. Caudicles and pollinia broadly winged, 1 mm long. Ovary ovoid, 1 mm long. Fruit unknown. Ecology: It is an epiphytic climber growing in partial shade on trees on river banks at c. 75 m a.s.l. Distribution: Endemic to Ulu Endau, Jahor, Malaysia. Specimens examined: Jahore, Ulu Endau: Sg. Salat- 3 July 1985 R. Kiew s.n. (UMP). 2 June 19898 R. Kiew RK2746 (UPM). In vegetative characters it most resembles H. curtisii K & G., which is also a slender climber, but which has small (13 by 13 mm), orbicular leaves, which are apiculate. It also differs from H. curtisii in indumentum (H. curtisii is hirsute) and inflorescence characters (Inflorescence of H. curtisii have peduncles to 5 mm longer are sessile); Kiew, 1987. In reproductive characters H. endauensis is most similar to H. lacunosa Bl. in its pendant, stalked inflorescence, in the shape of the corona, which has downward sloping lower lobes, and in the broadly winged caudicles (illustrated by Rintz, 1978). However H. endauensis is different in having glabrous corolla; that of H. lacunosa has inner surfaces covered by long, thick hairs. The leaves of H. lacunosa are also different. Usually they are longer than broad (2.5 - 3.5 by 1.5 - 2.5 cm) and have a thickened margine. In addition H. endauensis has smaller flowers (the corolla is 4 mm in diameter) than either H. curtisii (7mm) or H. lacunosa (8mm). Translation: Leaves glabrous ovate-cordate apiculate acute 2 to 2.5 cm, long and 2 to 2.5 cm, wide , diverse from peninsula Malaysia cogenerics.

Hoya engleriana Hosseus In Notizbaltt des Konigl. Botanischen Gartens und Museums zu Berlin 40 (1907) 315. Hosseus. Hoya Engleriana Hoss. n. sp. (Sect. Euhoya Miq.); epiphytica in arboribus, ramosa, ramis ramulisque elongatis, gracilibus, filiformibus, partim dependentibus,

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pubescentibus laxe vel densiuscule foliatis; foliis breviter vel brevissime petiolatis, breviter lanceolatis vel anguste ovato-lanceolatis, marginibus involutis, itaque primo aspectu speciem foliorum lineari-lanceolatorum imprimis versus inflorescentiam praebentibus, carnosulis, albido-pubescentibus, nervatione supra inconspicua, 1.5 cm longis, 0.4 cm latis, geniculis mobilibus; umbella terminali, vix vel breviter pedunculata, 4-flora, floribus suaveolentibus, pedicellatis, pedicello circ. 0.5 cm longo piloso; sepalis margine minute ciliatis, pilosis, carneis, 2 mm longis; corolla rotata, 1.6 cm diam., fere usque medium 5-lobata, carnosa, albida, extus glabra, intus dense brevissime granuloso-puberula lobis late ovato-deltoideis, acutis vel acuminulatis, fere horizontaliter patentibus, serius margine paullulo recurvis; gynostegio violaceo, marginibus obscurioribus. Siam: auf dem Doi Sutäp bei Djieng Mai in Ober-Siam (Lao-Provinz), um 1580 m. u. M. unterhalb des Gipfels auf Bäumen des immergrünen Urwaldes. Nah verwandt mit Hoya linearis Wall.: doch unterscheidt sie sich im Habitus, sowie in den bedeutend kleineren, anders beschaffenen Blättern (cf. Taf. II, Fig. A, H-N). Auszerdem besitzt Hoya Engleriana Hoss. Blätter mit gut ausgebildeten Gelenkolstern. Hoya Engleriana Hoss. wurde von mir auf dem Doi Sutäp lebend gesammelt. Die Pflanze wächst in der Nähe des Gipfels auf den Bäumen des immergrüen Urwaldes, so z. B. auf Anneslea fragans Wall. (einer Theacee), die zugleich auch eine ebenfalls im botanischen Garten in Dahlem in Kultur befindliche Pflanze, Agapetes Hosseana Diels beherbergt. Deshalb empfiehlt sich die Kultur beider in nicht zu heiszen Häusern. Es ist sin glücklicher Zufall, dasz die beiden nah verwandten Arten sich zu gleicher Zeit lebend im botanischen Garten in Berlin befinden. Hierbei fallen natürlich die Unterschiede vor allem im Habitus um so schärfer auf. Hoya linearis hat lange, eng einander anliegende herabhängende Zweige mit langen Blättern, die — den Zweigen anliegend — herabhängen. Nur im Jugendstadium machen die Einzelblätter Wachstumsbewegungen. Ein anagesprochenes Blattpolster ist nicht vorhanden. Das in Dahlem befinliche Exemplar, das bereits eine Anzahl Sämlinge ergeben hat, stammt aus Sikkim und ist von Professor Dr. Engler gesammelt. Im Gegensats nun zu Hoya linearis nimmt Hoya Engleriana einen viel gröszeren Raum als Pflanze fur sich in Anspruch, die einzelnen, verzweigten Zweige, die vor allem in älteren Stadien herabhängen, stehen zumeist weit ab. Die bedeutend kürzercn Blätter dagegen verländern unter günstigen äuszeren Faktoren ihre Lage; ausgewachsen reagieren sie bei autonomen Variationsbewegungeu, besonders güsstig auf Wärme und Lichteinflusz. Hoya Engleriana wird sicher nach Dssmodium gyrans, Flemmingia congesta und Trifolium pratense, alles Leguminosen, eins äuszerst wichtige Pflanze fur die Bewegungsfragen Angedeutet ist die verschisdene Lage der Blätter auf Tafel II, A. Die Pflanze wurde von mir lebend nach Europa gebracht und befindet sich z. Z. (Juli) in botanischen Garten in voller Blüte. Hoffentlich gelingt es Hoya Engleriana durch Stecklinge zu vermehren, damit die physiologischen Fragen gründlich untersucht werden können. Translation: epiphytic in the trees, branching, stems rooting, elongate, glabrous, threadlike, partially hanging down loosely pubescent, or densely foliaged; leaves short or shortly petiolate, shortly lanceolate or narrowly ovate-lanceolate, margines curled inward leaves of the species accordingly at first appearing chiefly turned in the direction of the inflorescence, fleshy, white pubescent, nerves above inconspicuously marked, 1.5 cm

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long, 0.4 cm wide, narrow mobile, umbels terminal, barely or shortly peduncled, 4 flowered, flowers fragrant, pedicelled, peduncles about 0.5 cm long hairy, sepals with minutely ciliate margins, hairy, fleshy 2 mm. long. Corolla rotate, 1.6 cm in diameter nearly 5 lobed all the way to the middle, fleshy, white, outside glabrous, inside densely and shortly granulose puberulus, lobes broadly ovate-deltoid, acute or acuminate, nearly horizontal spreading later the margines a little recurved, gynostegium violet, margines somewhat obscure. Siam: on the Doi Sutäp near Djieng Mai in upper-Siam (Lao-Province), about 1580 meters above sea level, below the summit on trees of the evergreen jungle. Near Hoya linearis allied with it: however distinguishing itself in the Habitat, just as in the significantly smaller, differently constituted leaves (cf. Taf. II, Fig. At, H-N). Auszerdem possesses Hoya Engleriana Hoss. Leaves with well trained supple padding. Hoya Engleriana Hoss. was collected freely by me on the Doi Sutäp. The Plant grows near the of the summit of the trees of the evergreen of jungle, so for example on Anneslea fragans Wallich. (a Theaceae), that at the same time the plant likewise in culture situated in the botanical garden in at Dahlem, also Agapetes Hosseana Diels lodges there. Therefore both of them are not recommended for culture in a small house. It is fortunate coincidence, that the two near relative types themselves at the same time are living in the botanical garden in Berlin. On this occasion the differences attract attention nature-like in the Habitus all the more keenly above all else. Hoya linearis has for a long time, allied narrowly to each other, hanging herbs branches with long leaves,- the branches allied—hanging herbs. Only in the youth-stage are there many single-leaves growth-movements. One concludes the leaf-cushions does not exist. That in Dahlem mounted example, that one number has already yielded a seedling, comes from Sikkim and is collected by professor Dr. Engler. In the opposition now to Hoya linearis takes Hoya Engleriana the plant claims for itself a much larger area for example, the single, branched branches, that above all else in different growth stages a hanging herb, stands out the most. The situation of their significantly short leaves against the changeable under favorable other factors; developed to counteract them with autonomous variation inducement, particularly favorable on warm and light rain. Hoya Engleriana certainly becomes after Dssmodium gyrans, Flemmingia congesta and Trifolium pratense, all Legumes, one other important plant for the discussion. Is the situation of the leaf of the drawing on layout II, A., The plant was brought back by me alive to Europe and I found it Z. Z. (July) in the botanical garden in full bloom. Hopefully Hoya Engleriana will succeeds through cuttings to multiply, so that the physiological questions are examined become understood. In Kew Bulletin (1911) 418. “Contributions to the Flora of Siam” W. G. Craib. Hoya Engleriana, Hosseus, Notizbl. Bot. Gart. Berl., No. 40, p. 315. Described from a specimen in cultivation at Dahlem collected on Doi Sootep, 1580 m. by Hosseus. In Aberdeen University Studies 135. “Flora of Siam” W. G. Craib. Hoya Engleriana, Hosseus, Notizbl. Bot, Gart. Berl., No. 40, p. 315; Hosseus, ii p. 427. Described from a specimen in cultivation at Dahlem collected on Doi Sootep, 1580 m. by Hosseus.

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In Flore Generale de Indo Chine 4 (1912) 132. J. Constantin. 7. H. Engleriana Hosseus, in Notizblatt Bot. Gart. Berlin, no. 40, p. 315. Plants epiphyte, rameuse. Tiges et rameaux de divers orders greles, allonges, filiformes, pubescents, couvertes de feuilles plus ou moins espalees. Feuilles petites, brievement petiolees, lanceolees ou ovales-lanceolees, a bords fortement enroules en dessous, sur le sec charnues, pubescentes, blanches; nervure mediene seule saillante en dessous; limbe long de 1.5 cm. sur 4 mm., articule et mobile a la base; petiole long de 2 mm. Inflorescence en ombelle terminale, brevement pedunculee a fleurs; peduncule poilu, long de 5 mm.; pedicellis de 5 mm.; fleurs odorantes. — Calice: sepales a bords tres cilies, etroits, longs de 2 mm.; glande entre les sepales. Corolle rotacee de 15 mm. de diam. a l’epanouissement, charnue, blanche, glabre en dehors, brievement puberulente et granduleuse en dedans; lobes 5, arrivant presque jusqu'au milieu, ovales-deltoides, aigus ou acumines, etales, presque horizontaux, a bords un peu incurves. Coronule a 5 pieces, petites, en étoile, charnues, a bec developpe vers le haut. Gynosteme violace a bords plus forces. Siam: Doi-sootep, pres Xieng-may (Hosseus). Feuilles courtes, petites, de moins de 1.5 cm. de long ................7. H. Engleriana. Translation: Epiphytic plant, Stems and branches small and of a different character, elongated filiform, pubescent, covered with leaves more or less wide spaced. Leaves very small, briefly petiolate, lanceolate or ovate-lanceolate, with edges strongly recurved, when dry, fleshy pubescent, white; midrib only prominent below; leaves 1.5 cm. by 4 mm. long, joints articulate and movable at the base; petiole 2 mm. long. Inflorescence in terminal umbels, briefly peduncled, having 4 flowers; peduncle hairy, shaggy 5 mm. long; pedicels to 5 mm.; flowers sweetly scented. Calyx sepals with edges very ciliate, narrow to 2 mm. long; gland between the sepals; corolla rotate of 15 mm. in diameter, when fully opened; fleshy white, glabrous briefly pubescent on the outside and glandular on the inside, 5-lobed nearly to the center, lobes ovate triangular, acute or acuminate, spread out almost horizontally with edges a little incurved. Corona a 5 scaled petite star, fleshy, with a beak at the top. Gynostegium purplish, at the edges darker. Siam: DoiSootep, near Cheng-Mai (Hosseus). Leaves short petite less than 1.5 cm. long. In A Handbook of Succulent Plants (1954). H. Jacobsen. Hoya Engleriana, Hoss, Siam. Epiphytic plant. L. 1.5 cm long, 4 mm wide. Upper surface convex. Flowers 4 together, terminal, 15 mm in diameter, white-violet. Prefers a light position in a warm house. In Dictionary of Gardening the Royal Horticultural Society (1965) 1016. H. Engleriana. Branches partly pendent. l. lanceolate or narrowly ovate-lanceolate, stalk short. fl. whitish ¼ in. across, fragrant; cor. fleshy deeply 5-lobed, usually in 4-fid. terminal umbels. Siam. Somewhat resembles H. linearis but smaller l. and different habit. In New York Botanical Garden Illustrated Encyclopedia of Horticulture 5 (1981) 1726. H. engleriana, of Thailand, has very short-stalked, linear-lanceolate to lanceolate or ovate-lanceolate leaves ½ inch to 1 ½ inches long. The flowers, a little over ½ inch across and white to pinkish with a pink corona, are mostly clusters of four. In The Complete Handbook of Cacti and Succulents. C. Innes. Hoya engleriana Hoss. A pendent species with thin stems, climbing or creeping. Leaves very small, elliptic,

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upper surfaces convex, dark green. Flowers in umbels, usually of 4 together, white with purplish center. From Thailand. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 87. “The Genus Hoya in Thailand” O. Thaithong. 6. H. engleriana Hosseus, Notizbl. Bot. Gart. Berl. 40, (1907) 315, t 2. , FIC 4 (1912) 132: Fl. Siam 3(1), (1951) 36. Occurrence: (N): Chiang Mai, Mae Hongson. Notes: Stem short; branches pendent, flower about 2 cm dia. when fully expanded; corolla rotate-shallowly campanulate; corona purple, about 5 mm dia.; fragrant.

Hoya epedunculata Schlechter In Botanische Jahrbücher 50 (1913) 133. “Die Asclepiadaceae von Deutsch Neu Guinea” R. Schlechter. 43. H. epedunculata Schltr. n. sp. — Suffrutex epiphyticus, parum ramosus, scandens, Rami filiformes, flexuosi, bene foliati, teretes, glabri, Folia patentia vel patula, petiolata, elliptica, acuminata, subcoriacea, utrinque glabra, pedunculo, glabra, crassiusculo, superne leviter sulcato. Inflorescentiae sessiles, umbelliformes, pauci- vel pluri-(ad 5-) florae pedicellis filiformibus, glabris. Flores in sectione inter majores, illis H. torricellensis Schltr. similes sed multo majores. Calyces parvuli foliola ovata, obtusa, margine minute ciliata, extus subverruculosa, quam corolla multoties breviora. Corolla late campanulata, patelliformis, usque ad medium fere 5fida, extus glabra, intus papilloso-puberula, marginibus dense ciliolata, lobis triangulis acuminatis. Coronae foliola patentia antice adscendentia, superne obovata, apice acuta, extus obtusissima, lateribus vix incrassata, Pollinia oblique oblongoidea, translatoribus brevibus, retinaculo rhomboideo, minuto. Ein wenig verzweigter, epiphytischer Schlinger mit schnurartigen, gut beblätterten Zweigen. Blätter 11 —18.5 cm lang, in der Mitte 4.5 — 7.5 cm breit, Blattstiele 1 — 1.3 cm lang. Blütenstande doldig, 1 — 5 blütig ohne gemeinsamen Stiel, Blütenstiele fadenförmig, kahl ca. 3.5 cm lang. Bluten in Sektion ziemlich grosz. Kelch sehr klein, Blättchen ca. 2 mm lang, kurz bewimpert. Korolla ausgebreitet, ca. 4.2 cm im Durchmesser, Lappen am Rande dicht bewimpert. Koronaschuppen von der Spitze bis zum äuszeren Rande etwa 3.5 mm lang. Nordöstl. Neu-Guinea: auf Bäumen in den Waldern des Torricelli-Gebirges, ca, 900 m. u. M. (Schlechter n. 20102. — Blühend im September 1909). Wie schon oben bemerkt wurde, ist die Art nahe verwandt mit H. torricellensis Schltr. Die Grunde, welche mich veranlaszten, beide Pflanzen als besondere Arten anzuchen, sind forgende: Bie H. epedunculata Schltr. sind die Blätter breiter, die Blütendolden sitzend, die Korolla sehr grosz, der Kelch und die Korollaabeschnitte gewimpert und die Koronaschuppen kürzer. Bie H. torricellensis Schltr. haben wir kleinere Blüten in langgestielten Dolden, nicht gewimperte Kelche und Korolla sowie längere Koronaschuppen. Auch die Pollinien sind heir breiter. Die Blüten sind rosenrot mit gelbich-Weiszer Korona. Translation: See entry below.

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In Hoyas of Northeastern New Guinea (1992) 89-90. “Translation of Die Asclepiadaceen von Deutsch Neu-Guinea: R. D. Kloppenburg. 43. Hoya epedunculata Schlechter n. sp. A climbing, epiphytic half shrub with few branches. Branches threadlike, flexible, well leaved, round, glabrous. Leaves spreading or outspread, petiolate, elliptic, acuminate, somewhat leathery, both sides glabrous, with the peduncle glabrous, moderately thick slightly grooved above. Inflorescences sessile, shaped like umbels, few or several flowers (near five), on threadlike, glabrous pedicels. Flowers among the largest in the section, similar to H. torricellensis but much larger. Leaflets (lobes) of the calyx small, ovate, obtuse, margins minutely ciliate, outside covered with somewhat wart like outgrowths, much smaller than those of the corolla. Corolla broadly campanulate (saucer shaped), 5-parted to near the middle, outside glabrous, inside papillose-pubescent, margins densely ciliate, lobes triangular, acuminate. Leaflets (lobes) of the corona spreading, at the front ascending, above obovate, with the apex acute, outside very obtuse, sides scarcely thickened. Pollinia obliquely oblong, with the translators short, retinaculum rhomboid, minute. A little branched epiphytic vine with cordlike, well leaved branches. Leaves 11 18.5 cm. long, in the middle 4.5 - 75 cm. wide, petiole 1- 1.3 cm long. Inflorescences umber like, 1 -5 flowered, without a common (combined) peduncle. Pedicels threadlike, bare, about 3.5 mm. long. Flowers in the section rather large. Calyx very small about 2 mm. long, shortly ciliate. Corolla flattened about 4.2 an in diameter, lobes at the edge thickly ciliate. Corona scales from the tip to the outer edge about 3.5 mm. long. Northeastern New Guinea: on trees in the forest of the Torricelli Mountains about 900 meters altitude (Schlechter #20102 -- Blooming in September 1909). As was already mentioned, the species is closely related to H. torricellensis Schlechter. The reason, that caused me to regard both plants as separate species are the following: with H. epedunculata Schlechter the leaves are wider, the inflorescence is sessile, the corolla is large, the calyx and the corolla lobes are ciliate and the corona scales are shorter. With H. torricellensis Schlechter we have smaller blooms in umbels with long peduncles, more ciliated calyx and corolla, as well as corona scales; also the pollinia are wider. The blooms are rose-red with white corona. In Dr. Schlechter’s Hoya Species (1993) 60. R. D. Kloppenburg. Hoya epedunculata Schlechter. A climbing epiphytic half shrub with few branches. Branches threadlike, flexible, well leaved, round, glabrous. Leaves spreading or outspread, petiolate, elliptic, acuminate, somewhat leathery, both sides glabrous, with the peduncle (petiole) glabrous, moderately thick, slightly grooved above. Inflorescences sessile, umbel like, few or several flowers (near five), on threadlike, glabrous pedicels. Flowers among the largest in the section, similar to H. torricellensis but much larger. Lobes of the calyx small, ovate, obtuse, margins minutely ciliate, outside covered with somewhat wart like outgrowths, much smaller than from the corolla. Corolla broadly campanulate (saucer shaped), 5 parted to near the middle, outside glabrous, inside papillose-pubescent, margins densely ciliate, lobes triangular, acuminate. Scales of the corona spreading, at the front ascending, above obovate, with the apex acute, outside very obtuse, sides scarcely thickened. Pollinia obliquely oblong, with the translators short, retinaculum rhomboid, minute.

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A little branched epiphytic vine with cord like, well leaved branches. Leaves 1118.5 cm. long, in the middle 4.5-7.5 cm. wide, petiole 1-1.3 cm long. Inflorescences umbel like, 1-5 flowered, without a common (combined) peduncle. Pedicels threadlike, bare, about 3.5 mm. long. Flowers in the section rather large. Calyx very small, about 2 mm. long, shortly ciliate. Corona flattened about 4.2 cm. in diameter, lobes at the edge thickly ciliate. Corona scales from the tip to the outer edge about 3.5 mm. long. Northeastern New Guinea: blooming in the forests of the Torricelli Mountains about 900 m. altitude (Schlechter #20102 - Blooming in September 1909). As was already mentioned the species is closely related to H. torricellensis Schlechter. The reason, that caused me to regard both plants as separate species are the following: with H. epedunculata Schlechter the leaves are wider, the inflorescences are sessile, the corolla is large, the calyx and the corolla lobes are ciliate and the corona scales are shorter. With H. torricellensis Schlechter we have smaller blooms in umbels with longer peduncles, calyx and corolla not ciliated, as well as longer corona scales, also the pollinia are wider. The blooms are rose- red with yellowish-white corona. Published by Dr. R. Schlechter in: 1913 Botanische Jahrbücher v.50 p.133.

Hoya erythrina Rintz In Malayan Nature Journal 30 (1978) 501. R. L. Rintz. 13) Hoya erythrina Rintz, sp. nov.* Type: Malaysia, Pahang, Sungai Teranum, Rintz 117 (L). Fig. 17. Distinguishing Features: Stems slender, deep red when young. Leaves coriaceous-fleshy, oblong with strongly corrugate margins and shallowly cordate basis; up to 15 cm long by 5 cm wide; 5-7 veined with a pair of veins parallel to the midrib glossy green or dull red above, red below. Peduncles reflexed, rigid, c. 2 cm long. Umbel positively-geotropic, convex with flexuous, uniform pedicels 2- 2.5 cm long; 1-8 flowers, open 4 days. Corolla lobes spreading, pubescent inside; c. 1.3 cm. diam; pale orange. Corona clear white. Pollinia and corpuscle narrow. Follicle c. 10 cm. long by 3 mm diam. Ecology: Endemic to Malaya, in hill forests in Pahang and Selangor from 400-700 m and at Bertam, Ulu Kelantan; common but not abundant along rivers on both sides of The Gap. Hoya erythrina Rintz, sp. nov., caulibus angustis, rubris in statu junioribus; foliis carnoso-coriaceis, oblongis, marginibus corrugatis, basi lente cordato, supra viridibus vel rubris, infra rubris 5 - 7 venatis; pedunculo reflexo rigido, umbello convexo, pedicellis 1 8; lobis corollae patentibus intus pubescentibus albo-aurantiaceis, corona albo; pollinia angusti; folliculis ad 10 cm longis, 3 mm latis. Typus: Rintz 117 (Pahang), Sg. Teranum 600 m. alt. 25 - 9 -1976 (L!). Also in Selangor, Kelantan. Translation: stems narrow, deep red when young; leaves thick leathery oblong, margines corrugate, base shallowly cordate, above deep green or red, below red with 5 - 6 veins, peduncle reflexed, rigid, umbel convex, pedicels 1-8; lobes of the corolla spreading, inside pubescent whitish-orange, corona white, pollinia narrow; follicles to 10 cm. long, 3 mm. wide. In Hoya Handbook (1992) 69. D. Kloppenburg & A. Wayman. Hoya erythrina Rintz. Why this beautiful hoya escaped notice in the hill forests of Malaya until Dr. R. L. Rintz described it in 1978 is a real mystery. It is so outstandingly unusual and attractive it could

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not have been easily overlooked. It must be that it is fairly rare in nature in the forests of Pahang and Selangor (areas similar to US counties, geographical divisions) where it is found at 400-700 m in altitude. It is said to be common but not abundant along rivers in the areas. This plant is very slow to start growth but with time it will become established and then put on rapid growth. It loves to twine and climb often with long, at first, leafless stems. As leaves develop they are usually deep shades of green and bronze, maturing into rather rigid deep green long undulant leaves with even deeper green venation. The undersides of these leaves is a rich dull maroon to pink, The foliage is very distinctive and beautiful often splotched or marked on the surface with pink markings. You will want to grow this one for the foliage alone! The pendant flower clusters are formed of convex umbels of buttery yellow flowers fussy on the part of the upper curved surface. this is set off with a slightly lighter pagoda shaped center. There is a warm undertone to the yellowish petals. The tip and the edges of the petals are turned under giving a squared off shape to the flowers. This is a plant that most collectors will find very attractive, unique and desirable. Although it is a little slow to start it is one that is worth cultivating and being patient with. I think this is one hoya that should not be over-potted nor over-watered. It might be that it will like more light than most hoyas and probably good humidity, since it grows along streams where there is ample light. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 87. “The Genus Hoya in Thailand” O. Thaithong. 7. H. cf. erythrina Rintz, Malay. Nat. J. 30 (1978) 501, Fig. 17. Occurrence: (PEN): Narathiwat. Notes: The material from Thailand has broader coronal scales than those from Malaysia.

Hoya erythrostemma Kerr In Kew Bulletin (1939) 460. (Siam) A. F. G. Kerr. Hoya erythrostemma Kerr (Asclepiadaceae-Marsdenieae); species H. ellipticae Hook. f. affinis, foliis majoribus acuminatis, corolla villosa, inter alia, differt. Frutex volubilis; rami subquadrati, sat gracilis, dispersim radicantes. Folia elliptica, basi acuta, apice leviter acuminata acuta, usque 10 cm. longa, 4 cm. lata, siccitate papyracea, glabra, costa cum nervis lateribus et transversis utrinque pagina praecipue supra prominulis, nervis lateralibus 5-6 paribus e costa angulo 60° abeuntibus et nervo marginali a margine 3-6 mm. distante conjunctis; petiolus sat robustus, glaber, 36 mm. longus, supra angustissime canaliculatus. Inflorescentia lateralis, umbellata; pedunculus glabra circiter 14 mm. longus; pedicelli graciles, explanati circiter 10 mm. diametro. Calyx 5-partitus; lobi ovati, acuti, glabri, margine minute erosi, 1.5 mm. longa. Corolla 5-lobatus, subrotata sub anthesin reflexa, supra praecipue ad marginem et lobos sat longe villosa; tubus explanatus, circiter 2 mm. longus; lobi late ovati, breviter acuminati, circiter 3 mm. longi, 5 mm. lati. Coronae segmenta cartilaginea, nitentia, lateraliter valde compressa, 3 mm. longa, basi 1.75 mm. alta, apice exteriore acuta, apice interiore antheris paulo breviora. Stigmatis caput umbelliforme, apice umbonatum.

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Folliculi desunt. Tasan, Kloss, 6909. Parkinson 1680, from Thebyu Chaung, South Tenasserim, also belongs here. It differs from the type in its longer petioles, which are up to 2 cm. in length. Translation: H. erythrostemma Kerr (Asclepiadaceae-Marsdenieae) species near H. elliptica Hooker, leaves larger acuminate, corolla velvety, among other differences. Twining shrub, stems somewhat square, moderately slender, with dispersed rootlets. Leaves elliptic, base acute, apex mildly acuminate, up to 10 cm. long, 4 cm. wide, when dry papery, glabrous, midrib with the lateral and transverse nerves on both sides especially above, lateral nerves 5-6 pairs at 60° from the midrib joining the marginal nerves 3-6 mm. from the margines; petioles moderately robust, glabrous, 3-6 mm. long, above narrowly wedge shaped. Inflorescence lateral, umbellate; peduncles glabrous, about 14 mm. long; pedicels slender, glabrous, 12-14 mm. long. Flowers white, corona dull red (from Kloss), flattened about 10 mm. in diameter. Calyx 5 parted; lobs ovate, acute, glabrous, margines minutely eroded, 1.5 mm. long. Corolla 5 lobed, somewhat rotate, below on opening reflexed, above with long velvet mainly at the margines and lobes, tube flattened about 2 mm. long; Segments of the corona horny, shinny, laterally very compressed, 3 mm. base 1.75 mm. tall, exterior apex acute, interior apex a little short of the anthers. Stigma head umbel shaped, apex domed. Follicles not seen. In Florae Siamensis Enumeratio (1951) 36. W. G. Craib & A. F. G. Kerr. Hoya erythrostemma Kerr in Kew Bull. 1939, 460. Surat. Chumpawn, Tasan, Kloss 6909 (type!) Distr. S. Tenasserim (Parkinson 1680!). A species distinguished from its congeners by its laterally compressed corona segments, villous corolla and relatively short corolla lobes. In Malayan Nature Journal 30 (1978) 507-509. R. L. Rintz. 18) Hoya erythrostemma Kerr, Kew Bull. (1939) 460, Type: Thailand, Surat, Chumpawn, Tasan, Kloss 6909 (K). – Fig. 22. Distinguishing Features: Leaves fleshy, elliptical with cuneate bases; 8-9 cm long by 2.5-3 cm wide; the leaves resemble those of H. parasitica but are not so thick and the veins are readily visible. Peduncle horizontal, rigid, 3-4 cm long. Umbel negativelygeotropic, convex with rigid, uniform pedicels c. 2 mm long; 1-30 flowers, open 4 days. Corolla spreading, pubescent inside; c. 1 cm diam; white. Corona lower lobe inflated and very thin horizontally; upper lobe deep red, lower lobe white. Corpuscle oval, Follicle c. 14 cm buy 4 mm diam. Ecology: Endemic to Malaysia and S. Thailand: very common along lowland rivers in Selangor, often draping the trees in the manner of H. parasitica. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 88. “The Genus Hoya in Thailand” O. Thaithong. 8. H. erythrostemma Kerr, Kew Bull. (1939) 460-461. Fl. Siam. Enum. 3(1) (1951) 36; Malay. Nat. J. 30 (1978) 508. Fig. 22. Occurrence: (PEN): Chumphon, Nakhon Si Tharmmarat, Songkhla (SW): Prachuab Kiri Khan.

Hoya esculenta Tsiang

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In Sunyatsenia 3 (1936) 176-177. Tsiang. Hoya esculenta (Rumphius) Tsiang, comb. nov. Sussuela esculenta Rumphius, Herb. Amb. 5: 467, t. 175, f, 2, 1747. Hoya diversifolia Blume, Bijdr. 1064. 1826; G. Don Gen. Syst. 4: 127, 1838; Decne. in DC., Prodr. 8: 636. 1844; Hook f., Fl. Brit. Ind. 4: 61. 1883; Ridley, Fl. Malay Penins. 2: 396. 1923. Hoya orbiculata Wall. ex Wight, Contr. Bot. Ind. 36. 1834; G. Don. Gen. Syst. 4: 125. 1838: Decne. in DC., Prodr. 8: 635. 1844. Malay Peninsula: Tanjong, Md. Nur 21757, April 26, 1929. Distribution: Burma: Malaya. This and the following species (H. liangii) have penninerved leaves and exterior angle of the coronal-lobes rotundate or very obtuse. The plate of Sussuela esculenta shows that the leaves are of the same shape but much smaller. Costantin’s figure in Leconte, Fl. Gen. Indo-Chine 4: 131, fig. 18, 6-8, shows a plant of doubtful status, so for time being, I eliminate the record for Indo-China.

Hoya esquirolii Leveille In Repertorium Specierum Novarum 11 (1912) 298. Leveille. 942. Hoya Esquirolii Levl., nov. spec. Ab H. nummularia Dcne. distincta floribus minutis umbellis sessilibus; flores albi 2 mm circiter lati et longi. Kouy-Tcheou: Lo-Fou, sur arber, mars 1909 (Jul. Caylerie, 3484); au bac de Pai-Ouai; pend en longues ficelles des rochers et des vieux troncs, 20 mai 1912 (Jos. Esquirol, 2801). Translation: flowers distinct from H. nummularia with minute sessile umbel Dcne. s; flowers white 2 mm. about wide and long. In Flore du Kouy Tcheou (1914) 42. Hoya Esquirolii Levl. Co fou sur abre: bac de Pia. ouai mars mai 1902-09. Jos. Esquirol 2801. 3482. Beihefte zum Botanischen Centralblatt 34/2 (1912) 14. D. Pain “Neue Asclep. von Sumatra und Celebes”. Leveille, H., Decades plantarum novarum, XCIII — C. (Rep. Spec. nov. XI. p. 295 — 307, 1912.) Editors note: Among the lengthy list is Hoya Esquirolii. In Journal of the Arnold Arboretum 15 (1934) 318. “Notes on the Ligneous Plants Described by Leveille from Eastern Asia” Hoya Esquirolii Leveille in Fedde, Rep. Spec. Nov. XI. 298 (1912) China. Kweichou: Lo-fou, sur arbe, J. Cavalerie, no. 3484, March, 1909 (Syntype; photo. in A. A.); au bac de Pia-ouai, pend en longues ficelles des rochers et des vieux troncs, J. Escuirol, no. 2801, May 20, 1912 (Syntype; photo in A. A.). Either specimen is without flowers for dissection; consequently a decisive opinion cannot be rendered concerning this plant, the foliage of which resembles to some extent that of H. nummularia Dcne,. with which it may be conspecific. In Catalogue of Names Publisher by Hector Leveille. (1978) 134. L. A. Lavener. 1437. Dischidia esquirolii (Levl.) Tsiang in Sunyatsenia 3: 183 (1936) & l.c. 4:127 (1939);

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Rehder in Jour, Arn. Arb. 18:241 (1937). Hoya esquirolii Levl. in Fedde, Rep. Sp. Nov, 11:298 (1912) & Fl. Kouy-Tcheou 42 (1914); Woodson in Jour. Arn. Arb. 15:318 (1934). In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. esquirolii Leveille – 67.

Hoya excavata Teijsmann & Binnendijk In Natuurkunndie Tijdschrift voor Nederlandsch Indie 25 (1863) 406. J. E. Teijsmann & S. Binnendijk. 68. Hoya Excavata. H. volubilis glabra, foliis subcarnosis, ovatooblongis, basi rotundatis, apice breviter apiculatis, sepalis ovatis , acutiusculis; corollae laciniis supra glandulosis, ovatis, acutis, erectiusculis, marginibus reflexis; coron. stam. foliolis subrotundatis, inferne apiculata-attenuatis, supra excavatis, subtus sulcatis, stigmate apiculato. Hab. ins. Boeroe district, Okie. Archipelagi Moluccani, Teijsmann. Frutex alte scandens. Caulis teres, glaber, volubilis. Petioli firmi, glabri.0.025 longi. Folia opposita, 0.15 longa 0.09 lata, ovato-oblonga, basi rotundata, vel obtusa, apice rotundata, breviter apiculata, subcarnosa, supra nitida, utrinque nervis 6, ad petiolum callosoglandulosa Pedunculi petiolis fere aequilongi, nutantes, superne crassiores, virides, glabri; flores in receptaculis brevissimis fasciculati. Pedicilli pedunculo duplo longiores, filiformes, virescente-albidi. Calyx parvus, 5-partitus, laciniis acutis, virescens, membranacens, ciliolatis. Corolla subcarnosa, 5 fida, albida, intus basi violacco-stellata, brevissime papillosa, laciniis ovatis, acutis, erecto-patentibus, vel apice reflexis; coronae stamineae foliola carnosa, cereacea, supra concava, in fando excavationis elevatopunctata, subtus marginibus revolutis, quasi fissura longitudinali excisa, angulum interius in dentem acutum, erectiusculum productum. Antherae sagittatae, membrana tennis. Pollinia basi per crura brevia affixa. Retinacula complicata, deltoidea, stigma abbreviatoprismaticum. Ovaria duo. Uit den botanischen tuin te Utrecht, werd in 1848 eene plant ontvagen, met den naam H. cunninghami, die zoo nabij deze nieuwe soort komt, dat men ze in de bloemen voor eene en dezelfde soort zoude bouden. Het verschil is voornamelijk te vinden in de bladeren, zijude die van H. Cunnighami dik, de nerven op de boven-en ondervlakte naauwelijks zigtbaar, 0.08 lang en 0.03 breed. Het receptaculum is gerekt en schubbig. De bloem is een weinig kleiner en de reuk heeft eenige overeenkomst met Philadelphus coronarius, terwijl deze soort van Boeroe een sterken Vanilje reuk bezit. Translation: Hoya twining, glabrous, leaves somewhat fleshy, oval-oblong, base rounded, apex briefly apiculate, sepals oval, apexes briefly apiculate, sepals ovate, somewhat acute; lobe of the corolla glandular, ovate acute, somewhat erect, margines reflexed; leaflets of the staminal corona somewhat rounded, lower apiculate-attenuate, above excavated, below grooved, stigma apiculate. Habitat on the Island of Borneo, Okie district, Molucca archipelago, Teijsmann. A high climbing scrub. stems round, glabrous, twining. Petiole firm, glabrous,0.025 long. Leaves opposite 0.15 long 0.09 wide, ovate-oblong, base rounded, or obtuse, apexes rounded, briefly aciculate, somewhat fleshy, above shinny, both sides with 6 nerves, with the petiole callous-glandulous. Peduncle petioles nearly equal long, nodding extra fleshy green, glabrous; flowers from briefly fascilated receptacles. Pedicels are twice as long,

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filiform, becoming greenish-white. Calyx small, of 5 parts. lobes acute, greenish membranaceous ciliate. Corolla somewhat fleshy, 5 lobed, whitish, inside near the base with bluish white hairs, shortly papillose, lobes ovate, acute, spreading upright, or with the apex reflexed; leaflets of the staminal corona fleshy, leathery, concave above, with the base excavated elevated and punctate, margins below revolute, as if cut with a longitudinal fissure, internal angle with a acute tooth, erectly produced. Anthers arrowshaped, a thin membrane. Pollinia affixed at the base portion. Retinacula complicated, deltoid, stigma abbreviated Prism-shaped. Ovaries 2. A plant was received from the Botanical Garden in Utrecht in 1848 with the name Hoya cunninghami, that is so close to this new species that as concerns the blooms one should regard it as one and the same species. The difference is especially to be found in the leaves; those of H. cunninghami are thick and the veins on the lower surface are scarcely visible, being 0.08 long and 0.03 wide. The receptacle is lengthy and scaly. The flower is a little smaller and the fragrance has a similarity to Philadelphus coronarius (mock orange), while this species from Borneo has a strong vanilla small.

Hoya exilis Schlechter In Botanische Jahrbücher 50 (1913) 121. “Die Asclepiadaceen von Deutsch Neu Guinea” R. Schlechter. 28. H. exilis Schltr. n. sp. — Suffrutex gracillimus, exilis, parum ramosus, scandens. Rami filiformes, flexuosi. laxe foliati, teretes, glabri. Folia patentia vel patula, elliptico-lanceolata, obtuse longi-acuminata, textura pro genere tenuia, utrinque glabra, petiolo glabro, superne leviter sulcato. Inflorescentiae umbelliformes, 6 — 15 florae, pedunculo pedicellisque gracillimis filiformibus glabris. Flores in sectione inter minores, subglabri. Calycis foliola parvula, lanceolato-ovata, usque ad tertiam partem basilarem 5-fida, extus glaberrima, intus basi minute puberula, caeterum glabra, lobis recurvis, rhomboideo-ovatis, acutis. Coronae foliola patentia, superne oblonga, apice adscendentia, breviter acuminata, extus obtusa, lateribus rotundato-incrassata. Pollinia oblique ovalia, translatoribus brevissimis, retinaculo rhomboideo, minuto. Ein äuszerst zierlicher Schlinger mit fadenförmigen, locker beblätterten Zweigen, Blatter sehr dünn von Textur, 5 - 10 cm lang, in der Mitte 1.5 - 3.3 cm breit, Blattstiele 0.4 - 0.7 cm lang. Blütenstände doldig, 6 - 15 blütig, auf sehr schankem, 5 - 6 cm langen Stiel. Blütenstiele sehr fein fadenförmig, 1.5 - 2 cm lang. Kelch sehr klein, Zipfel ca. 1 mm lang. Korolla 1.1 cm in Durchmesser. Koronaschuppen kurz und fleischige, vor der Spitze bis zum äuszeren Rande kaum 2 mm überschreitend. Nordöstl. Neu-Guinea: auf Bäumen längs der Gebirgsbäche von Albo, ca. 300 m. u. M. (Schlechter n. 16172. — Blühend im Juni 1907); auf Bäumen in der Waldern des Kani-Gebirges, ca. 700 m. u, M. (Schlechter n. 17865. — Blühend im Juni 1908). Eine schon äuszerlich durch die sehr dünnen Zweige und die auffallend dünne Textur der Blatter leicht zu erkennende Art. Die geringe Blütengrösze bringt sie neben H. chloroleuca Schltr., doch sie in den Koronaschuppen sehr verschieden vor jener, Die Blutenfärbung ist grünlich-weisz. Translation: See article below. In Hoyas of Northeastern New Guinea (1992) 57-58. “Translation of Die Asclepiadaceen von Deutsch Neu-Guinea: R. D. Kloppenburg. 28. Hoya exilis Schlechter n. sp.—An

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extremely slender, small, not very branched climber. Branches threadlike, flexible, loosely leaved, round, glabrous. Leaves spreading or outspread, elliptic-lanceolate, obtuse, long-acuminate, texture for the genus thin, both sides glabrous, petiole glabrous, above slightly grooved. Inflorescences shaped like umbels, 6-15 flowered. Pedicels of the peduncle very slender, threadlike, glabrous. Flowers among the smallest in the section, almost glabrous. Leaflets (lobes) of the calyx small, lanceolate ovate, subacute, glabrous, much shorter than those of the corolla. Corolla somewhat rotate, 5-parted all the way to one third from the base, outside glabrous, inside at the base minutely puberulous, central portion glabrous, lobes recurved, rhomboid-ovate, acute. Leaflets (scales) of the corona spreading, above-oblong, with the apex ascending, shortly acuminate, outside obtuse, sides rounded-fleshy. Pollinia obliquely-ovate, with the translators very small, retinaculum rhomboid, minute. An extremely pretty clinging vine with threadlike loosely leaved branches. Leaves very thin textured, 5 - 10 cat long, in the middle 1.5 - 3.3 cm. wide. Petioles 0.4 -0.7 cm. long. Inflorescences umbellate 6-15 flowered, on a very slender peduncle, 5 -6 cm. long. Pedicels very fine, threadlike, 1.5-2 cm. long. Calyx very small, lobes about 1 mm. long. Corolla 1.1 cm. in diameter. Corona scales short and fleshy, from the apex to the outer rim barely over 2 mm. Northeastern New Guinea: on trees along mountain brooks of the Albo, about 300 meters altitude (Schlechter #16172—Blooming in June 1907); on trees in the Kani Mountains about 700 meters altitude (Schlechter #17865—Blooming in June 1908). A beautiful species, easy to recognize by means of its very thin branches and the diminished, thin texture of the leaves. The small bloom size brings it near H. chloroleuca Schlechter, however it is very different from that one in the corona scales. The coloring of the blooms are greenish-white. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. exilis Schlechter (Schlechter 17430, 19410) – 15. In Hoya Dr. Schlechter’s Hoya Species (1993) 62. R. D. Kloppenburg. Hoya exilis Schlechter. An extremely slender, small not very branched climber. Branches threadlike, flexible, loosely leaved, round, glabrous. Leaves spreading or outspread, ellipticlanceolate, obtuse, long- acuminate, texture for the genus thin, both sides glabrous, petiole glabrous, above slightly grooved. Inflorescences shaped like an umbel, 6-15 flowered, pedicels of the peduncle very slender, threadlike, glabrous. Flowers among the smallest in the section, almost glabrous. Lobes of the calyx small, lanceolate-ovate, subacute, glabrous, much shorter than from the corolla. Corolla somewhat rotate, 5parted all the way to one third from the base, outside glabrous, inside at the base minutely puberulous, central portion glabrous, lobes recurved, rhomboid-ovate, acute. Scales of the corona spreading, above oblong, with the apex ascending, shortly acuminate, outside obtuse, sides rounded-fleshy. Pollinia obliquely-ovate, with the translators very small, retinaculum rhomboid, minute. An extremely pretty clinging vine with threadlike loosely leaved branches. Leaves very thin textured, 5-10 cm. long, in the middle 1.5-3.3 cm. wide. Petioles 0.4-0.7 cm. long. Inflorescences umbellate 6-15 flowered, on a very slender peduncle, 5-6 cm. long.

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Pedicels very fine threadlike, 1.5-2 cm. long. Calyx very small, lobes about 1 mm. long. Corolla 1.1 cm. in diameter. Corona scales short and flashy, from the apex to the outer rim barely over 2 mm. Northeastern New Guinea: on trees along mountain brooks of the Albo, about 300 m. altitude (Schlechter #16172 - Blooming in June 1907); on trees in the Kani Mountains about 700 m. altitude (Schlechter #17865 - Blooming in June 1908). A beautiful species easy to recognize by means of its very thin branches and the diminished, thin texture of the leaves. The small bloom size brings it near H. chloroleuca Schlechter, however it is very different from that one in the corolla scales, The coloring of the flowers are greenish-white. Published by Dr. R. Schlechter in:1913 Botanische Jahrbücher V.50 p.121, “Die Asclepiadaceen von Deutsch Neu Guinea".

Hoya filiformis Rechinger In Repertorium Specierum Novarum 5 (1908) 132. K. Rechinger. 17. Hoya filiformis Rechinger, nov. spec. Caule scandente tenui, filiformi (in sicco 1— 2 mm diametro) glaberrimo ut tota planta), foliis omnibus oppositis geminis, tenuibus penninerviis, lanceolato-ovatis, acuminatis, acumine prostrato. Foliis tenuibus plane reticulato-venosis, glaberrimis lucidis. Inflorescentiis umbelliformibus, pedunculo brevi gracili axillari 7 — 8 mm longo, pedicellis tenuissimis pedunculo aequilongis. Floribus minutis albis glabris, calycis lanciniis minutissimis lanceolatis acuminatis. Lobis corollae e basi lata obtuse angustatis planis. Fructibus maturis 13 — 14 cm longis. Samoa Insel Upolu, in grosser Menge im Kammgebiete ober Utumapu auf neideren Bäumen. Juli 1905 (No. 924, 1500, 1701 Rechinger). Blattextur dünn, Blätter onhe Blattstiel 6 cm lang, 2 cm breit, davon entfällt auf die Träufelspitze nahezu 1 cm. Blattstiel 8 mm long. Blätter zart aber doch deutlich fiedernervig, die Nerven sind auf der Blattoberund Unterseite sichtbar, das der Blattbasis zunächst aus dem Medianus entspringende Nervenpaar ist fast bis über die Mitte des Blattes hinaus randläufig. Blüten weiss, wenig duftend, denen der H. pycnophylla mihi ähnlich, aber noch kleiner. Blüttendolden etwa 2 cm im Durchmesser. Bemerkenswert durch die langausgezogenen Spitzen der Laubblätter und denen dünne Textur. Die ganze Pflanze ist volkommen kahl. Translation: A thin stemmed climbing plant, threadlike (in the dry state 1 - 2 mm in diameter) the total plant glabrous, leaves entire in opposite pairs, thin penninerved, lanceolate-ovate, acuminate, acumen procumbent. Leaves thin visibly reticulate veined, very glabrous bright. Inflorescence like an umbel, peduncle short, glabrous axillary 7 — 8 mm long, pedicels very thin of equal length to the peduncle. Flowers small white, glabrous, calyx lobes very small lanceolate acuminate. Lobes of the corolla broad at the base, obtuse narrowly flat. Fruit at maturity 13 — 14 cm long. Island of Upolu, Samoa, in great quantity in the Hill Region above Utumapu on the low trees. June 1905 #924, 1500 and 1701 Rechinger. Leaf texture thin, leaf exclusive of the petiole 6 cm long, 2 cm wide, 1 cm near the tapered point turns down. Petioles 8 mm long. Leaves are delicately feather nerved, nevertheless the nerves are distinct on the upper surface and visible on the lower surface, first of all from the median of the leafbases extend outward in pairs almost to the edge of the leaf. Flowers white, not much scent, and to me much like those of H. pycnophylla but

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yet smaller. The umbel is 2 cm in diameter. Noteworthy through the long twisted apex of the leafblade and the fine thin texture. The entire plant is completely bald. In Denkschriften d. kais Akad. d. Wiss Math-Natur. 75 (1908) 334. K. Rechinger. “Botanisch-Zoologische Ergebnisse von den Samoainseln”. * Hoya filiformis Rechinger in Fedde, Repert. novar. specier., Bd. V, p. 132 (1908). Caule scandente tenui, filiformi (in sicco 1 —2 mm diametro) glaberrimo ut tota planta, foliis omnibus oppositis geminis, tenuibus penninerviis, lanceolato-ovatis, acuminatis, acumine prostrato, tenuibus, plane foliis omnibus oppositis geminis, tenuibus penninerviis, lanceolato-ovatis, acuminatis, acumine prostrato, tenuibus plane reticulato-venosis, glaberrimis lucidis. Inflorescentiis umbelliformibus, pedunculo brevi gracili axillari 7 — 8 mm longo, pedicellis tenuissimis pedunculo aequilongis. Floribus minutis albis glabris, calycis lanciniis minutissimis lanceolatis acuminatis. Lobis corollae e basi lata obtuse angustatis planis. Fructibus minutis albis glabris, calycis laciniis minutissimis lanceolatis acuminatis Lobis corollae basi lata obtuse angustatis planis. Fructibus maturis 13 — 14 cm longis. Insel Upolu: In groszer Menge auf dem Kammgebiet ober Utumapu auf neideren Baümen. Juni, Nr. 924, 1500, 1701. Blattextur dünn, Blattlamina 6 cm lang, 2 cm breit, davon entfällt auf die Träufelspitze nahezu 1 cm. Blattstiel 8 mm long. Blätter zart aber doch deutlich fiedernervig, die Nerven sind auf der Blattober-und-unterseite sichtbar, das der Blattbasis zunächst aus dem Medianus entspringende Nervenpaar ist fast bis über die Mitte des Blattes hinaus randläufig. Die Blüten sind weiss, wenig duftend, denen der H. pycnophylla Rechinger ähnlich, aber noch kleiner. Die Blüttendolden etwa 2 cm im Durchmesser. Ausgezeichnet durch die langausgezogenen Spitzen der Laubblätter und durch dünne Texture. Die ganze Pflanze is vollkemmen kahl. Translation: Almost identical to the above:

Hoya finlaysonii Wight In Contributions to the Botany of India (1834) 38. R. Wight & Wallich. 17. H. Finlaysonii (Wight:) volubilis glabra, ramis teretibus, foliis oblongo-lanceolatis obtuse acuminatis supra (siccitate) reticulatis subtus subaveniis (5-7 uncias longis 1 ½- 2 latis) brevi petiolatis. — Hoya, Wall.! Asclep. n. 42. — Penang ?; Wallich et Finlayson. No station is attached to the specimens, but from the same plant existing in Wallich’s and Finlayson’s herbaria, it is probably from Penang. — (R. W.) Translation: twining glabrous, branches round, leaves oblong-lanceolate obtuse acuminate above (dried) reticulate, below somewhat veinless (5-7 inches long 1 ½ -2 wide) petioles short. I have seen the Hoya, from Wallich. Asclepias number 42. — maybe from Penang. ; Wallich & Finlayson. In General System of Gardening & Botany 4 (1837) 127. G. Don. 31 H. *Finlaysoniana (Wight, l. c.) twining, glabrous; branches terete; leaves oblong-lanceolate, bluntly acuminated, reticulated above in the dried state, veinless beneath, on short petioles. Perennial woody shrub. Native, probably to Penang. Hoya. Wall. ascl. no. 42. Leaves 5-7

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inches long, and 1-2 broad. Finlayson’s Hoya. Shrub twining. *Note: incorrect spelling. In Synopsis Plantarum 6 (1840) 892. D. N. F. Dietrich. 17. H. Finlaysonii Wight; volubilis glabris; ramis teretibus; fol. oblongo-lanceolatis obtuse acuminatis supra reticulatis subtus aveniis. In Penang. woody. Translation: twining glabris; branches round; leaves oblong-lanceolate obtuse acuminata above reticulate below veinless. From Penang (Malaya). woody. Prodramus System Veg. (DeCandolle) 8 (1844) 638. Decaisne. 31. H finlaysonii (Wight, contrib., p 38), volubilis glabris, ramis teretibus, foliis oblongo-lanceolatis obtuse acuminatis supra (siccitate) reticulatis subtus subaveniis (5-7 uncias longis 1 ½- 2 latis) brevi petiolatis. Penang ?; Wallich et Finlayson. Translation: Same as type description of Wight’s above. In Flora van Nederlandsch Indie Batavae 1 (1856) 527. F. A. W. Miquel. 38. Hoya finlaysonii Wight Contr. p. 38. Decaisn. l. c. p. 638. Volubilis glabra, ramis teretibus, folia oblongo-lanceolata obtuse acuminata, supra (in sicco) reticulata, subtus subavenia, 5 - 7 poll. longa. breviter petiolata. - Poeloe Penang? (Wall) Note this is under “Species dubiae:” Translation: essentially as above. In The Flora of British India 4 (1883) 62. J. D. Hooker (under doubtful and excluded species) H. finlaysonii, Wight Contrib. 38; Wall. Cat. 8166; Dcne l. c.; stem stout leaves 5-7 by 2 in ovate-lanceolate obtusely acuminate densely thick penninerved, nerves prominent on both surfaces arched, intermarginal very distinct, petiole short stout. Singapore, Wallich; Siam, ? Herb. Finlayson. In Flora of the Malaya Peninsula 2 (1903) 574-575. King & Gamble 17. Hoya finlaysonii, Wight Contrib. 38 (1834). A climbing shrub: stem stout; branchlets pale brown, smooth, lenticellate, flexuose. Leaves coriaceous, thick, hard and tough when dry; ovate- or oblong-lanceolate, obtusely acuminate at apex, more or less rounded at base; both surfaces at first minutely papillose, afterwards glabrescent or dotted, when fresh greenish-red above, dark red beneath; margins with a wavy nerve close to the edge; 5 to 7 in. long, 1.5 to 2.5 in. broad; nerves very prominent and raised on both surfaces when dry; midrib stout, with a large gland just above the petiole; main nerves about 5 to 7 pairs besides the marginal one, at about 60° with the midrib, nearly straight to join a second and prominent looped intramarginal nerve; secondary nerves many, irregular; reticulations irregular netted; petiole very thick, .25 to .5 in. long. Umbels very manyflowered, without any prominent tubercular rachis; peduncles .75 to 2 in. long, rather slender; pedicels almost capillary, .5 to .75 in. long, buds 5-angled, hemispheric below, fattened and depressed above! .25 in. across; flowers with cream-colouled and pink corolla and wavy-white corona. Calyx thin, nearly hyaline, with a few brown veins; lobes obtuse, .05 in. long, pubescent within. Corolla 4 in. broad when open, pubescent within; lobes triangular-ovate-acute, usually inflexed. Corona of 5 spreading inflated processes; the lower lobes ovate-acute, concave at first, afterwards more or less ridged, 2-winged

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below; the upper-lobe erect, short, acute. Staminal-column short, attached to the top of' the corolla-tube; anther-cells divergent; appendages short, rounded, scarious; pollen-masses oblong-falcate, obtuse at tip, attached by very short thick caudicles to the conical pollen-carriers. Style-apex 5-angled, with 5 rays leading to a short apiculus. Follicle 1, very slender, 7.5 in. long, .25 in. broad, straight, acuminate; pericarp thin, glabrous, striate. Seeds cylindric, 3 in. long, acute at base, truncate at tip, and bearing a 1 to 1.5 in. long silky coma, teste very thin, brown; cotyledons fleshy; lanceolate, .15 in. long; radicle cylindric, 1 in. long. Wall. Cat. 8166; Dcne. in DC. Prodr. VIII. 638; Fl. Ind. Bat. II. 527; Hook. f. Fl. Br. Ind. IV. 62. Perak: Wray 2724; King’s Collector 4800. Singapore: Wallich 8166.—Distrib. Siam? (Finlayson). The specimens in the Kew Herbarium have no flowers, but we see no reason to doubt the identification of the leaves of our specimen. In Flora of the Malaya Peninsula 2 (1923) 394; 397. H. Ridley. (10) H. finlaysonii Wight, Contrib. 38; King, l. c. 574. Stem fairly stout. Leaves coriaceous, ovate or oblong lanceolate, blunt acuminate, base rounded or narrow blunt, greenish red above, dark red beneath; nerves 5 to 7 pairs joining an intermarginal, elevate as are reticulations when dry; 5 to 7 in. long, 1.5 to 2.5 in. wide; petioles thick, .25 to .5 in. long, a gland just above it. Peduncles 1.75 in. long, rachis very short; pedicels .75 in. long. Flowers numerous, cream-colour and pink, and corona white; .4 in. across. Corolla pubescent inside, lobes triangular-ovate acute, Corona-lobes ovate acute. Follicle 1, very slender, 7.5 in. long, .25 in. wide. Seeds cylindric, .3 in. long, base acute, tip truncate, plume 1.5 in. long. Hab. Singapore (Wallich 2724). Perak, Goping (Kunstler). Rare. In Florae Siamensis Enumeratio 3 (1951) 36. W. G. Craib & A. F. G. Kerr. Hoya finlaysonii Wight, Contrib. Bot. Ind. 38 (1834); F.B.I. iv. 62, inter spp. dub. F.M.P., No. 19 574 (784); F.M.P. ii 397. Rachaburi. Prachuap, Hui Tap Sake, 60 m., Winit 614! Surat. Ban na Yuang 1! Puket. Pang-nga, Nop Pring, c. 100m., evergreen forest, Kerr 19357! Dist. Pen. Mal.! (type! Penang ?) In Malayan Nature journal 30, 3/4 (1978) 511. R. L. Rintz. 21) Hoya finlaysonii Wight, Contr. 38 (1834). Type: Malaysia, Melaka, Wight (K). — Fig, 25, Distinguishing features: Leaves thick and rigid, elliptical with cuneate bases and rough, finely serrate margins; up to 21cm long by 6cm wide; veins prominent, wide, dark green on a lighter green background. Peduncle horizontal, rigid, 2 —3cm long. Umbel negatively-geotropic, convex with rigid, uniform pedicels c. 2.5cm long; 1— 40 flowers, open 12 hrs. Corolla reflexed, finely pubescent inside; c. 8mm long by 6mm diam; yellow with deep red tips. Corona white. Follicles c. 12cm long 6cm diam. Ecology: Along rivers in lowland forests of Pahang, Perak and Selangor; not common. Distribution: S. Thailand, Sumatra, Borneo. In The Hoya Handbook (1992) 70. D. Kloppenburg & A. Wayman. Hoya finlaysonii Wight. Each hoya seems to have some distinctive and outstanding character. In the case of this hoya it is the striking foliage. At once you can pick this one out from all

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the rest. The leaves are thick and rigid, elliptic with narrowing bases. Most of all it is recognized by the contrast between the light green leaves with prominent deep emerald green venation and the deep green edge to each leaf. They are spectacular, they stand out. The plant would, in my estimation, be worth growing for the foliage alone. In its native environment it is not a common plant. It comes from Malaya, South Thailand and Borneo. This plant will attract you by its foliage alone. The leaves are medium sized but often much larger than our copy depicts, especially when it reaches maturity. I have found that it roots slowly and takes some time before it establishes itself and really begins to grow rapidly. It may be my conditions, however I feel it is one that is worth waiting for, and once established displays strong growth. Because of the long internodes (space between leaf pairs) it will help to wrap the plant around itself to give a more compact appearance. In hanging baskets let some stems twine around the hanger and then go on to droop back downward. The flower clusters are compact balls of reflexed waxy flowers. The centers are creamy white with some yellow tones and contrast well with the bronze-maroon reflexed petal lobes. This plant is a conversation piece and another species you will surely want in your collection In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 88. “The Genus Hoya in Thailand” O. Thaithong. 9. H. finlaysonii Wight, Contib. Bot. Ind. (1834) 38. Fl. Siam. Enum. 3(1), (1951) 36; FMP 2 (1934)* 397; Malay. Nar. J. 30 (1978) 511 &514, Fig. 25. Occurrence: (SW): Pranchuab Kiri Kahn. (PEN): Surat Thani, Phuket, Phang-nga, Trang.

Hoya fischeriana Warburg In Fragmentia Florae Philippinae 1 (1904) 129. (Perkins) R. Schlechter & O. Warburg. 3. Hoya Fischeriana Warb. n. sp. Folio crasse coriaceo late lanceolato 12 — 19 cm longo 4 — 7 cm lato basi et apice acuto glabro 5-nervio, nervis internis apicem fere attingentibus, reticulo nervorum distincto, petiolo 2 — 3.5 cm longo 2 — 3 mm crasso; pedunculo ca. 5 cm longo 2 mm crasso, dimidio superiore florigero cylindrico 3 mm crasso; pedicellis glabris capilliformibus ca. 1.5 cm longis, sepalis late lanceolatis 1 mm longis glabris, corolla 7 mm in diametro extus glabra intus subpapillosa profunde lobata, lobis rhomboideis, coronae stamineae lobis patulis late lanceolatis utrinque acutis, apice interno, elevato, apice externo haud recurvo; fructuum pedicellis 2 cm longis, fructibus 12 cm longis 4 mm latis, coma fere 3 cm longa. Nord Luzon, Prov. Isabela, Malunu (Warburg no. 11955). Diese mit H. cinnamomifolia Hook verwandte, aber durch viel kleinere Blüten und schmälere lanzettliche, beiderseits spitze Koronallappen ausgezeichnete Art ich Ehren das Tabakpflanzers Fischer benannt, der in Malunu mich in liebenswürdiger Weise beherbergt hat. Translation: Leaves thick leathery broadly lanceolate 12 — 19cm long 1 — 7cm wide base and apex acute glabrous 5-nerved, nerves inside almost reaching the apexes, nerve reticulations distinct, petiole 2 — 3.5cm long 2 — 3mm thick; peduncles about 5cm long 2 cm thick, upper half cylindrical 3mm thick; pedicels glabrous hair-like about 1.5cm long, sepals broad lanceolate 1mm long glabrous, corolla 7mm in diameter outside

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glabrous inside somewhat papillose deeply lobed, lobes rhomboid, lobes of the staminal corona spreading broadly lanceolate both ends acute, internal apex elevated, outer apex by no means recurved; fruit pedicels 2cm long, fruit 12cm long 4mm wide; seeds tawny linear flattened 4mm long 3/4mm wide, coma nearly 3cm long. Northern Luzon Isabela Province, Malunu (Warburg # 11955). This is related to Hoya cinnamomifolia Hook , but differs through the smaller blooms and both ends of the slender lance-like coronal lobes I have named this species in honor of the wife of my highly esteemed friend, the Malunu Tobacco Planter, Fischer, with whom I have lodged. In An Enumeration of Philippine Flowering Plants 3 (1922) 352. E. D. Merrill. Hoya fischeriana Warb. in Perk. Frag. Fl. Philip. (1904) 129. Luzon (Isabela, Ilocos Norte), Mindoro, Biliran, Dinagat, Mindanao (Surigao, Davao), Copeland 421m McGregor 333, B.S. 32906, 32905 Ramos, 18529 McGregor, 35256, 34526 Ramos & Pascasio. On trees at low altitudes. Endemic. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. fischeriana Warb. – 32. Philippine Hoya Species 3rd Ed. (1996) 71. R. D. Kloppenburg. Hoya fischeriana Warburg in Perkins Fragmentia Florae Philippinae 1 (1904) 129. Type: Warburg #11955, Malunu, Isabela Prov., Luzon. Key #14 Leaves thick coriaceous, broadly lanceolate, 12-19 cm. long, 4-7 cm. wide, both base and apex acute, glabrous with 5 nerves inside almost reaching the apex, reticulated, all distinct. Petiole 2- 3.5 cm. long 0.2-0.3 cm. in diameter. Peduncle about 5 cm. long, 0.2 cm. thick. Rachis cylindrical 0.3 cm. in diameter. Pedicel glabrous, filiform about 1.5 cm. long. Sepals broadly lanceolate 0.1 cm. long, glabrous. Corolla 0.7 cm. in diameter, outside glabrous, inside somewhat papillose, deeply lobed, lobes rhomboid. Corona scales spreading , broadly lanceolate both ends acute, apexes of inner lobe elevated, outer lobe not recurved. Fruit peduncles 2 cm. long 0.04 cm. in diameter. Seeds tawny, linearly flattened out 0.4 cm. long, 0.075 cm. broad. Comma almost 3 cm. long. Luzon (Isabela, Ilocos Norte) Mindoro, Biliran, Dinagat, Mindanao (Surigao, Davao). Enumeration of Philippine Plants 3 (1923) 352 Merrill; The Hoyan 9 #2 (1987) 23- 26. Copeland #421; Mc Gregor 333, #18529 1914; Ramos #32905 (BS), #32906 (BS); Ramos & Pascasio #35256 1959 (PHU,UC),.Ramos & Edano #47136 1929 (BO,B,UC) #85130 (A), #75526 1928 (BO,UC), #49395 1927 (UC); Edano #40187 1959 (PNH), ? ; Clemens #16790 1926 (UC); #17136 1930. In Botanische Jahrbücher 50 (1913) 114-115 R. Schlechter “Die Asclepiadaceae von Dutch Neu Guinea”.

Hoya flagellata Kerr In Icones Plantarum 35 (1940) t.3407. Kerr. H. flagellata Kerr; species nova, H. caudatae Hook. f. affinis, foliis floribusque minoribus, corollae lobis pro rata longius

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caudatis distinguenda. Suffrutex epiphyticus; rami graciles, glabri. Folia ovata vel lanceolata, basi rotundata et minute auriculata, apice acute, margine plus minusve revoluta, 4.7-8.5 cm. longa, 2-2.5 cm. lata, coriacea, siccitate pallide brunnea, glabra nisi supra interdum pills minutis sparsissime instructa, costa supra subtusque aegre visibili, nervis lateralibus utrinque obscuris; petiolus crassus, rugosus, glaber, 3-5 mm. longus. Inflorescentia lateralis, pedunculata; pedunculus gracilis, 20-25 mm. longus, pills deflexo-patentibus copiose indutus; pedicelli graciles, circiter 9 mm. longi, indumento ut illud pedunculi. Alabastrum apice in cornu circiter 4 mm. longum productum. Calyx alte 5-fidus, extus parce pilosus; lobi ovati, longe acute acuminati, apice acuti, 2 mm. longi, basi 0.75 mm. lati. Corolla rotate, subtus glabra, supra marginem versus pills mollibus longiusculis praedita, esplanata circiter 15 mm. diametro; tubus circiter 2 mm. longus; lobi late triangulares, apice in caudam 5 mm. longam producti, cauda exclusa circiter 1.5 mm. longi. Coronae segmenta radiato-patentia, elliptica, circiter 2.25 mm. longa, supra plana, subtus convexa, angulo inferno in linguam brevem acutam erectam producto, angulo externo subacuto. Antherae appendix membranacea, apice in processum anguste linearem circiter 5 mm. longum producta; corpusculum parvum, ellipticum. Carpella circiter 1.5 mm. alta; stigmatis caput umbonatum, leviter bifidum. Folliculus teres, glaber, leviter striatus, siccitate pallide cinereus maculis fuscopurpureis variatus, apice sensim attenuatus, circiter 7.5 cm. longus, medio 3.5 mm. diametro; semina compressa, circiter 5 mm. longa, 0.75 mm. lata, apice comosa. Thailand (Siam). Terutao, edge of mangrove forest, Kerr 14164. This species is known only from the one, rather scanty collection. It is closely allied to H. caudata Hook. f., found further to the south, in Malacca and Penang. Terutao, the provenance of H. flagellata, is an island on the coast of Satul.—A. F. G. Kerr. Translation: new species, allied to Hoya caudata Hooker f., leaves and flowers small, lobes of the corolla distinguished by being long caudate in proportion. A somewhat woody epiphyte; branches slender, glabrous. Leaves ovate or lanceolate, base rounded and minutely eared, with the apex acute, margine more or less revolute, 4.7 - 8.5 cm. long, 2 - 2.5 cm. wide, leathery, when dry pale brown, glabrous except that above it is sometimes provided with minute sparse hairs, midrib scarcely visible above, lateral nerves obscure on both sides; petioles thick, wrinkled, glabrous, 3-5 mm. long. Inflorescence lateral, pedunculate; peduncles slender, 20-25 mm. long, clothed with copious flatly deflexed hairs; pedicels slender about 9 mm. long, with indumentum same as the peduncles. Flower bud with a apical spur 4 mm. long. Calyx deeply 5 parts, outside sparsely pilose (felt-like); lobes ovate, long acute acuminate, apex acute, 2 mm. long. base 0.75 mm wide. Corolla rotate, glabrous below, above at the margines with long soft pliable hairs flattened about 15 mm. in diameter, tube about 2 mm. long; lobes broadly triangular with a tail like appendage 5 mm. long, exclusive of the tail 1.5 mm. long. With the segments of the corona radiating flatly, elliptic, about 2.25 mm. long. flat above, convex below, with the interior angle tongue like briefly acute produced erect, exterior angle somewhat acute. With the anther appendix membranaceous, apex of the process narrowly linear about 5 mm. long; corpusculum small elliptic. Carpels about 1.5 mm. tall; stigma capitate umbonate, slightly bifid. Follicles round, glabrous, slightly striate, when dry pale ash gray colored with variable spots of brownish purple, apex gradually attenuate, about 1.5 cm. long, in the middle 3.5 cm. in diameter; seeds

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compressed, about 5 mm. long, 0.75 mm. wide, apex comose. In Florae Siamensis Enumeratio (1951) 36. W. G. Craib & A. F. G. Kerr. Hoya flagellata Kerr in Hook. Ic. Pl. xxxv. t. 3407 (1940). Puket. Terutao, edge of mangrove forest, Kerr 14164 (type!). This interesting species comes close to Hoya caudata Hook. f., but it has smaller flowers and the corona lobes have relatively longer tails. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 88. “The Genus Hoya in Thailand” O. Thaithong. 10. H. flagellata Kerr, Hook. lc. (1840) Pl. XXXV, t. 3407. Fl. Siam. Enum. 3(1) (1951) 36. Occurrence: Satun, Trang. Notes: Corona purplish red throughout.

Hoya flavescens Schlechter In Botanische Jahrbücher 50 (1913) 112-113. R. Schlechter. 11. H. flavescens Schltr. n. sp. — Suffrutex epiphyticus, ramosus scandens. Rami filiformes, flexuosi, laxe foliati, teretis, glabri. Folia patentia vel patula, oblonga vel lanceolato-oblonga, acuminata, basi subcuneato-rotundata, carnosula, glabra, petiolo brevi, carnosulo, superne leviter sulcato. Inflorescentiae graciliter pedunculatae, umbelliformes, 10 — 20 florae, pedunculo pedicellisque gracilibus, glabris. Flores in sectione mediocribus, illis H. ischnopus Schltr. similes. Calycis foliola ovata, obtusa, glabra, quam corolla multo breviora. Corolla subrotata, usque infra medium 5-lobata, extus glabra, intus dense et minute papillosa, marginibus intus subvilloso-puberula, ima basi subbarbellata, lobis ovatis, obtusiuscule acuminatis, marginibus subrecurvulis. Coronae foliola horizontalia, patentia, superne elliptico-oblonga, antheris paululo breviora. Pollinia oblique oblongoidea, translatoribus perbrevibus, retinaculo rhomboideo, quam pollinia fere 3-plo minore. Ein zierlicher Schlinger mit schnurartigen, locker beblätterten Zweigen. Blätter 10 — 16 cm lang, in der Mitte 3.7 — 4.3 cm breit. Blattstiele 0.5 — 1.5 cm lang. Blütenstände doldig, 10 — 20 blütig, auf 5.5 — 6.5 cm langen Stielen. Blütenstiel sehr fein, kahl, bis 1.7 cm lang. Kelchzipfel sehr klein, ca. 1.5 mm lang. Korolla ca. 1.6 cm im Durchmesser. Koronaschuppen von der Spitze bis zum äuszeren Rande knapp, 0.3 cm long. Nordöstl. Neu-Guinea: auf Bäumen in den Wäldern des Kani-Gebirges, ca. 800 m ü M. (Schlechter n. 17623. — Blühend im April 1908) Durch die am Rande dicht behaarten Korollaabschnitte ähnelt die Art der H. marginata Schltr., doch hat sie viel schmälere und kleinere Blätter, innen dicht papillöse Blüten mit bedeutend breiteren Koronaschuppen. Translation: See text below. In Hoyas of Northeastern New Guinea (1992) 31-32. R. D. Kloppenburg. (Translation of Die Asclepiadaceen von Deutsch Neu Guinea). 11. Hoya flavescens Schlechter. n. sp.. - An epiphytic half shrub, branched, twining. Branches threadlike, flexible, loosely leaved, round, glabrous. Leaves spreading or outspread, oblong or lanceolate-oblong acuminate., base somewhat cuneate rounded, fleshy, glabrous, petiole short, somewhat fleshy above lightly grooved. Inflorescences slenderly pedunculate, umbellate, 10-20

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flowered, peduncle and pedicels slender, glabrous. Flowers in the section midsize, similar to H. ischnopus Schlechter. Leaflets (lobes) of the calyx ovate, obtuse, glabrous, much shorter than from the corolla. Corolla almost ovate, 5 lobed divided to below the middle, outside glabrous, inside densely and minutely papilose, with the margins inside somewhat villose-puberulous, at the base somewhat tufted, lobes ovate, somewhat obtusely acuminate, margins slightly recurved. Scales of the corona horizontal, spreading, above elliptic-oblong, with the apex obtusely acuminate, outside obtuse, sides thickly rounded, anthers a little shorter. Pollinia obliquely oblong, with the translators very short, retinaculum rhomboid, three times shorter than pollinia. A pretty clinging vine with cord like, loosely leaved branches, leaves 10-16 cm. long, in the middle 3.7-4.3 cm. wide. Petioles 0.5-1.5 cm. long. Inflorescences umbellate, 10-20 flowered, with peduncles 5.5-6.5 cm. long. Pedicels very fine, bare to 1.7 cm. long. Calyx lobes very small, about 1.5 mm. long. Corolla about 1.6 cm. in diameter. Corona scales from the apex to the outer end, barely 0.3 cm. long. Northeastern New Guinea: on trees in the forest of the Kani mountains, about 800 m. altitude. (Schlechter #17623 -- blooming in April 1908) Through the thick hairs on the margins of the corolla lobes, it is related to the species H. marginata Schltr., but it has much narrower and with smaller leaves, the flowers on the inside are papillose an the inside and with considerable wider corona scales. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. flavescens Schlechter (Schlechter 17623) – 15. In Dr. Schlechter’s Hoya Species (1993) 64-65. R. D. Kloppenburg. 11. Hoya flavescens Schlechter. n. sp. An epiphytic half shrub, branched, twining. Branches threadlike, flexible, loosely leaved, round, glabrous. Leaves spreading or outspread, oblong or lanceolate-oblong acuminate., base somewhat cuneate rounded, fleshy, glabrous, petiole short, somewhat fleshy above lightly grooved. Inflorescences slenderly pedunculate, umbellate, 10-20 flowered, peduncle and pedicels slender, glabrous. Flowers in the section midsize, similar to H. ischnopus Schlechter. Calyx scales ovate, obtuse, glabrous, much smaller than corolla. Corolla almost ovate, 5 lobed divided to below the middle, outside glabrous, inside densely and minutely papillose, margins inside somewhat villose-puberulous, at the base somewhat tufted, lobes ovate, obtusely acuminate, margins slightly recurved. Corona scales horizontal, spreading, above elliptic-oblong, with the apex obtusely acuminate, outside obtuse, sides thickly rounded, anthers a little shorter. Pollinia obliquely oblong, translators very short, retinaculum rhomboid, three times shorter than pollinia. An ornamental vine with cord like, loosely leaved branches, leaves 10-16 cm. long, in the middle 3.7-4.3 cm. wide. Petioles 0.5-1.5 cm. long. Inflorescence umbellate, 10-20 flowers, on peduncles 5.5-6.5 cm. long. Pedicels very fine, bare to 1.7 cm. long. Calyx very small, about 1.5 mm. long. Corolla about 1.6 cm. in diameter. Corona scales from the apex to the outer end, barely 0.3 cm. long. Northeast New Guinea: blooming in the forest of the Kani mountains, about 800 m. altitude. (Schlechter #17623 -- blooming in April 1908) Through the thick hairs on the margins of the corolla lobes, it is related to the species H. marginata Schltr., but it has much narrower and smaller leaves, and blooms that are papillose an the inside and with considerable wider corona scales. Fig. 3

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G-N. G= Branches/w leaf. H= Corolla segments inner surface. J= gynostegium with corona. K L M = Corona scales. N= Pollinarium.

Hoya flavida P.I. Forster et D. J. Liddle In Austrobaileya 4/1 (1993) 53-54. P. I. Forster & D. J. Liddle. Hoya flavida P.I. Forster et D. J. Liddle, sp. nov. affinis H. anulatae Schltr. a qua foliis succulentioribus, corolla campanulatirotata lutes vel flavida lobis ovatis, et lobis coronae staminalis margins exteriors rotundato differt. Typus: cultivated at Emerald Creek near Mareeba (ex plant collected on Mt. Gallego, Guadalcanal Province, Solomon Islands), 24 February 1992, D. J. Liddle IML118 (holo: BRI (1 sheet + spirit)). Epiphytic fleshy vine to several meters long, latex white. Stems cylindrical, glabrous; internodes up to 120 mm long and 2.5 mm diameter. Leaves glabrous, petiolate; lamina elliptic-ovate, up to 140 mm long and 45 rum wide, discolorous, secondary veins 4 per side of midrib just visible on both surfaces, tertiary venation obscure; upper surface dark green; lower surface pale green; tip acute to short acuminate; base cuneate; petiole grooved along top, 10 - 12 mm long and c.1.5 mm diameter; colleters 2, at lamina base. Cyme umbelliform to racemiform, up to 140 rum long; peduncle up to 135 mm long and 1.5 mm diameter, glabrous; bracts ovate, 0.3-0.4 mm long, 0.3- 0.4 mm wide. Flowers 3-4 mm long, 10-15 mm diameter; pedicels 15-22 mm long, c. 1 mm diameter, glabrous. Sepals lanceolate, 1.2-1.4 mm long, 0.5-0.6 mm wide, glabrous. Corolla campanulate-rotate, yellow to yellow-pink, externally glabrous, internally with dense short white trichomes that are uniformly distributed except for on the lobe edges where slightly dense but not forming a conspicuous band; tube 3-4 mm long, 7-8 mm diameter; lobes ovate, 5-5.5 mm long, 4.5-5 mm wide. Staminal corona 2.8-3 mm long, 5.5-5.8 mm diameter, inserted on column flush with corolla, yellow-pink; each lobe c. 2.5 mm long, 2.5-2.8 mm high, 1.3-1.5 rum wide, outer edge somewhat rounded-obtuse and not upraised above the level of the style-head, inner edge acute. Staminal column c. 2 mm long and 2.5 mm diameter; anther appendages lanceolate, c. 0.8 rum long and 0.5 mm wide; alar fissure c. 1 mm long. Style-head c. 1 mm diameter. Pollinaria c. 0.32-0.36 mm long, 0.27-0.32 mm wide; pollinia oblong, 0.29-0.33 mm long, 0.12-0.13 mm wide, with pellucid germination mouth on outer edge; corpusculum oblong, 0.09-0.11 mm long, 0.05-0.06 mm wide; caudicles winged, c. 0.09 mm long and 0.05 mm wide. Fruit and seed not seen. Fig. 2. Specimens examined: Papua New Guinea. Bougainville Province: South slopes of Crown Prince Range, Apr. 1936, Voyce D2 (K). Cultivated. cultivated at Emerald Creek near Mareeba (ex plant collected on Mt. Gallego, Guadalcanal Province, Solomon Islands). Feb. 1992, Liddle IML423 (BRI). Distribution and habitat: Known only from Bougainville and Guadalcanal Islands in the Solomon Islands chain. Plants occur as epiphytes in lowland rainforest. In addition to the plants in cultivation from Mt. Gallego, we have also collected plants from the Lungga River area on Guadalcanal that are probably referable to this species; however, they have yet to flower for us to be sure. Notes: H. flavida is a free flowering plant that is now reasonably well spread in cultivation as H. sp. 'Mt. Gallego'. It is probably allied to H. anulata Schltr. from mainland New Guinea and Australia (Forster & Liddle 1990, 1992) but differs from that

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species in the more succulent leaves, the yellow or yellowish campanulate-rotate corolla with ovate lobes; and the staminal corona lobes with rounded outer edges. Etymology: The specific epithet alludes to the yellowish corollas. Acknowledgments: Figures 1 and 2 were drawn by W. Smith (BRI). Our visit to Papua New Guinea, enabling a visit to the LAE herbarium, was made possible by the award of a Christensen Research Institute Fellowship and we are grateful to M. Jebb, Director of that institute for his support. G. Stocker was of great assistance with our visit to Lae. L. A. Craven (CANB) kindly provided Latin translations of the diagnoses. Plants or information on localities were supplied by G. Dennis, B. Ghen and the late P. Tsang. The Directors/Curators of A, B. BM, BO, BSIP, CANB, K, L, LAB, MICH, SING, NY, W and WRSL allowed access to collections in their care, either on loan or on visits to their institutions. References: Forster, P. l. & Liddle, D. J. (1990). Hoya R. Br. (Asclepiadaceae) in Australia - an alternative classification. Austrobaileya 3: 217-234. — (1991). Variation in Hoya australis R. Br. ex Traill (Asclepiadaceae). Austrobaileya 3: 217-234.. — (1992). Taxonomic studies on the genus Hoya R. Br. (Asclepiadaceae) in Papuasia, 15 Austrobaileya 3: 627- 641. Translation: like Hoya anulata Schlechter from which it differs by its more succulent leaves, campanulate corolla pale yellow or yellow ovate lobes, and with the staminal coronal lobe exterior margins rounded. In Hoya Section Acanthostemma (Blume) Kloppenburg (1994) 43-44. R. D. Kloppenburg. Hoya flavida P. I. Forster et D. J. Liddle, sp. nov. affinis H. anulata Schltr. a qua foliis succulentioribus, corolla campanulatata lutea vel flavida lobis ovatis, et lobis coronae staminalis margine exteriore rotundato differt. Typus: cultivated at Emerald Creek near Mareeba (ex plant collected on Mt. Gallego, Guadalcanal Province, Solomon Islands, 24 February 1992, D. J. Liddle IML 118 (holo: BRI ( 1 sheet + spirit)). Epiphytic fleshy vine to several meters long, latex white. Stems cylindrical, glabrous; internodes up to 120 mm long and 2.5 mm diameter. Leaves glabrous, petiolate; lamina elliptic-ovate, up to 140 mm long and 45 mm wide, disclorous, secondary veins 4 per side of midrib just visible on both surfaces, tertiary venation obscure; upper surface dark green; lower surface pale green; tip acute to short acuminate; base cuneate; petiole grooved along top, 10-12 mm long and c. 1,5 mm diameter; colleters 2, at lamina base. Cyme umbelliform to racemiform, up to 140 mm long; peduncle up to 135 mm long and 1.5 mm diameter, glabrous bracts ovate, 0.3-0.4 mm long, 0.3-0.4 mm wide. Flowers 3-4 mm long, 10-15 mm diameter; pedicels 15-22 mm long, c. 1 mm diameter, glabrous. Sepals lanceolate, 1.2-1.4 mm long, 0.5-0.6 mm wide, glabrous. Corolla campanulaterotate, yellow to yellow -pink externally glabrous, internally with dense short white trichomes that are uniformly distributed except for on the edges where slightly denser but not forming a conspicuous band; tube 3-4 mm long, 7-8 mm diameter; lobes ovate, 5-5.5 mm diameter, inserted on column  flush with corolla, yellow pink; each lobe c. 2.5 mm long, 2.5-2.8 mm high, 1.3-1.5 mm wide, outer edge somewhat rounded-obtuse and not upraised above the level of the stylar-head, inner edge acute. Staminal column c. 2 mm

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long and 2.5 mm diameter; anther appendages lanceolate, c. .8 mm long and 0.5 mm wide; alar fissure c. 1 mm long Style-head c. 1 mm diameter. Pollinaria c. 0.32-0.36 mm long, 0.27-0.32 mm wide, with pellucid germination mouth on outer edge; corpusculum oblong, 0.09-0.11 mm long and 0.05 mm wide. Fruit and seed not seen. Herbarium Sheets: Bougainville Prov. S. slopes of Crown Prince Range, Apr. 1936, Voyce #D2 (K); Cultivated Emerald Creek ex Mt. Gallego, Guadalcanal, Solomon Is., IML 423 (BRI).

Hoya flexuosa Spreng. In Sprengel System Vegetabilum 1 (1825) 843. Spreng. flexuosa * 5. H. foliis cordatis oblongis venosis, pedunculo flexuoso, corolla imberbi. Nov. Holl. Translation: Hoya leaves cordate oblong veined, peduncles flexible, corolla beardless. from New Holland. In General System of Gardening 4 (1837) 128. G. Don. 3. T. flexuosa (r. Br. l. c.) umbels alternate, sessile; common peduncles flexuous; leaves cordate-oblong, veiny; corolla beardless. (woody perennial. F.) Native of New Holland, within the tropic. Hoya flexuosa, Spring. syst. I. P. 843. Flexuous Tylophora. Shrub tw. In Flora Australiensis 4 (1867) 336. Bentham. 9. T. flexuosa, R. Br. Prod. 460. A rather slender twiner, glabrous or the inflorescence pubescent. Leaves ovate-lanceolate, acutely acuminate cordate and broad, the larger ones 2 to 3 in. long or even more, the upper ones small and narrow. Flowers very small, in little clusters or sessile umbels along a slender flexuose peduncle, often longer than the leaves. Pedicel filiform, 1 to 2 lines long. Calyxsegments about 3/4 line long. Corolla scarcely 1 1/2 lines diameter. Corona-segments rather thick, but not very prominent, acuminate, the adnate points reaching to about half the length of the anthers. — Dcne. In DC Prod. Viii. 612; Hoya flexuosa, Spreng. Syst. I. 843. In Australia. Upper Victoria, F. Mueller; islands of the Gulf of Carpentria. In Queensland Flora (1900) 1006. F. M. Bailey. T. flexuosa (flexuose), R. Br. Prod. 460; Benth, Fl. Austr. iv. 336. A rather slender twiner, glabrous or the inflorescences pubescent. Leaves ovate-lanceolate to linear-lanceolate, acutely acuminate, cordate when broad, the larger ones 2 to 3 in. long or even more, the upper ones small and narrow. Flowers very small, in little clusters or sessile umbels along a slender flexuose peduncle, often longer than the leaves. Pedicels filiform, 1 to 2 lines long. Calyx-segments about ¾ line long. Corolla scarcely 1 ½ line diameter. Corona-segments rather thick, but not very prominent, acuminate, the adnate points reaching to about half the length of the anthers. — Decne. in Prod. viii. 612; Hoya flexuosa, Spreng. Syst. i. 843. Hab.: Islands of the Gulf of Carpentaria, R. Brown.

Hoya forbesii King & Gamble In Flora of the Malay Peninsula (1903) 574. King & Gamble. 16. Hoya forbesii, King & Gamble, n. sp. A climbing shrub, rooting on trees and probably epiphytic; branchlets somewhat angular, lenticellate, with pale brown shining bark. leaves coriaceous when

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dry; elliptic-oblong or obovate, shortly and bluntly acuminate at apex, rounded at bass; upper surface glabrous, shining; lower surface minutely papillose; both pale brown when dry; margines somewhat recurved; 4 to 7 in. long, 2.5 to 4 in. broad; midrib strong, much raised beneath, bearing on the upper surface a gland just above the petiole; main nerves about 7 pairs, irregular but usually at 40° with the midrib, more or less curved and branching; secondary nerves frequent, especially near the top between the main nerves; reticulations transverse, irregular, clearly visible only on the upper surfaces; petiole very thick, bent, articulate in the middle, about 5 in. long. Umbels apparently few-flowered, on 75 in. long rachises with small tubercles a the end of short (about .75 in.) peduncles; pedicels filiform, 5 in. long, puberulous; buds flattened, .25 in. in diam.; flowers having grayish-white corollas with purple tips (Fortes). Calyx membranous; lobes about twice as long as tube, ovate, ciliate, puberulous without; no scales Corolla .4 in. in diam., puberulous without; lobes broadly triangular-ovate-acute, incurved. Corona of 5 inflated processes closely adnate to too staminal-column; lower lobes horizontal, narrow-oblong, slightly ridged above, 2-valved below, nearly 1 in. long; upper lobe a minute erect tooth. Staminal-column short; anthers incumbent over the style-apex; cells divergent; appendages acute, scabrous; pollen-masses flattened, falcate, attached by short straight caudicles to the narrow pollen-carriers. Style-apex very thin, deeply 5-lobed with a conical tip. Fruit not known. Perak: Scortechini 1679, 1680.—Distrib. Sumatra, on Kaba Volcano 5000 feet (Forbes 2896A). In the accompanying key: Flowers small, .25 in. in diam. in bud:- Leaves large, usually over 5 in. in length and 1.5 in. in breadth; main nerves very prominent:— Main nerves at 40° with the midrib; reticulations prominent, chiefly on the upper surface , but not sharp. In The Flora of the Malay Peninsula 2 (1923). H. Ridley. (9) H. Forbesii King & Gamble, l.c. 574. Branches thick, slightly angled. Leaves coriaceous, elliptic-oblong or obovate acuminate blunt, base narrowed rounded, stout, midrib and nerves 7 pairs, elevated when dry, 4 to 7 in. long. 2.5 to 4 in. wide; petiole thick. Peduncle .75 in long, rachis short; pedicels .5 in long. Flowers .4 in. across, grayish white. Corolla purpletipped. lobes triangular acute incurved, puberulous outside. Corolla lower lobes narrowoblong. Hab. Perak (Scortechini). Rare. Distrib. Sumatra. In Malay Nature Journal 30(¾) (1978) 517-518. R. L. Rintz. 23) Hoya forbesii King & Gamble, J.A.S. Beng. IV (1903) 574. Type: Sumatra, Gunong Kaba at 1500m, Forbes 2896a (K). ---Fig. 28. Distinguishing Features: Leaves thick rigid, broadly elliptic with cuneate bases, the lower surface finely papillose; up to 16cm long by 6.5cm wide. Peduncle horizontal, rigid, 23cm long. Umbel negatively-geotropic, convex with rigid, uniform pedicels c. 2cm long; 1-20 flowers, open c. 24hrs and falling after 2 days. Corolla spreading, finely and sparsely pubescent inside and out; c. 1.5 cm diam; pale green or yellow. Corona entirely white. Corpuscle narrow. Ecology: very rare; known only from Perak and Batu caves (the interior canyon and the eastern summit), Selangor. Distribution: S. Sumatra.

Hoya formosana Yamazaki

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In The Journal of Japanese Botany 43 (1968) 223. Takasi Yamazaki. 9. Hoya formosana (Chinese Script). Hoya formosana Yamazaki, sp. nov. Caulis scandens glaber 4 mm diameter. Folia carnosa late ovata 7-9 cm longa 4-5 cm lata apice abrupte attenuata basi rotundata in siccate pallida supra glabra basi ad costas sparse scabrida subtus glabra ad nervis scabrido-punctata, nervis primariis paucis 2-3 jugis conspicuis arcuatis, petiolis crassis 2-3cm longis fauce superiore surcatorum scabrido-punctatis ceteris glabris. Inflorescentia axillaris umbellata folio subaequilonga 5 cm diameter 25-40 florifera, pedunculis 3-3.5 cm longis glabris, pedicellis 2-2.5 cm longis fusco-pubescentibus. Bracteae lineari-oblongae 2-2.5 mm longae apice rotundatae margine puberulae. Calyx campanulatus 5 partitus late ovatis-obtusis extus sparse fusco-pilosis margine punctatopilosis 2 mm longis 1.3 mm latis Corolla rotata 1.2 cm diameter utrinque glabra 5 fissa, petalis late petalis late ovatis, laciniis ovatis apice obtusis vel rotundatis margine sparsissime punctatis 5mm longis et latis. Corona stellate patens crassa 6 mm diameter, lobis ovalibus 2 mm longis 1.2 mm latis posterioribus obtusis anterioribus in dentem obtusem antherae incumbentem productis, appendicibus antherarum membranaceis rotundatis. pollinia erecta oblonga 0.3 mm longa, caudicibus brevissimis, retinaculo oblongo apice obtuso 0.15 mm longo. Pistillum conicum quinquangularis glabrum. Hab. Formosa. Hengchun (... ) (Yanagawa, typus in TI). Translation: Stems climbing glabrous 4mm in diameter. Leaves fleshy broadly ovate 79cm long 4-5cm wide apex abruptly attenuate base rounded when dry flat very glabrous base near the midrib sparsely rough below glabrous with nerves roughly punctate, primary nerves few 2-3 pairs conspicuously curved, petioles thick 2-3cm long above the beginning of the young shoot roughly punctate otherwise glabrous. Inflorescences axillary umbels somewhat the same length as the leaves 5cm in diameter with 25-40 flowers, peduncles 3-3.5cm long glabrous, pedicels 2-2.5cm long brownish pubescence. bracts linear oblong 2-2.5mm long apex rounded margine puberulous. Calyx cup-shaped of 5 parte broadly ovate obtuse or rounded margines punctate-hairy 2mm long 1.3mm wide. Corolla rotate 1.2cm in diameter both surfaces glabrous of 5 parts, petals broadly ovate apex obtuse or rounded margines sparsely punctate 5mm long and wide. Corona star-like flat thick 6mm in diameter, lobes oval 2 mm long 1.2mm wide posterior obtuse anterior tooth-like obtuse anther incumbent, appendix of the anther membranaceous rounded. Pollinia erect oblong 0.3 mm long, caudicles short, retinaculum oblong with an obtuse apex 0.15mm long. Pistil conic of 5 angles glabrous. Habitat in Formosa. In Flora of Taiwan (Vascular Plants) 6 (1973) 114. 122. Asclepiadaceae. H. carnosa (lf) R. Br. H. formosana Yamazaki in Jour. Jap. Bot. 43:223 (1968) Yang Lit. Pl. Taiwan 673.

Hoya fraterna Blume In Museum Botanicum Lugduno-Batavum 1 (1849) 44. C. L. Blume. 104. Hoya (Physostemma) fraterna Bl: volubilis, glabra; foliis coriaceis ovato-oblongis ellipticisve acuminatis basi rotundatis subvenosis supra petiolum calloso-glandulosis; umbellis longissime pedunculatis multifloris; corollae intus sericeo-velutinae laciniis trianunlari-ovatis acutis; coronae stamineae foliolis supra convexis angulo exteriore

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obtuso recto. Bl. in Rumphia IV.—H. coriaceae maxima affinis, eujus flores majores, segmenta calyces breviore et foliola coronae stamineae angulo exteriore minus obtuse atque subreclinata.— In calcareis Kuripan Javae occidentalis. Translation: twining, glabrous; leaves leathery ovate-oblong elliptic acuminate base rounded somewhat veined above the petiole callous-glandulose; umbels many flowered on long peduncles. corona inside silky-velvety lobes triangular-ovate acute; leaflets of the staminal corona convex above exterior angle erect obtuse. Blume in Rumphia IV — Very near H. coriacea but larger flowers, calyx segments smaller and leaflets of the staminal corona at the exterior angle less obtuse and less bent down. In Botanical Magazine (Curtis’s) (1850) 4684. Hoya (Physostemma) fraterna; foliis amplis ellipticis crassissime coriaceis acutis basi subcordatis et calloso-glandulosis obscure remote penninerviis margine recurvis, petiolo costaque subtus praecipue crassis, pedunculo folio 3-4 plo breviore, umbella multiflora compacta, sepalis ovalibus obtusis concavis, corollas rotatae lobis deltoideis patenti-recurvis sericeo-velutinis, coronae stamineae foliolis brevi-ovatis apice recto obtuso. Hoya fraterna. Blume, Mus. Bot. Lugd. Bat. p. 44. A very fine new and very distinct species of Hoya, first detected in Java by Blume and since by Thomas Lobb, and sent by him to his employer Mr. Veitch, in whose stove at Exeter it has grown very vigorously and yielded its very handsome flowers during a great part of the summer and autumn. Some of the leaves measure a foot in length: our coloured figure is taken from a portion of the plant yielding smaller foliage; but these leaves are remarkable no less for their great size than they are for their firmness and thickness, and the very indistinct remote pinnated nerves, scarcely seen except when the leaf is held between the eye and the light, or when the leaves are dried for the herbarium; then the shrinking of the parenchyma brings the veins more distinctly into view, and slows them to be pinnated, anastomosing, and slender. The petioles and costa beneath are peculiarly thick. The upper side of the corolla, disc excepted, is downy, or between silky and velvety, and of a pale yellowish buff-colour, but five stains or spots are seen radiating from the centre towards the sinuses, which are always wet and clammy, which clamminess appears to be due to a flow of honey from beneath each of the leaves of the crown or nectary, and give a rich brown tone of colour to the whole umbel of flowers. It is named fraterna by Blume on account of its affinity to H. coriacea from which it is however abundantly distinct, as still be seen by a reference to the latter plant figured at our Tab. 4518. Descr. A clincher its terete stems and branches, rooting, near the insertion of the petioles, bearing opposite leases, on rather short but very thick petioles; varying from six inches to a foot in length, singularly thick, and firmly fleshy, subcoriaceous, elliptical, very glabrous and even, the margins recurred, the apex rather acute, the base emarginate or subcordate, dark green and glossy above, pale above opaque beneath, where the mid- rib is very broad and prominent; lateral veins scarcely at all visible except the leaf be held between the eye and the light, wen they are seen to be pinnated, distant, slender, anastomosing, towards the margin. Peduncle much shorter than the leaves, moderately stout, thickened at the base, bearing at the apex a dense umbel of rather large, brownishred flowers. Sepals five, oval, concave. Corolla rotate, pale buff, with five red-brown blotches, five-lobed, the lobes triangular, silky reflexed. Leaflets of the corona pale buff,

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rotundato-ovate, thick fleshy, concave above with a blood-red spot at the base, grooved beneath. Ovaries two, oblong. Translation: leaves large elliptic thickly leathery acute base somewhat cordate and with a callused gland obscure remote penninerved margines recurved, petiole and midrib thick mainly below, peduncle short blade 3-4 in., umbel many flowered compact, sepals oval obtuse. lobes of the corolla deltoid flat-recurved silky-velvety, leaflets of the staminal corona briefly-ovate apex erect obtuse. In Fleur des Serres 8 (1852) 815. J. E. P. Hoya fraterna. (Hoya fraternal.) Charact. Specif. — H. Physostemma foliis amplis ellipticis crassissime coriaceis acutis basi subcordatis et calloso-glandulosis obscure remote penninerviis margine recurvis, petiolo costaque subtus praecipue crassis, pedunculo folio 3-4 plo breviore, umbella multiflora compacta, sepalis ovalibus obtusis concavis, corollas rotatae lobis deltoideis patenti-recurvis sericeo-velutinis, coronae stamineae foliolis brevi-ovatis apice recto obtuso. Hook. Hoya fraterna, Blume. Mus. Bot. Lugd. Batav. I, p.44 (1), fide Hook. in Bot. Mag. t. 4684 (Icon. hic iterata). (1) N’etant pas sur de la justesse de certe determination specifique, nous croyons devoir inserer en note, comme element de comparison, Particle descriptif consaere par Blume a son Hoya fraterna. “Hoya (Physostemma); volubilis, glabra; foliis coriaceis ovato-oblongis ellipticisve acminatis basi rotundatis subvenosis supra petiolum calloso-glandulosis; umbellis longissime pedunculatis multifloris; corollae intus sericeo-velutinae laciniis trianunlari-ovatis acutis; coronae stamineae foliolis supra conrexis angulo exteriore obtuso recto. Bl. in Rumphia IV.—H. coriaceae maxima affinis, eujus flores majorie, segmenta calyces breviore et foliola coronae stamineae angulo exteriore minus obtuse atque subreclinata.” Le nom specifique de cette remarquable Asciepiadee fait allusion a l’affinite tres intime qui l'unit a l'Hoya coriacea, espece deja figuree dans ee recueil (ci-dessus, tab. 578). En supposant nenmoins, ee qui nous semble un peu douteux, que la determination de ces deux plantes soit bien exacte, il suffit d’un simple coup-d'oeil pour les distinguer l'une de l'autre. Sans parler des diversites de couleur florale, de la forme du calice et des pieces de la forme du calice et des pieces de la couronne staminale l’Hoya fraterna serait aisement caracterise par l'ampleur isolile et l'epaisseur de ses feuilles, dont les plus grandes atteignent jusqu’a 30 centimetres de longueur et dont les nervures disparaissent a l’etat frais, dans la profondeur d’un parenhyme ferme et succulent. Du reste, l’hisioire horticole des deux est a peu pres identique. Decouvertes dans les forets de Java par l’illustre botaniste Blume, introduites de ees memes lieux par Thomas Lobb, dans les serres de ses patrons, MM. Veitch, la premiere fleurit d'abord en 1849, la seconde (fraterna), plus tardive, developpa ses belles corolles durant l’ete et l’automne de 1852. Ces corolles, faconnees en etiole a cinq rayons, offrent sur leur face interne, legerement convexe et delicatement veloutee, une teinte buffle clair: cinq macules de meme teinte, mais plus foncees en couleur, s’etendent des einq pointes de la couronne staminale vers les sinus du limbe de la corolle et sont constamment lubrifiees par un suintement de nectar qui se fait a la base de eette couronne. J. E. P. Editors note: Latin translations can be found above. Translation: (French). The specific name of this Asclepiad remarkably makes allusion to

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the very intimate affinity that it has to Hoya coriacea, species already figurative in the compilation (above, tab. 578). while supposing nenmoins, it seems to us a little doubtful, that the determination of these two plants is well exact, it is sufficient a simple stroked'oeil to distinguish them one of the other. San to that the diversities of the floral color, of the shape of the calyx and pieces of the shape of the calyx and pieces of staminal corona of Hoya fraterna would be characterized comfortably by ampleness isolile and the thickness of its leaves, of which the biggest reach up to 30 centimeters of length and whose ribs disappear has the natural state, in the depth of a parenchyma closes and succulent. Of the remainder, the horticultural history of the two is more or less identical. Discovered in the hills of Java by the famous botanist Blume, introduced of from these same places by Thomas Lobb, in greenhouses of his bosses, Messers Veitch, the first blooms in 1849, the second (fraterna), more belated, developed its beautiful corollas lasting the summer and the fall of 1852. These shaped corollas blanched some have five radiuses, offer on their face interns, slightly convex and finely velvety, a tent clear tan: five stain grandma tent, but darker in color, spreading tips of the staminal crown toward sinuses of the limb of the corolla and are lubricated constantly by an oozing of nectar that makes itself the basis of whole crowns. J. E. P. In Tuinbouw Flora 1 (1853) 69. W. H. DeVriese. 6. Hoya fraterna Bl. De Hoya, welke men Aanverwante Hoya zou kunnen noemen, uit hoofde van hare overeenkomst met Hoya coriacea, is beschreven en afgebeeld door Blume (Mus. Bot. Lugd. Bat. I. p. 44 en Rumphia, Tom; IV.) en door Hooker (Bot. Mag. t. 4684, herhaald in de Flore des serres de, l'Europe, door van Houtte, VIII. 179). Hoya fraterna verschilt van H. coriacea vooral door de kleur van de bloemkroon, den vorm van den kelk en van de kroon der meeldraden. De bladen zijn zeer groot, vaak 0.03 lang, en zeer dik en saprijk. Zij werd ontdekt door Blume in de bosschen van Java, overgebragt naar Europa door Thomas Lobb, en bloeide in den zomer en herfst van 1852 bij Vetch, te Exeter. Deze soort is ongetwijfeld eene der schooste Hoya's van onze kassen. Translation: The Hoya, which we could call related (brotherly) Hoya, expressing the similarity to Hoya coriacea, has been described and pictured by (Mus. Bot. Lugd. Bat. I. p. 44 and Rumphia, Tom; IV.) and by Hooker (Bot. Mag. t. 4684, repeated in the Flore on serres de l'Europe, by Van Houtte, VIII. 179). Hoya fraterna mainly differs from H. coriacea by the corolla, shape of the calyx and the staminal corona. The leaves are very large, often 0.03 long, and very fleshy and succulent. It was discovered by Blume in the woods of Java, brought to Europe by Thomas Lobb, and flowered in the summer and autumn of 1852 at Veitch in Exeter. This type is doubtless one of the most beautiful Hoya's in our greenhouses. In Flora van Nederlandsch Indie 1 (1856) 522. F. A. W. Miquel & Zollinger. 19. Hoya fraterna Bl. Volubilis radicans glabra, folia coriacea, e basi rotundata ovato-oblonga acuminata, supra petiolum calloso-glandulosa, subtus utrinque 4—5 costulis patule erectis extrorsum venosis pertensa, 4½—5 poll. longa; umbellae longiter pedunculatae multiflorae, corollae flavescenti-fuscae (rubello-carneae) intus sericeo-velutinae laciniae triaugulari-ovatae acutae, coronae stamineae phylla supra convexa, angulo exteriore obtuso recto. Hoya fraterna Blume Mus. bot. I. p. 44. Rumphia IV. p. 31.—Hook. Bot. Mag. tab. 4684. Van Houtte Fl. des Serres VIII. tab. 815. Habitus Hoyae laurifoliae.—

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Java. Translation: Twining rooting glabrous, leaves leathery, with the base rounded ovateoblong acuminate, above a callous gland on the petiole, beneath 4 to 5 veins opening on the outside spread at a 45° angle from both sides of the midrib. 4 ½ — 5 inches long; many flowered umbels with long peduncles, dark yellow corona (reddish-flesh colored) inside silky-velvety leaflets triangular-ovate acute, with the leaflets of the staminal corona convex above, exterior angle obtuse erect. In Annales Systematicae 5 (1858) 505. W. G. Walpers. 2. Hoya (Physostemma) fraterna. — Hook. in bot. mag. tab. 1681. — F. amplis ellipticis crassissime coriae, acutis b. subcordatis et calloso-glandulosis obscure remote penninerviis marg. recurvis, pet, costaque subt. praecipue crassis, pedunculo folio 3 — 4 pol breviore, umbella multifl. compacta , sep. ovalib. obtusis concavis, cor. rotatae lobis deltoideis patentirecurvis sericeo-velutinis, coronae stamineae foliolis brevi-ovatis ap. recto obtuso. H. fraterna Mus. bot. Lugd. Bat. p. 44. Planta ab ill. Blume primo in ins. Java detecta. — In calcareis Kuripan Javae occidentalis. Translation: Leaves large elliptic very thick acute base somewhat cordate and callousglandulose obscurely and remotely penninerved with margines recurved petiole and midrib noticeably thick, peduncle short blade 3 — 4 inches, umbels many flowers. compact, sepals ovate obtuse concave, lobes of the corolla rotate deltoid spreadingrecurved silky-velvety, leaflets of the staminal corona short ovate apex erect obtuse. In The Garden (1880) 354. Besides these there are H. fraterna, the leaves of which are nearly 1 foot in length, the flowers being pale brown red, and the corona yellow. In Handbook of Succulent Plants Herman Jacobsen. Hoya fraterna, Bl. Java ... L. very large and thick, broadly elliptic; fl. rose-red, with yellow corona. In Flower Garden 3 (1884) 639. (Paxton’s). (Ed. notation: almost an exact copy of article in Botanical Magazine). 639. Hoya fraterna. Blume. A hothouse climbing plant Java. Flowers buff-coloured. Belongs to Asclepiads. Introduced by Messers. Veitch. A very fine new and very distinct species of Hoya, first detected in Java by Blume, and since by Mr. Thomas Lobb, and sent by him to his employer Mr. Veitch, in whose stove at Exeter it has grown very vigorously, and yielded its very handsome flowers during a great part of the summer and autumn. Some of the leaves measure a foot in length our coloured figure is taken from a portion of the plant yielding smaller foliage; but these leaves are remarkable no less for their great size than they are for their firmness and thickness, and the very indistinct remote pinnated nerves, scarcely seen except Even the leaf is held between the eye and the light, or when the leaves are dried for the herbarium; then the shrinking of the parenchyma brings the veins more distinctly into view, and shows them to be pinnated, anastomosing, and slender. The petioles and costa beneath are peculiarly thick. The upper side of the corolla, disk excepted, is downy, or between silky and velvety, and of a pale yellowish buff colour, but five stains or spots are seen radiating from the centre towards the sinuses, which are always wet and clammy, which clamminess appears to be due to a flow of honey from beneath each of the leaves of the crown or nectary, and give rich brown tone of colour to the whole umber of flowers. It

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was named fraterna by Blume, on account of its affinity to H. coriacea, from which it is however abundantly distinct. A climber, with terete stems and branches, rooting near the insertion of the petioles, bearing opposite leaves, on rather short but very thick petioles; varying from six inches to foot in length, singularly thick, and firmly fleshy, subcoriaceous, elliptical, very glabrous and even, the margins curved, the apex rather acute, the base emarginate or subcordate, dark green glossy above, pale and opaque beneath, where the midrib is very broad and prominent; lateral veins scarcely at all visible except the leaf be held between the eye and the light, when they are seen to be pinnated, distant, slender; anastomosing towards the margin. Peduncle much shorter than the leaves, moderately stout, thickened at the base, bearing at the apex a dense umber of rather large, brownish red flowers. Sepals five, oval, concave. Corolla rotate, pale buff with five red brown blotches, five-lobed, the lobes triangular, silky, reflexed. Leaflets of the corona pale buff, rotundate-ovate, thick, fleshy, concave above, with a blood-red sport at the base, grooved beneath.— Bot. Mag., t. 4684. In Exkersionsflora von Java “Flora of Java” 3 (1912) 99. S. H. Koorders. (KEY) 9d. Blätter 11—14 cm lang, lederig, eiförmig-oblong, am Grunde abgerundet und oberseits an der Insertion des Blattstieles kallös-drüsig, unterseits mit 4—5 Paar hervoretenden Seitennerven und grober Netzaderung. Dolden vielblütig, langgestielt. Korolle gelblichbraun odor fleischfarben, innen seidensammethaarig. Coonazipfel oben konvex. Windender, z. T. mit Wurzeln kletternder (?) Stauch, habituell an Hoya laurifolia erinnernd. Miq. l. c. 522; Bot. Mag. tab. 4684. West-Java: Im Kalkgebirge bei Kuripan (Blume in Herb. Leiden). Im Buitenzorger Herbar habe ich diese Art nicht gefunden. Das Leidener Original dieser Art halte ich mit Zweifel nur für ein kleinblütiges Exemplar von der in derselben Gegend von Java wachsenden Hoya coriacea Bl. ........................................................ H. fraterna Bl. Translation: In Exkersionsflora from Java flora of Java 3 (1912) 99. S. H. Koorders. (KEY) 9d. Pages 11-14 cm long, leathery, ovate-oblong, at the acuminate and otherwise at the insertion of the leaf-stalk a callused gland, on the undersides with 4-5 pairs of veins on both sides and coarse midrib. Umbel of many flowers, with long peduncles. Corolla yellow-brown or flesh-colored, inside silky-velvety. Corona scales above convex. Twining, z. T. with roots climbing (?) The stem, and habitat remind one of at Hoya laurifolia. Miq. l. c. 522; Offered. Likes. tab. 4684. West-Java: In the lime-mountain with Kuripan (flower in sharp. Ailments). I have not found this type in the Buitenzorg Herbarium. I only consider the Leiden original of this type with doubt as a few bloom copy of the Hoya growing in the same area from Java Hoya coriacea Bl. ................................................................... Hoya Fraterna Bl. In Standard Cyclopedia of Gardening (Horticulture) (1930) 1603. F. M. Bailey. H. fraterna, Blume. Climbing, rooting near intersection of petiole: lvs. 6-12 in. long, very thick, elliptic, glossy above pale beneath: fls. brownish red, in dense umbels; corolla rotate, pale buff with 5 red-brown blotches; lobes of crown round-ovate, concave with blood-red spot at base, Java. B.M. 4684. J.F. 4:385.

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In The Hoya Handbook (1992) 71. D. Kloppenburg & A. Wayman. Hoya fraterna Blume. Fraterna is very distinctive, having one of the longest leaves in the Hoya genus. Not only is the foliage large, it is also thick and very rigid, with almost no venation to be seen. It is a strong grower and not a plant for limited areas. The photocopied leaf is typical but leaves 12" long are not uncommon. The reflexed fuzzy flowers (described as being between silky and velvety) are typical of species of this group, which includes among others Hoya kerrii, Hoya obovata, Hoya meliflua, and Hoya diversifolia. All of this group have a compact central crown in which the outer lobes are rounded off. In addition all these species are strong vigorous plants with thick bold fleshy leaves. This species like the others mentioned above are very easy to grow and rapidly grow into large plants. They all exudes a honeydew as the flowers age, which often stains the petals. This staining is visible in the colored photograph. As a word of caution I would advise you not to hang flowering plants above valuables where the honey-dew may drip on them. Fraterna is a native of the Java forests. This is one species that I find difficult to bring into flower. Its rank growth, long internodes and infrequent flowering preclude its inclusion in many collections. If you have the room and would like to include a species with the longest leaves, this will be a must for you. The flowers are very similar to H. obovata but are even larger. In Dictionary of the Royal Horticultural Society. H. fraterna. Stem terete, l. elliptical, 612 in. long, thick. leathery, fleshy, glabrous, shining dark green above, margins recurved, apex acute, base sub-cordate, midrib broad, prominent. fl. brownish-red, in dense umbels; peduncles thickened at base, shorter than l. Java. (B.M. 4684.) In Fraterna 2nd Quarter (1993) 11 R. D. Kloppenburg. Hoya fraterna Blume is described in "Annales Museum Botanicum Lugduno- Batavum" V.I p. 44 ii 1849. This is 24 years after H. coriaceae Blume was described. The name of this species is mentioned in the above text as noted. Blume placed this species in the Section Physostemma along with H. coriaceae Blume; H. vitellina Blume; H. polystachya Blume and H. clandestina Blume. The text under #104 reads as follows: 104. Hoya (Physostemma) fraterna Bl.: volubilis, glabra; foliis coriaceis ovatooblongis ellipticisve acuminatis basi rotundatis subvenosis supra petiolum callosoglandulosis; umbellis longissime pedunculatis multifloris; corollae intus sericeo-velutinae laciniis triangulari- ovatis acutis; coronae stamineae foliolis supra convexis angulo exteriore obtuso recto. Bl. in Rumphia IV. - H. coriaceae maxime affinis, cujus flores majores, segmenta calycis breviora et foliola coronae stamineae angulo exteriore minus obtusa atque subreclinata. - In calcareis Kuripan Javae occidentalis. Translation: 104. Hoya (Physostemma) fraterna Blume: twining, glabrous, leaves leathery ovate-oblong elliptical acuminate with the base rounded somewhat veined, above the petiole a little glandular callous; with long peduncled many flowered umbels, with the corona inside silky-velvety with lobes triangular ovate acute; with the leaflets of the staminal corona convex above, the exterior angle obtuse erect. Blume in Rumphia IV - very close to Hoya coriaceae whose flowers are larger, with segments of the calyx smaller and leaflets of the staminal corona outer lobe (angle) less obtuse and somewhat

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reclined (turned downward). In calcareous soil at Kuripan Western Java. In The Hoya Handbook, 2nd Book R. D. Kloppenburg. Hoya fraterna Blume. Fraterna is very distinctive, having one of the longest leaves in the Hoya genus. Not only is the foliage large, it is also thick and very rigid, with almost no venation to be seen. It is a strong grower and not a plant for limited areas. The photocopied leaf is typical but leaves 12" long are not uncommon. The reflexed fuzzy flowers (described as being between silky and velvety) are typical of species of this group, which includes among others Hoya kerrii, Hoya obovata, Hoya meliflua, and Hoya diversifolia. All of this group have a compact central crown in which the outer lobes are rounded off. In addition all these species are strong vigorous plants with thick bold fleshy leaves. This species like the others mentioned above are very easy to grow and rapidly grow into large plants. They all exudes a honeydew as the flowers age, which often stains the petals. This staining is visible in the colored photograph. As a word of caution I would advise you not to hang flowering plants above valuables where the honey-dew may drip on them. This is one species that I find difficult to bring into flower. Its rank growth, long internodes and infrequent flowering preclude its inclusion in many collections. If you have the room and would like to include a species with the longest leaves, this will be a must for you. The flowers are very similar to H. obovata but are even larger.

Hoya fungii Merrill In Lingnan Science Journal 13 (1934) 68-9. E. D. Merrill. Hoya fungii sp. nov. Frutex scandens ramis ramulisque teretibus, glabris, internodis elongatis; foliis coriaceis, penninerviis, ellipticus ad oblongis, 8 ad 14 cm. longis, 4.5 ad 8 cm. latis, sicco pallidis, opacis, utrinque glabris concoloribusque vel subtus paullo pallidiorbus, apice acutis ad breviter acuminatis, basi rotundatis, costa crassa, subtus valde elevata, nervis lateralibus utrinque circiter 7, haud perspicuis, arcuato-anastomosantibus, costam angulo circiter 30° relinquentibus, reticulis laxis, obscuris; petiolo 1.5 ad 3 cm. longo, glabro; pedunculo glabro, saltem 4 cm. longo, sursum incrassato; cylindrico, cicatricibus multis instructo; floribus albidis, numerosis, umbellatis, circiter 1.5 cm. diametro, pedicellis gracilibus, 3 ad 4 cm. longis; leviter consperse ciliatis; sepalis oblongis, acutis, circiter 2.5 mm. longis, extus leviter consperse, subadpresse, subferrugineo-pubescentibus; corolla extus glabra, intus dense pallide furfuracea, lobis rotundatis, circiter 5 mm. longis et 4 mm. latis; corona stellata, glabra, nitida, lobis oblongo-lanceolatis, acuminatis, circiter 3.5 mm. longis. Hainan, Chim Shan, Fan Maan Ts'uen, Ling-shui District; H. Fung 20137 (type), May 3-20, 1932, scandent, 6 mm. long, flowers white, fragrant; Po-ting, S. P. Ko 5223, April 26, 1932; Seven Finger Mountains, H. Y. Liang 61746 May 3, 1932, climbing on trees, flowers cream, white or pinkish. This falls in the group with group with Hoya bonii Cost. of Indo-China, but has slightly larger, white or pale, not violet flowers, while the corolla lobes not at all triangular but are broadly rounded. Translation: Climbing shrub, branches rebranching, round, glabrous, long internodes, leaves leathery, penninerved, elliptic to oblong, 8 to 14 cm. long, 4.5 to 8 cm. wide, when dry pale, opaque, both sides glabris uniform color or below a little paler, apexes acute and

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shortly acuminate, base rounded, midrib thick, below strongly elevated, lateral nerves on both sides about 7, not at all evident, arched meeting at the margins, about 30° angle with the midrib, reticulations loose, obscure; petiole 1.5 to 3 cm. long, glabrous; peduncle glabrous; at least 4 cm. long, sparsely thickened, cylindrical, with many scars; flowers dull white, numerous, umbellate, about 1.5 cm. in diameter, pedicels somewhat slender, 3 to 4 cm. long, with cilia lightly dispersed; sepals oblong, acute, about 2.5 mm. long, outside lightly dispersed, somewhat depressed, near rusty pubescence; corolla outside glabrous, inside densely pale scurfy lobes rounded, about 5 mm long and 4 mm wide; corona star-shaped, glabrous, shinny, lobes oblong-lanceolate, acuminate, about 3.5 mm. long. In Sunyatsenia 3 (1936) 176. Tsiang. Hoya fungi Merrill in Lingnan Sc. Journal. 13: 68. 1934. Hainan: Lingshui, H. Fung 20137 (isotype in Herb. Lingnan), May 3-20, 1932; Ngai Yuen, C. Wang 34053, Sept. 18, 1933; Lingshui, S. P. Ko 52213m April 26, 1932; Seven Fingers Mt., H. Y. Liang 61746, May 3 1932; Ngai Yuen, F. C. How 70603, April 26, 1933. Kwangsi: Lung Chow, S. P. Ko 55130, 1935. Distribution: Hainan; new to Kwangsi. Very near preceding two species (H. dasyantha & H. elmeri) but the leaves and flowers are distinctly larger. In Flora Kaiwantensis (1943) 267. Genekei Masamune. Hoya fungi Merr., in Lingn.- J. XIII. p. 68 (1934); Tsiang, in Sunyat. III. p. 175 (1936); Tanaka et Odashima, in J.-Trop. X p. 379 (1938) In Flora Republicae Popularis Sinicae (1977) 483. Tomus 63. Hoya fungi Merr. in Lingnan Sci. Jour. 13: 68. 1934; Tsiang in Sunyatsenia 3: 175. 1936: ............, 3: 506. ... 4966. 1974; ......... , 3: 273. 1974. In The Hoya Handbook 1 (1992) 72. D. Kloppenburg & A. Wayman. Hoya Fungi Merrill. The soft appearance of the foliage of this plant along with the dark green veins attract you immediately to this plant. The leaves are large, often twice as large as our photocopy shows, they are covered with soft hairs (pubescence). The green is an unusual shade, I'd say a medium deep emerald green, set off with very dark green venation. The new leaves are bronze green in color, also the stems, especially on new growth are a bronze purple. These maroon like tones add a pleasant addition to an already beautiful plant. This is a cold tolerant plant that comes to us originally from the island of Hainan, off the south coast of China. The island forms the western side of the gulf of Tonkin, opposite Hanoi, Vietnam. If necessary it along with Hoya carnosa types and the Hoya compacts can survive the cold months with less heat than most hoyas. Fifty degree F. would be fine. It will actually take temperatures lower than this, but it will be happy in the cold periods with the fifty degree temperatures. In addition to the beautiful foliage, the most out standing feature of this plant is its globular clusters of 60 flowers. They are much like Hoya carnosa clusters, but with twice as many flowers. I find that normally the individual flowers are slightly larger than carnosa flowers. My flowers are always open a little deeper pink in color than our photo depicts. Colors of flowers will vary with the temperatures under which they develop and other environmental factors. It would be normal for pale pink flowers in hot bright

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areas to fade to a pale color even before they open. This plant is a medium strong grower, so plan on it occupying considerable space as it matures. Curl the long stems around the pot and hanger to keep it confined and to give a more compact form. I would say that given time and space this species will be a must for most growers. It is very easy to grow and easy to bring into bloom If you are familiar with growing Hoya carnosa, this one should be a natural for you. As I have said the flower clusters are larger, its foliage unique and very attractive. Combine these qualities with its cold tolerance and you have a plant that will fit into a wide range of acceptable conditions, and is likely to win your heart.

Hoya fusca Wallich In Plantae Asiaticae Rariores 1 (1930) 68. Wallich. Hoya fusca. Tal. 75. Radicans, ramosissima; foliis oblongis, acuminatis; corolla laevissima, coronae foliolis subtus sulco exaratis. Habitat super rupibus arborumque truncis montium valli Napaliae vicinarum; etiam in Napalia inferiore secus ripas fluminis Rapti. Floret Augusto; fructus profert Octobri. Planta fruticosa, magna, latissime radicans, ramosissima, omnino laevis, omnibus partibus copiose lactescens. Rami longi, cylindrici pennam cygneam, usque ad digitum modicum crassi, plus minusve penduli, epidermide cinerea, pallida obtecti, ad foliorum insertionen parum tumidi, et vestigio annulari ciliarum notati. Folia approximata, opposita, patentissima, valde crassa et coriacea, firma, plana oblonga, acumine angusto, semipollicari terminata, marginibus parallelis convexis, basi obtusa, supra atroviridia, lucida, ad insertionen petioli utrinque corpusculo cinereo, emarcido (loco acervuli soliti ciliarum), subtus pallida, subargentea, costa valde elevata, nervis trans versis, capillaceis, intra marginem arcuatim anastomosantibus; 6—7 pollicaria. Petioli crassi, convexi, supra parum sulcati, semipollicares Flores magni, fusci, nitidi, inodori, valde numerosi, densissime dispositi in umbellam hemisphaericam, lateralem, extra-axillarem, diame trum 3-polllcarem emetientem, erectam, pedunculatam, vix ad tertiam inferiorem folii partem elevatam, subinvolucratam. Pedunculus communis crassus, cylindricus, petiolo dimidio longior; partiales teretes, pollicares, subclavati, exteriores ad basin muniti bracteola ob longa, obtusa, ciliata. Calyx quinquefidus, parvus; laciniae ovatae, obtusae, ciliatae, basi extus gibbosae. Corolla rotata, plana, patentis sima, demum reflexa; laciniae lanceolatae, subacuminatae, marginibus subrevolutae, subtus concaviusculae. Columna pentagona, truncatae pallide fusca, 5-loba; lobi depressi, subtus profunde sulcati, supra concaviusculi, angulo interiore producto in dentem subulatum, longum anthere ovat e, obtusae, membranula cinctae oppositum. Massae Pollinis luteae, tenues, longiusculae, lineari-clavatae, erecto-divergentes, basi acuti apici corpusculi minuti, sulcati, fusci insertae. Stigma 5-angulare, vertice in umbilicum obtusum productum, antheris occultum Folliculus saepius solitarius, cylindricus, laevis, 4-pollicaris; maturus haud observatus. Translation: Spreading branched very little, leaves oblong, acuminate; corolla very smooth, leaflets of the corona grooved and furrowed below. Lives above on rocks in the forest and on the trunks in the mountain valleys in the vicinity of Nepal, furthermore below Nepal along the banks of the Raspti river. Florets narrow; fruits prolifically in October.

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A fruiting plant, large broadly spreading, very little branched altogether smooth, and all parts with copious milk. Stems long cylindric winged, up to 1 inch or less thick, plus or minus pendulous, epidermis grayish, covered with pale whitish. Inserted on the thickened petiole, and traces of the annual marked cilium. Leaves approximately opposite, very outspread, strongly thick and leathery, firm, flat, oblong narrowly acuminate ½ inch terminal, margins parallel convex, base obtuse, above purple veined, shinny on both sides of the petiole insertion a gland withered (at this place a solitary cilia) below flat, somewhat green, midrib very much elevated, various transverse nerves, capillaries near the margins anastomosing, 6 to 7 inches. Flower large, brownish, shinny without smell, very numerous, very dense in hemispherical lateral umbels, beyond the axils, 3inches in diameter, erect, pedunculate, becoming elevated in part 1/3 in the lower part of the leaf. Peduncles commonly thick, cylindrical, petiole somewhat longer, circular, 1 inch, somewhat cut off, exterior at the base with many bracts, oblong obtuse, ciliate. Calyx of 5 parts, small, leaflets ovate, obtuse ciliate, base outside with bumps. Corolla rotate, flat, somewhat shinny, at length reflexed, flaps lanceolate, somewhat acuminate, margins somewhat revolute, below concave. Column pentagonal, truncate pale brown, 5 lobed, lobes depressed, below profoundly grooved, above concave, interior angle produced into a sharp pointed tooth, with long ovate anthers and obtuse enclosing the opposite membrane. Pollinia masses yellow, thin, somewhat long linear a little club shaped, erect divergent, base acute corpuscles minute grooved, brown, inserted. Stigma 5 angled, produced with a obtuse umbilical tip, anthers hidden. Follicle often solitary, cylindric, smooth, 4 inches, did not observe at maturity. In Contributions to the Botany of India (1834) 37. R. Wight & Wallich. 7. H. fusca (Wall.:) volubilis glabra, foliis coriaceis vix carnosis lineari-oblongis acuminatis (5-8 unicas longis 1-1 ½ latis) transversim venosis, pedunculatis brevibus multifloris, corolla glabra segmentis obtusis, cor. st. foliolis ovatis obtusis angulo interiore attenuato erecto stigma apiculatum superante — Wall. ! pl. As. rar. 2. p. 78. t. 175; Asclep. n. 30. — Nepal; Silhet; Wallich. The flowers are rather small, brownish coloured, and very numerous.— (R. W.). Editors Notation: The reference above of Wight’s for Wallich’s Pl. As. rar. is incorrect see this 1830 description above. Translation: twining glabrous, leaves leathery barely fleshy linear-oblong acuminate (5 to 8 inches long 1 to 1 ½ wide) transversely veined, peduncles short umbel-like, corolla glabrous segments obtuse, corona star-like leaflets ovate obtuse with the interior angle erect attenuate stigma apiculate overtopping. In Synopsis Plantarum (1840) 891. D. N. F. Dietrich 7. H. fusca Wall. volubilis glabra, fol. lineari-oblongis acuminatis; pedunc. Multifloris; cor. Glabra laciniis obtusis. In Nepalia. Translation: twining glabrous; leaves leathery linear-oblong acuminate; peduncle multiflowered; corona glabrous, leaflets obtuse. In General System of Gardening and Botany 4 s.23 (1837) 126. G. Don. 7. Hoya fusca (Wall. pl. rar. asiat. 2. p. 78. t. 175.) twining, glabrous; leaves coriaceous, hardly fleshy, linear-oblong, acuminated, transversely veined; peduncles short, many flowered; corolla

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glabrous, with obtuse segments; leaflets of corona ovate, obtuse: with the inner angles erect and attenuated, laying upon the stigma, which is apiculated. Perennial shrub. Native of Nepal and Silhet. The flowers are rather small, brownish coloured, and very numerous, Leaves 5-6 inches 1 or 1 ½ broad. Brown-flowered Hoya. Shrub tw. In Synopsis Plantarum 6 (1840) 891. D. N. F. Dietrich. 7 H. fusca Wall. volubilis glabra; fol. coriaceis lineari-oblongis acuminatis; pedunc. multifloris; cor. glabra laciniis obtusis. In Nepalia. Woody. Translation: twining glabrous, leaves leathery linear-oblong acuminate; multiflowered peduncles; corolla glabris leaflets obtuse. From Nepal, woody shrub. In Prodromus System Veg. 8 (1844) 639. DeCandolle. 36. H. fusca (Wall. pl. as. rar.2, p. 78 t. 175)., volubilis glabra, foliis coriaceis vix carnosis lineari-oblongis v. elliptooblongis basi rotundatis apice longiscule acuminatis transversim venosis, pedunculatis petiolum aequantibus umbellatim multifloris, pedicellis crassis, corolla introsum papillosa, laciniis obtuscullis subreflexis, cor. stam. foliol. ovatis obtusis angulo interiore lanceolato erecto stigma apiculatum superante. Climbing Shrub. in Nepalia, Silhet. Wall. herb. ind. cat. n. 8157, B. Wight cont. 37. (v. s. h. Mus. par.) Editors note: Here again the volume and tab are incorrectly entered, evidently repeating Robert Wight’s mistake. Translation: twining glabrous, leaves leathery barely fleshy linear-oblong or ellipticoblong base rounded apex long acuminate transversely veined, peduncle and petiole the same length umbels many flowered, pedicels thick, corolla on the inside papillose, leaflets a little obtuse somewhat reflexed, leaflets of the staminal corona ovate obtuse interior angle lanceolate erect, stigma apiculate rising above. Climbing shrub from Nepal, Silhet. In Handbook of Indian Flora 2 (1866) 242. R. Wight (W. Drury). (10) H. fusca. (Wall.) Ident. Wall. Pl. As. Rar. II. p. 78. — Dec. prod. VIII. p. 639. Engrav. Wall. l. c. t. 175. Spec. Char. Twining: leaves coriaceous, scarcely fleshy, linear-oblong, acuminate, transversely veined: peduncles short, many-flowered: corolla glabrous with obtuse segments: leaflets of the staminal crown ovate, obtuse, inner angle attenuated, erect, exceeding the apiculated stigma; flowers small, brownish. Silhet. In Flora of British India 4 (1883) 58. J. D. Hooker. 24. H. fusca, Wall. Pl. As. Rar. i. 68, t. 75, and Cat. 8157; glabrous, leaves 6-9 in. linear-oblong acuminate fleshy midrib very thick, peduncles short and pedicels very stout, corolla pubescent within. Wight Contrib. 37; Decne. in DC Prodr. viii. 639. Nepal, Wallich; Sikkim, alt. 1-4000 ft. Khasia Mts. and Pegu. A stout climber. Leaves 1 ½ - 2 ½ in. dia,., sides nearly parallel; nerves numerous, quite horizontal; petiole thick ½ - 1 in. Peduncles sometimes terminal, ½ - 1 in., peduncles long. Sepals broadly ovate, obtuse. Corolla ½ in. diam., yellow-brown, lobes spreading. Coronalprocesses short, very thick, obtuse, concave above, inner angle ending in an erect or recurved spur as long as the anther-tip. Follicles 4-5 in. long, by ½ in. diam., rather thickwalled, straight. Seeds 1/8 in. long.

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In Flore General des Indo China 4 (1912) 137. J. Costantin (M. H. LeComte). H. fusca Wall. Pl. Asiat. Rar., 1, p. 68, tab. 75; Wight Corntr., p. 37; Dcne in DC. Prodr., VIII, p. 639; Hook. f. Fl. Brit. Ind., IV, p. 58. Tige grimpante, vigoureuse, glabre. Feuilles charnues, oblongues ou oblongues-lanclolees, arrondies a la base, acuminees a la pointe, glabres d’ordinaire, ou avec des polls rares en dessous; nervure mediane avec quelques glances a la base, saillante en dessus; nervures seconidaires 8-10 paires, apparentes sur le see, perpencliculaires, confluentes par une nervure marginale; reseau tertiaire parfois visible; limbe long de 7-16 cm. sur 3-5 cm.; petiole glabre ou pubescent, long de 1.5-2 cm. inflorecscence: ombelle tres multiflore, pedonculie, parfois terminale, ailleurs axillaire; pedoncule long de 1.5-3 cm., parfois aile, glabre; pedicelles greles, ailes, longs de 20 mm., fleurs larges de 8-10 mm. epanouies. --- Calice a sepales ovales, obtus. Corolle a lobes jaune brun, etales, pubescents en dedans. Coronule formee de 5 processus courts, en bateau, profonds, obtus, termines a l’angle inteme par un bec assez long, dresse et recourbe, depassant un peu le sommet de l’anthere. Fruit: follicules longs de 10-12 cm. sur 8 mm., droits, a paroi un peu epaisse; graines longues de 4 mm. Cambodge: Mts. Knang-Krepeuh, province de Thepong (Pierre). ** Pas de nervures laterales a la base du limbe; nervures laterales visibles le long de la nervure mediane et obliques ou perpendiculaires par rapport a celle—ci. * Nervures laterales perpendiculaires a la nervure mediane; feuilles charnues (7-1 6 cm. X 3-5 cm.) ………………………………………………………...............15. H. fusca. Translation: Trailing, vigorous, stem glabrous. Fleshy, oblong leaves or oblonglanceolate, rounded at the base, acumindes at the tip, glabrous, or with the rare puberulous underneath; median rib with some glands at the base, prominent in over; secondary veins 8-10 pairs, obvious (visible), perpencliculaires, confluentes by a marginal vein; tertiary network sometimes visible; long limb of 7-16 cm. on 3-5 cm.; petiole glabrous or pubescent, 1.5-2 cm. long. inflorecscence: umbel very multi flowered, peduncle, terminal sometimes, elsewhere axillaire; stalked 1.5-3 cm. long, sometimes winged, glabrous; spindly pedicels, winged, 20 mm. long, large flowers 8-10 mm. bloomed. --- Calyx is oval, obtuse sepals. Corolla has lobes brown yellow, spread out, pubescents inside. Corona formed of 5 short processes, boat-shaped, deep, obtuse, finish with inner angle with a long beak, raise and bent, passing a little the top of the anther. Fruit: long follicles of 10-12 cm. on 8 mm., right, has partition a little thick; long seeds of 4 mm. Cambodia: Mts. Knang-Krepeuh, province of Thepong (Pierre). * * Lateral ribs don't have the basis of the limb; this one has visible lateral ribs with the veins either perpendicular or oblique to the midrib by one report. *Nerves lateral perpendicular to the median nerve; fleshy leaves (7-16 cm. x 3-5 cm.) ....………………………………………………………... ...15. H. fusca. In Symbolae Sinicae 7 (1936) 1001. . H. fusca Wall., Plt. As. rar.. I., 68, t. 75 (1830). NW-Y.; Im wtp. Regenwahle des birm. Mons. im Tale Gümbalo bei Tschamutong am Salwin, Granit, …… 2300 m. v. E., 17. VIII. 1916 (9869). Warscheinlich auch diese in der …… St. am Salwin unter Tjiontson, 1675 m. Translation: In wtp. Forests of Burma. Growing in the valley Gümbalo near Tschamutong at the Salwin, granite, 2300 meters elevation, 17. VIII. 1916 (9869). Probably also these in the St. at the Salwin under Tjiontson, 1675 meters.

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In Sunyatsenia 3 (1936) 172. Tsiang. Hoya fusca Wallich, Pl. As. Rar. 1: 68, t. 75. l831; Wight, Contr. Bot. Ind. 37. 1834; G. Don, Gen. Syst. 4: 126. 1838; Decne. in DC., Prodr. 8:639. 1844; Hook. f., Fl. Brit. Ind.. 4: 58. 1883; Costantin in Lecomte, Fl. Gen. IndoChine 4:137. 1912. Yunnan: Lava bed, W. of Teng-yueh, G. Forest 9082, 1912; Shangpa Hsien, alt.. 2300 m., H. T. Tsai 54300, Sept. 17 1933; Ping-pien Hsien, H. T. Tsai 62175, June 2, 1934; same locality, F. H. Yang. s.n. 1934. Kwangsi: Sup-man-ta. Shan, alt. 3000 ft., R. C. Ching 8308, Oct. 29, 1928; Shang-sze, W. T. Tsang 22375, May 25, Distribution: Sikkim-Himalaya; Indo-Chine; Nipaul; Pegu; new to China. The Kwangsi collection is in fruit having straight follicles to 10.5 cm. long, 5 mm. thick corresponding to the description. Dr. Handel-Mazzetti also collected it in Yunnan under No. 9869, but of this we have only a photograph. This and the following six species have penninerved leaves. Sunyatsenia 4 (1939) 124-125. E. D. Merrill. Hoya fusca Wallich, Pl. As. Rar. 1: 68, t. 75. 1831; Tsiang in Suny. 3: 172. 1936. Yunnan: Ma-Kwan alt. 1800 m., H. T. Tsai 51935, March 1933; Ping-Pien, H. T. Tsai 62175, June 1934; Shang-Pa. alt. 2300 m. H. T. Tsai 54300, Sept. 1933; Dzung-Duei, Cham-Pu-Tung, alt. 2000 m., C. W. Wang 66992, Oct. 1935; Chen-Kang, alt. 2700 m., C. W. Wang 72288, March 1936. Distribution: Sikkim-Himalaya; Indo-China: Nepal; Pegu; China-Yunnan and Kwangsi In Florae Siamensis Enumeratio (1951) 36. W. G. Craib & A. F. G. Kerr. Hoya fusca Wall. Pl. As. Rar. i. 68, t. 75 (1830); F. B. I. iv. 58; F. I. C. iv. 137. Payap. Doi Angka, c. 1480 m., Garrett 869! Distr. Nepaul ! (type), Sikkim ! Burma ! Cambodia ! Laos !. In Flora of the Eastern Himalayas (1971) 109. H. Hara. Hoya fusca Wallich. Pl. Asia. Rar. I: 68, t. 75 (1830) Fl Brit. Ind. 4: 58 (1883) — Tsiang in Sunyat. #: 172 (1936). Yektin – Akasay, 1400--2300m (nov. 30, 1036, fr.); Damthang—Temi, 2200—1800 m (Mar. 25, 1963, fr.). Distr. E. Himalaya (Nepal, Sikkim). Khasia. Burma. Thai, IndoChina, and S. China. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 88. “The Genus Hoya in Thailand” O. Thaithong. 11. H. fusca: Wall., Pl. As. Rar. i. (1830) 68, t. 75. FBI 4 (1883) 58; FIV 4 (1912) 137; Fl. Siam. Enum. 3 (1) (1951) 36. Occurrence: (N): Chiang Mai. (Ne) Phetchabun. (E): Nakhon Ratchasima. Notes: Branches 0.7-1 m long, diffuse from short stem; leaves chartaceous; flower about 1 cm diam.; corolla glabrous, yellow, slightly reflexed; corona purple; inner end of coronal scale not covering stigma. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 71. "The Asclepiadaceae of Central Myanmar" Nyo Maung. 11. Hoya fusca Wall. Pl. Rar. 1 (18320 68. Epiphytic climbing vine, leaves oblong-elliptic, 6.5-12.5 x 1.5-3.5 cm, thick umbel with peduncle 2.5-3.5 cm long; coronal scales short globosely ovoid with concave surfaces. Distribution: Kachin state.

Hoya fuscomarginata N. E. Brown

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In Kew Bulletin 577. (1910) 278. N. E. Brown. Hoya fuscomarginata, N. E. Brown [Asclepiadaceae-Marsdenieae]; affinis H. Pottsii, Traill, sed foliis multo majoribus acutis vel gradatim subacuminatis nec abrupte acuminatis, petiolis crassioribus, corolla ochracea vel flavo-virente et coronae lobis purpureotinctis facile distinguitur. Folia crasse carnosa; lamina 17-22 cm. longa, 7-8 cm. lata, ovato-lanceolata, interdum leviter obliqua, acuta; vel gradatim subacuminata, obtusa vel subcuneata, 3-5 nervia, glabra., viridia., marginibus fuscis; petiolus 2.5-4.5 cm. longus, 7 mm. crassus, cinereus. Umbelli multiflori. Pedunculi 3.5-4.5 cm. longi, 2.5 mm. crassi. Pedicelli graciles, 2 cm. longi. Sepala minuta, 1 mm. longa, ovata, acuta, ut pedicelli et pedunculi parce et minutissime puberula. Corolla rotata, 1.2-1.3 cm. diametro, glabra, ochracea vel flavo-virens, lobis 5 mm. longis et latis late ovatis abrupte breviter acuminatis patentibus apice recurvis. Coronae lobi 3.5 mm. longi, stellato-patentes, ovati. utrinque acuti leviter purpureo-tincti. Origin unknown. It was purchased in Brussels by Mr. F. W. Moore of the Royal Botanic Gardens, Glasnevin, from Mr. Pauwels, who stated that is had been imported with Orchids from British Guiana. But it can scarcely be a native of that country, as no species of Hoya is known to inhabit America. Translation: near Hoya pottsii Trail, but leaves are many times larger acute or gradually somewhat but not abruptly acuminate, petioles somewhat thick, corolla ocher or yellowgreen and lobes of the corona easily distinguished by the purple tint. Leaves thick fleshy; blade 17-22 cm. long, 7-8 cm. wide. ovate-lanceolate, sometimes slightly oblique, acute or gradually somewhat acuminate, obtuse or somewhat wedge-shaped, 3-5 nerves, glabrous, green, margines dark brown; petioles 2.5-4.5 cm. long, 7 mm. thick grayish. Many-flowered umbels. Peduncle 3.5-4.5 cm. long, 2.5 mm. thick. pedicels slender, 2 cm. long. Sepals minute, 1 mm. long, ovate, acute, with pedicels and peduncles sparsely and minutely puberulous. Corolla rotate, 1.2-1.3 cm. in diameter, glabrous, yellowish-brown or yellow-green, lobes 5 mm. long and wide ovate abruptly and shortly acuminate outspread with margins recurved. Lobes of the corona 3.5 mm. long, outspread star-like, ovate, both ends acute, lightly tinged purple. In Cyclopedia of Horticulture (1930) 1613. L. H. Bailey. H. fusco-marginata, N. E. Br. Lvs. fleshy, ovate-lanceolate, 7-9 in. long, tawny on margins: fls. ochre-yellow to yellowish green, in many-fld. umbels on peduncles 2 in. long. Country unknown. In Dictionary of the Royal Horticulture Society (1965). Hoya fusco-marginate. Lvs. ovate-lanceolate, about 8” long, fleshy, margins tawny, fl. pale yellow to yellowishgreen; umbels many-flowered; peduncles about 2” long.

Hoya gigantanganensis Kloppenburg In Fraterna 4th Quarter (1992) 2. Hoya gigantanganensis Kloppenburg sp. nov. Type #36787 PNH collected by Yoshio Kondo & G. Edano 26 March 1957, at Gigantangan, Leyte, Philippines. Section: Acanthostemma. Suffrutex epiphyticus, ramosus, scandens. Rami flexuosi, lax foliate, teretes, glabri, bene foliati, radicanti. Folia patentia vel patula, lanceolata acuminate, basi

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subcuneato-rotundata, glabra, petiolo subbrevi, superne leviter sulcato; pedunculo longissimus, glabris; pedicellis gracilibus, glabris. Flores in section mediocribus. Calycis segments ovate, glabra, margins ciliolata, corolla multi breviora. Corolla revoluta, usque ad teretiam partem basilarem 5-fida, lobis triangularis, acuminatis, extus glabris, intus apice glabrato excepto pubescentis reversis subulatis cristalinis obtectis. Coronae foliata extus decurva, oblanceolato-elliptica, antice acuminate postice obtuse subtus in auriculas 2 oblongas foliolum superantes extensa, antheris inferiors. An ornamental climbing vine with leaves of similar shape to H. bordenii Schlechter but with pinnate venation. In addition to the distinctive foliage this species has a very long (20 cm.) glabrous, terete peduncle. The corona is very upright with the inner lobes overtopping the anthers. The outer lobe and bifid side appendages are at 90 degrees to the floral axis. The bilobed extensions are beautifully linearly lined; surround and extend beyond the outer scale apex which is cut off rather abruptly . The anther appendages are swollen and noticeable. The bilobed extensions are modified into a skirt, thick, fleshy and inflated, extending under the outer apex of the anther appendage, not however, as in the Otostemma section. Leaf Blade: 1 1-16 cm. x 2.4- 2 cm., pinnate venation, both surfaces glabrous. Petiole: heavy, curved 1.3-1.6 cm. long, probably grooved above. Stems: stout, rooting glabrous. Peduncles: extremely long 20 cm. cat 0.2 cm. In diameter, round, glabrous. Rachis: columnar. Pedicel: curved, filiform, terete, 1.5-2.8 cm. x 0.05 cm. Calyx: somewhat triangular, narrowly rounded apex, 0.21 cm. long and 0.17 cm. at the widest (near the base); ciliate, ligules are present. Ovaries: 0.12 cm. tall 0.13 cm. at the base. Corolla: outside glabrous, Inside pubescent except for the apical area, apex acute. Sinus to sinus 0.33 cm.; sinus to apex 0.37 cm.; apex to center 0.57 cm. making the flower diameter flattened 1.14 cm.; sinus to center 0.25 cm.; widest portion of corolla lobe 0.32 cm. Corona: central portion upright even in dry specimen. Inner lobe reaching above the anther appendage, inner lobe ligulate. Other details above. Sulcate below formed by the bilobed extensions. Stigma Head: (staminal apex area): short triangular with a mealy central head. Pollinia: 0.57 mm. long, 0.15 mm. wide. Translators short. Retinaculum 0.19 mm. long. Translation: Perennial epiphyte, branching, climbing. Stems flexible, loosely leaved, round glabrous, well leaved, rooting. Leaves erect or somewhat erect, lanceolate acuminate, base somewhat wedge shaped , rounded, glabrous; peduncle very long, glabrous; pedicels narrow, glabrous, flowers medium sized for the section. Calyx segments ovate, glabrous, margins ciliate, much smaller than the corolla. Corolla revolute, divided all the way to 1/3 toward base of 5 parts, lobes triangular, acuminate, outside glabrous, inside except for the glabrous apex pubescent, the other side somewhat broad, crystalline over spread, leaflets of the corona outside curved downward, oblanceolate, elliptic, inside apex acuminate, at the base obtuse below with tongues, 2 oblong leaflets extended above, anthers inferior.

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In Hoya Section Acanthostemma (Blume) R. D. Kloppenburg (1994) 45-47. R. D. Kloppenburg. Hoya gigantanganensis Kloppenburg in Fraterna 4th Quarter 1992. Type #36787 PNH collected by Yoshio Kondo & G. Edano, 26 March 1957, at Gigantangan, Leyte, Philippines. Section Acanthostemma (BL) Kloppenburg. An ornamental climbing vine with leaves of similar shape to H. bordenii Schlechter but with pinnate venation. In addition to the distinctive foliage this species has a very long (20 cm.) glabrous, terete peduncle. The corona is very upright with the inner overtopping the anthers. The outer lobe and bifid side appendages are at 90 degrees to the floral axis. The bilobed extensions are beautifully linearly lined; surround and extend beyond the outer scale apex which is cut off rather abruptly. The anther appendages are swollen and noticeable. The bilobed extensions are modified into a skirt, thick, fleshy and inflated, extending under the outer apex of the anther appendage not, however, as in the Section Otostemma Schlechter. Leaf Blade: 11-20 cm. x 2.4-6.2 cm., pinnate venation, both surfaces glabrous. Petiole: heavy, curved 1.3-1.5 cm. long, probably grooved above. Stems: stout, rooting, glabrous. Peduncle: extremely long 20 cm. ca. 0.2 cm. in diameter, round, glabrous. Rachis: columnar. Pedicel: curved, filiform, terete, 1.5-2.8 cm. x 0.05 cm. Calyx: somewhat triangular, narrowly rounded apex, 0.21 cm. long and 0.17 cm. at the widest (near the base) ciliate, ligules are present. Ovaries: 0.12 cm. tall 0.13 cm. at the base. Corolla: outside glabrous, inside pubescent except for the apical area, apex acute. Sinus - Sinus: 0.33 cm.; Sinus-Apex 0.37 cm.; Apex - Center 0.57 making the flower diameter flattened 1.14 cm.; Sinus - Center 0.25 cm.; widest portion of corolla lobe 0.32 cm. Corona: central portion upright even in dry specimen. Inner lobe reaching above the anther appendage, inner lobe ligulate. Other details above. Sulcate below formed by the bilobed extensions. Stigma Head: (staminal apex area): short triangular with a mealy central head. Pollinia: 0.57 mm. long, 0.15 mm. wide. Translators short. Retinaculum 0.19 mm. long. References: Philippine Hoya Species 2nd. Ed. (1993) 61. In Philippine Hoya Species 3rd Edition (1996) 73-74. R. D. Kloppenburg. Hoya gigantanganensis Kloppenburg in Fraterna 4th Quarter 1992.Type #36787 PNH collected by Yoshio Kondo & G. Edano, 26 March 1957, at Gigantangan, Leyte, Philippines. Section Acanthostemma (Blume) Kloppenburg. An ornamental climbing vine with leaves of similar shape to H. bordenii Schlechter but with pinnate venation. In addition to the distinctive foliage this species has a very long (20 cm.) glabrous, terete peduncle. The corona is very upright with the inner lobes overtopping the anthers. The outer lobe and bifid side appendages are at 90 degrees to the floral axis. The bilobed extensions are beautifully linearly lined; surround and extend beyond the outer scale apex which is cut off rather abruptly. The anther appendages are swollen and noticeable. The bilobed extensions are modified into a skirt, thick, fleshy and inflated, extending under the outer apex of the anther appendage not, however, as in the Otostemma Section.

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Leaf Blade: 11-16 cm. x 2.4-6.2 cm., pinnate venation, both surfaces glabrous. Petiole: heavy, curved 1.3-1.5 cm. long, probably grooved above. Stems: stout, rooting, glabrous. Peduncle: extremely long 20 cm. ca. 0.2 cm. in diameter, round, glabrous. Rachis: columnar. Pedicel: curved, filiform, terete, 1.5-2.8 cm. x 0.05 cm. Calyx: somewhat triangular, narrowly rounded apex, 0.21 cm. long and 0.17 cm. at the widest (near the base) ciliate, ligules are present. Ovaries: 0.12 cm. tall 0.13 cm. at the base. Corolla: outside glabrous, inside pubescent except for the apical area, apex acute. Sinus Sinus 0.33 cm.; Sinus-Apex 0.37 cm.; Apex - Center 0.57 cm. making the flower diam. flattened 1.14 cm.; Sinus - Center 0.25 cm.; widest portion of corolla lobe 0.32 cm. Corona: central portion upright even in dry specimen® Inner lobe reaching above the anther appendage, inner lobe ligulate. Other details above. Sulcate below formed by the bilobed extensions. Stylar apex: short triangular with a mealy central head. Pollinia: 0.57 mm. long, 0.15 mm. wide. Translators short. Retinaculum 0.19 mm. long. Meaning: gigantanganensis named for the palace of discovery Gigantangan Leyte.

Hoya gigas Schlechter In Botanische Jahrbücher 50 (1913) 136. "Die Asclepiadaceen von Neu Guinea” R. Schlechter. 49. H. gigas Schltr. n. sp.. — Suffrutex ramosus, alte scandens Rami filiformes, flexuosi, laxe foliati, teretes, glabri minute et molliter tomentosuli. Folia erecto-patentia vel patula oblonga vel elliptico-oblonga, acuminata, basi rotundata, superne glabra, subtus brevissime tomentosuli, petiolata, petiolo tomentosulo, superne leviter sulcato. Inflorescentia, pedunculatae, umbelliformes, 3—7 florae, pedunculo pedicellisque aequilongis brevissime tomentosulis. Calycis foliola suborbicularia, im aequimagna, brevissime tomentosula, quam corolla multo breviora. Corolla subrotata in genere maxima, usque supra medium 5-fida, extus sparsim puberula, intus microscopice papillosa, basi intus anulo villoso circumdatia, lobis triangulis, acutis, margine ciliatis. Coronae foliola suborbiculari-ovalia antice breviter rostrata, medio concava, extus obtusissima. Pollinia falcato-clavata, translatoribus curvatis, polliniis subaequilongis, retinaculo rhomboideo polliniis paulo minore. Eine prächtige hoch-kletternde Liane, mit schnurartigen locker beblättern Zweigen. Blätter 9—14 cm lang, in der Mitte 4.2—5.5 cm breit, Blattstiele l.5—2 cm lang. Blüten stände doldig 3—7 blütig auf 2.5—3 cm langem, sehr kurz filzigern Stiel. Blütenstiele 2.5—3 cm lang, sehr kurz filzig. Blüten in der Gattung sehr grosz denen der H. lauterbachii R. Schum. ähnlich und fast gleichgrosz. Kelchblätter ungefähr 6—8 mm im Durchmesser. Korolla sehr breit, schüsselförmig. ausgebreitet etwa 8.2 cm im Durchmesser. Gynostegiumsäule etwa 1.4 cm hoch. Koronaschuppen breit und kurz, von der Spitze bis zum äuszeren Rande 7 mm lang Nordöstl. Neu-Guinea: auf Bäumen in den Wälden auf den Gomadjidji, am Waria, ca. 450 m. ü. M. (Schlechter n. 19389. — Blühend im Mai 1909). In dieser Art liegt eine nahe Verwandte der H. Lauterbachii K. Schum. mit etwat; gleichen Blütendimensionen. Die beiden Artet sind unschwer auseinander zu innen

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besonders nach der Mitte zu dicht sammelhaarig, Die Blüten der varliegenden Art sind rot, die Korona gelb mit braunem Band. Translation: See below: Hoyas of Northeastern New Guinea. In Exotica III 9th Ed. (1978) 1631. A. B. Graf. Hoya gigas (New Guinea); one of many Hoyas we collected on Government expedition in the Finisterre Mountains; a beautiful liane with 4-6 cm waxy flowers reddish brown with light tips. In New York Botanical Garden Illustrated Encyclopedia of Horticulture 5 (1981) 1726. H. gigas, of New Guinea, is a high climber. Its broad-elliptic to ovate leaves, up to 5 inches long, have hairy under surfaces. The starry, cupped flowers, 1 ½ to 2 ½ inches in diameter, are reddish brown with paler apexes to the petals. In Hoyas of Northeastern New Guinea 1 (1992) 97-98. R. D. Kloppenburg. (Translation). 49. Hoya gigas Schlechter new species. A high climbing branched half shrub, branches threadlike, flexible, loosely leaved, round, glabrous, minutely and softly tomentose. Leaves erect spreading or outspread, oblong or elliptic oblong, acuminate, base rounded, glabrous above, below very shortly tomentose, petiolate, with the petiole minutely tomentose, above lightly grooved. Inflorescences pedunculate, shaped like an umbel. 3-7 flowered, pedicels of the peduncle of equal length, minutely tomentose. Lobes of the calyx somewhat rounded, unequal, very shortly minutely tomentose, much shorter than from the corolla. Corolla almost rotate, largest in the Genus, 5 parted all the way to above the middle, outside sparsely puberulous , inside microscopically papillose (pimply), at the base a ring of long soft straight ascending hairs, lobes triangular acute, with ciliate margins. Scales of the corona almost round-oval, with the apex shortly beaked, in the center concave, outside very blunt (rounded). Pollinia sickle shaped, with the translators curved, a little shorter than the pollinia, retinaculum rhomboid, a little shorter than the pollinia. A magnificent high climbing vine, with cord like, loosely In this species there is a close relationship to H. lauterbachii K. Schumann with about the same size flowers (bloom dimensions). The two species are easy to tell apart, however for with H. lauterbachii K. Schumann, the leaved branches. Leaves 9-11 cm. long, in the middle 4.2-5.5 cm. wide. Petiole 1.5-2 cm. long. Inflorescences in umbels, 37 flowered on a 2.5-3 cm. long, very shortly tomentose peduncle. Pedicels 2.3-3 cm. long, very shortly tomentose. Blooms very large for the section, similar to H. lauterbachii K. Schumann, almost of equal size. Sepals approximately 6-8 mm. in diameter. Corolla very broadly bowl shaped, about 8.2 cm. in diameter when spread out. Gynostegium about 1.4 cm. high. Corona scales broad and short, from the apex to the outer rim 7 mm. long. Northeastern New Guinea: on trees in the forests of the Gomadjidji, on the Waria, about 450 meters altitude (Schlechter #19389 - Blooming in May 1909).corolla is more deeply bowl shaped and inside especially toward the middle, has thick velvet like hairs. The blooms of the specie are red with a yellow corona, with a brown band. In Tropica 4 (1992) 1020. A. B. Graf. Hoya gigas (New Guinea); one of the many Hoyas we collected on government expedition in the Finistere Mountains; a beautiful

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liane with 4-6 cm waxy flowers reddish brown with light tips. Tropical. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. gigas Schlechter (Schlechter 19389) –15. In Dr. Schlechter’s Hoya Species 1 (1993) 66-67. R. D. Kloppenburg. Hoya gigas Schlechter new species. A high climbing branched half shrub, branches threadlike, flexible, loosely leaved, round, glabrous, minutely and softly tomentose. Leaves erect spreading or outspread, oblong or elliptic oblong, acuminate, base rounded, glabrous above, below very shortly tomentose, petiolate, with the petiole minutely tomentose, above lightly grooved. Inflorescences pedunculate, shaped like an umbel. 3-7 flowered, pedicels of the peduncle of equal length, minutely tomentose. Lobes of the calyx somewhat rounded, unequal, very shortly minutely tomentose, much shorter than from the corolla. Corolla almost rotate, largest in the Genus, 5 parted all the way to above the middle, outside sparsely puberulous , inside microscopically papillose (pimply), at the base a ring of long soft straight ascending hairs, lobes triangular acute, with ciliate margins. Scales of the corona almost round-oval, with the apex shortly beaked, in the center concave, outside very blunt (rounded). Pollinia sickle shaped, with the translators curved, a little shorter than the pollinia, retinaculum rhomboid, a little shorter than the pollinia. A magnificent high climbing vine, with cord like, loosely leaved branches. Leaves 9-11 cm. long, in the middle 4.2-5.5 cm. wide. Petiole 1.5-2 cm. long. Inflorescences in umbels, 3-7 flowered on a 2.5-3 cm. long, very shortly tomentose peduncle. Pedicels 2.33 cm. long, very shortly tomentose. Blooms very large for the section, similar to H. lauterbachii K. Schumann, almost of equal size. Sepals approximately 6-8 mm. in diameter. Corolla very broadly bowl shaped, about 8.2 cm. in diameter when spread out. Gynostegium about 1.4 cm. high. Corona scales broad and short, from the apex to the outer rim 7 mm. long. Northeastern New Guinea: on trees in the forests of the Gomadjidji, on the Waria, about 450 meters altitude (Schlechter #19389 - Blooming in May 1909). In this species there is a close relationship to H. lauterbachii K Schumann with about the same size flowers (bloom dimensions). The two species are easy to tell apart, however for with H. lauterbachii K. Schumann, the corolla is more deeply bowl shaped and inside especially toward the middle, has thick velvet like hairs. The blooms of the specie are red with a yellow corona, with a brown band.

Hoya glabra Schlechter In Beiblatt zu den Botanischen Jahrbücher #92 40 (1908) 13. R. Schlechter. Hoya glabra Schltr. n. sp.; volubilis, alte scandens, ramosa, epiphytica; ramis ramulisque filiformibus elongatis, glabris, laxe foliatis; foliis patulis ovatis acuminatis, basi rotundatis, glabris, textura carnosis, 14—16 cm longis, infra medium 7.5—9.5 cm latis, nervis 3 primariis praecipue conspicuis, petiolo crasso 2.5—3.2 cm longo; cymis pedunculatis umbelliformibus 15—25 floris; pedunculo crasso tereti, glabro, 5—6 cm longo, pedicellis filiformibus glabris, 1.5—2 cm longis, glabris; floribus in genere mediocribus purpureis; calycis segmentis ovatis obtusis, glabris, margine tantum

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minutissime ciliolatis, vix 0.2 cm longis, corolla subrotata c. 1 cm diametiente, usque infra medium 5-lobata, lobis ovatis acutis reflexis, apice tantum subinconspicue puberulis, ceterum glabris; coronae phyllis carnosis horizontalibus, superne ellipticis antice breviter rostratis, callo elliptico infra apicem ornatis, subtus oblongis obtusis, medio late foveolatis; antheris trapezoideis, appendice hyalina ovata obtusa, coronae folia vix excedente; polliniis oblique cuneato-oblongoideis, margine exteriore carinatomarginatis, translatoribus perbrevibus, retinaculo minuto; stigmatis capite conico. Borneo: auf Bäumen in der Wäldern an Long-Wahau (R. Schlechter n. 13458, blühend im August 1901). Mit H. latifolia Bl. am Nächsten verwandt. Translation: twining, high climbing, branched, epiphytic, branches and branchlets threadlike elongate, flexible, glabrous, loosely leaved; leaves outspread ovate acuminate, base rounded, glabrous, fleshy texture, 14—16 cm long in the middle 7.5—9.5 cm wide, 3 primary nerves especially conspicuous, petiole thick 2.5—3.2 cm long; cymes pedunculate formed like umbels of 15—25 flowers; peduncle thick round, glabrous, 5— 6 cm long, pedicels threadlike glabrous, 1.5—2 cm long, glabrous; flowers midsize in the genus, purple; calyx segments ovate, obtuse, glabris, margines only minutely ciliate, barely 0.2 cm long. corolla somewhat rotate about 1 cm in diameter, 5 lobed to below the middle, lobes ovate acute apex reflexed, apexes barely somewhat conspicuously puberous, otherwise glabrous; leaflets of the corona fleshy horizontal, above elliptic in front briefly beaked, with a beautiful ornate elliptic apex below, below oblong obtuse, in the middle broadly channeled, anthers trapezoidal appendages ovate hyaline leaflets of corona barely exceeding; pollinia oblique cuneate-oblong, exterior margines with keels, translators very small, retinaculum minute, stigma head conic. Borneo: on trees in the forest near long-Wahau (Schlechter n. 13458, blooming in August 1901) Most closely allied to H. latifolia Blume. Compilers Notation: Blume did not name a H. Latifolia, the species was named by George Don. In Journal of the Royal Asiatic Society Straits Branch (Spec. #) (1921) 506. H. Ridley. 5. Hoya glabra Schltr. in Engl. Bot. Jarb. 40 (1908) Beibl.92: 14. Borneo. Schlechter 13458. Endemic. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. glabra Schlechter (Schlechter 13458) – 14. In Dr. Schlechter’s Hoya Species (1993) 68-69. (Translation) R. D. Kloppenburg. Hoya glabra Schlechter Published by Dr. R. Schlechter in: Botanische Jahrbücher , Beiblatt 40 #92 (1908) 13 "Borneo Is., Molucca”. Twining high climber, branched, epiphytic; branches and branchlets threadlike, elongate, flexible, glabrous, loosely leaved; leaves outspread ovate acuminate, with the base rounded, glabrous, texture leathery, 14-16 cm. long, below the middle 7.5-9.5 cm. wide, with the 3 primary nerves especially conspicuous; petiole thick 2.5-3.2 cm. long; cymes pedunculate, shaped like umbels, 15-25 flowered,; peduncle thick, round, glabrous 5-6 cm. long, pedicels threadlike, glabrous 1.5-2 cm. long, glabrous; flowers in general midsize, purplish; with the segments of the calyx ovate obtuse, glabrous, margins to a degree only, ciliate, barely 0.2 cm. long; corolla somewhat rotate about 1 cm. in diameter,

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5 lobed all the way to below the middle, lobes ovate acute, apex reflexed, with the tip to a degree only somewhat inconspicuously puberulous (downy), otherwise glabrous; with the scales of the corona fleshy, horizontal, above the middle broadly pitted; anthers trapezoidal, with the appendages hyaline, ovate obtuse, with the apex of the corona barely exceeding; pollinia oblique cuneate-oblong, exterior margin keeled, translators very short, retinaculum minute; with the head of the stigma conic. Borneo: On trees in the Forest at Long-Wahau (Schlechter #13458, flowering in August 1911). Most closely allied with H. latifolia Blume.

Hoya globiflora Ridley In Journal of Federated Malayan States Mus. 5 (1915) 164. “Prenn. Mal.” H. Ridley. Koh Pennan. Disfrib.—Indo-Malaya. 47. HOYA GLORIFLOR. Sp. nov. Stems pale corky, 2 mm. in diameter. Leaves ovate subacute with rounded bases 6.5 cm. long, 3.5 cm. wide, nerves 3 pairs and nerrules few visible when dry, one pair from the base ascendinig, the others short horizontal soon broken up, petiole very thick, 6 mm. long. Peduncles stout 3 to 4 cm. long, raceme 1 cm. long, stout, occasionally branched. Flowers innumerable forming a large globose umbel 4-5 cm. across when dry. Pedicels 1.5 cm. long. Sepals 5, short ovate obtuse. Corolla 1 cm. across waxy white a pink tinge, lobes ovate sub-obtuse. Corona large, upper lube short erect tooth-like, lower ovate, spreading obtuse. apex emarginate above depressed. Staminal column short. Anther cells incumbent over the style apex. Pollen masses linear oblong straight blunt f1uttened, caudicles very minute (hardly any) Carrier small, triangular, dark brown. Koh Pennan. No. 5756. Creeper.

Hoya globosa LeConte In Flore General De l’Indo-chine (1912) 131. H. globosa N. H. LeConte. (here in Figure 18. a leaf drawing under 10; also in key under 21 but not described. Hoya globulifera Blume In Museum Botanicum Lugduno-Batavum 1 (1849) 44. C. L. Blume. 102. Hoya globulifera Bl.: volubilis, glabri; foliis subsessilibus ex ovato ellipticus acutis basi rotundatis carnosis planis subaveniis supra ad basin macula alba notatis subtis punctatis; umbellis pedunculatis subglobosis; corollae intus sericeo-velutinae laciniis triangulariovatis acutis; coronae staminae foliolis lanceolatis supra canaliculatis angulo exteriore obtuso. — In sylvis litoralibus Novae Guineae. Flores magnitudine qua in H carnosa, fusco-purpurei. Translation: twining, glabrous leaves almost sessile and ovate elliptic acute with the base rounded fleshy flat lightly veined above toward the base large white spots below punctate; umbels pedunculate almost globose; lobes of the corona inside intensely hairy

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triangular ovate acute; leaflets of the staminal corona lanceolate above channeled with the exterior angle obtuse. — From the trees near the shore on New Guinea. Flowers the size of H. carnosa., dark purple. In Flora Indiae Bataviae 2 (1857) 520. Zollinger & Miquel. 12. Hoya globulifera Bl. Volubilis, folia subdensa brevi-petiolata ovato-elliptica acuta subacuminata, carnosa; subavenia, glabra, supra ad basin macula alba notata, subtus punctata, 3 — 3½ pollicares, suprema internodiis valde elongatis distantia, pedunculi pollicares ex apice incrassato umbelliflori, pedicelli tenues pedunculum aequantes, corollae fusco-purpureae intus sericeo-velutinae triangulari-ovatae acutae, coronae stamineae phylla lanceolati extrorsum obtusa supra canaliculatis. Blume Mus. bot. I. p. 44. Nieuw-Guinea, in de wouden der kunststrecken (Zippelis). Translation: Twining, leaves somewhat dense shortly petioled ovate-elliptic acute somewhat acuminate, fleshy; lightly veined, glabrous, above near the base a marked white spot, below punctate, 3 — 3 ½ inches, topmost distant internodes exceedingly elongated, peduncles an inch from end coarse umbel flowered, pedicels thin same length as the peduncles, with the corolla dark-purple inside thickly hairy triangular ovate acute, leaflets of the staminal corona lanceolate outside obtuse above channeled. ................ New Guinea in the trees of the coastal woods. In Annales du Jardin Botanique de Buitenzorg 1 (1876) 37. R. H. C. C. Scheffler. Hoya globulifera Bl.; Miq., l. c., p. 520. Annales Botanices Systematicae 3 (1852) 66. W. G. Walpers. 6. H. globulifera Blume, Mus. Bat. p. 44, no. 102 † — Volubilis glabra; foliis subsessilibus ex ovato ellipticis acutis basi rotundatis carnosis planis subaveniis, supra ad basin macula alba notatis, subtis punctatis; umbellis pedunculatis subglobosis; corollae intus sericeo-velutinae laciniis triangulari-ovatis acutis, coronae staminae foliolis lanceolatis, supra canaliculatis, angulo exterior obtusa. — Habitat in sylvis litoralibus Novae Guineae. Translation: Twining glabrous; leves almost sessile from a acute rounded base ovateelliptic, fleshy flat almost veinless; above near the base a marked white spot below punctate; umbels pedunculate almost globose, flaps of the corolla inside densely velvety triangular-ovate acute. leaflets of the staminal corona lanceolate, above channeled with the exterior angle obtuse. Habitat in the shore forests of New Guinea.

Hoya globulosa Hooker In Floral Magazine (1880) 406. Hoya globulosa, Hook. f. Under the name of Hoya Globulosa, the Cranston Seed and Nursery Company. Hereford, exhibited at the meeting of the royal Horticultural Society, on April 13th, a charming Hoya, so pretty and distinct as to merit and receive a First-class Certificate on Merit. Some doubt being expressed as to whether is was this species or not, the plant was referred to Kew for the opinion of the authorities there, and Sir J. D. Hooker has informed the Company that “the Hoya you wrote about is a plant discovered by myself in India in 1849, and specimens of it collected by me are in the Herbarium here. I have called it H. globulosa, Hook. f.” We can confidently recommend it as a most useful and valuable plant, blooming

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profusely and thriving well under the same treatment as other Hoyas. The flowers are of a bright straw colour, the coronal protuberances white, the interstices of a shining brownish-pink, forming a very pleasant contrast; trusses large and freely produced; and the leaves of a shining green colour, with dark green veins. In the Gardener’s Chronicle 2 (1882) 732. J. D. Hooker f. Hoya globulosa, Hk, f. ined. (fig. 115, p. 741). This is newly introduced from Sikkim Himalaya, where it was discovered by Sir Joseph Hooker, who is the sponsor for the name, though he has not hitherto, unless we are mistaken, published it. The plant was sent out by the Cranston Nursery Company, Hereford, who have exhibited it on several occasions, and who kindly furnished the materials for our illustration. In The Gardener’s Chronicle (1882) 570. Fertilization of Hoya globulosa. Mr. W. G. Smith refereed to this subject and exhibited Hoya flowers with flies attached to the glutinous discs of the pollen masses. This hoya is highly fragrant; this fragrance is very attractive to insects and it is by insect agency alone that flowers can be fertilized in the state of nature. The flowers of the Hoya differ from Orchids in their perfectly regular form, but they agree somewhat in their pollen grains, which are grouped in masses of two placed in five small pouches. The five little gelatinous dark coloured disks, belonging to the bases of the pollen masses, are the only parts visible in the open flower. In Hoya there are five pairs of pollen-masses arranged regularly round the centre of each flower. When insects alight on the flowers (attracted by the powerful fragrance) it is almost inevitably happens that one foot slips and gets caught by one of the five gelatinous discs. In the effort of the insect to escape, two, three, or even four to its other feet are almost sure to get similarly caught. The insect then tries with all its power to free its limbs, and if these efforts are successful the pollen masses are drawn out of the pouches with the feet. The basal appendages of each pair of pollen masses are elastic, and when in the pouches are like an extended spring, but the instant the masses are drawn out by the insect’s foot the spring closes, the two pollen masses quickly cross each other and hold tightly on to the insect’s little claws. If the insect is week it cannot withdraw its legs at all, and so perishes in the flower; but if strong it flies away with one to five pairs of pollen-masses clasped round its feet. Sometimes an insect breaks part of a leg off in its efforts to escape. The five stigmas are not ready to receive the pollen at the time the pollen is ripe, so that it is only when the insect re-alights on to some neighboring Hoya flower in a different stage of growth that cross-fertilization takes place. In Flower Garden 2 (1883) 95-96. Paxton. Hoya globulosa. Although this species has much in general character, in its habit of growth, and conformation of its flowers, with several others of the family — the Cunningham Hoya (H. Cunninghami), for instance— still, from the cultivator’s point of view, it is sufficiently distinct to make it an acceptable addition to our warm house climbers. One advantage possessed by this and other twining Hoyas is that they are not such coarse rampant growers as many stove climbers are, consequently do not too much shade the plants the plants grown beneath them in houses where they are suspended on the rafters overhead. H. globulosa will no doubt succeed under like conditions to most of the other species, thriving in ordinary loam or peat, with too much pot-room. It will most likely not require more warmth than that of an

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intermediate house. It has been recently sent out by Cranston Nursery Company, Hereford, who exhibited it London. The plant is thus described in the Gardener’s Chronicle, p. 732, vol. xvii. Leaves oblong, leathery, rounded or subcordate at the base, acuminate at the apex. Flowers in globose umbels, ground-colour pale straw or cream, coronal appendages pink at the base. A native of the Sikkim Himalaya. Compiler’s Notation: No Hoya with the name of Cunninghami has ever been written up (described). In Flora of British India 4 (1883) 60. J. D. Hooker. 34. H. globulosa, Hook f. in Gard. Chron. 1882, ii 732, f. 115; more or less hairy, leaves 5-7 in. long-petioled elliptic or oblong cuspidate or acuminate base rounded midrib very stout, nerves distinct, peduncle long, pedicels villous, corona nearly glabrous within. Floral Magazine, 1860* t. 406. — Hoya n. 30 and 33; Herb. Ind. Or. H. f. & T. Sikkim Himalaya; alt. 1-3000 ft. J. D. H., Clark. Assam; Muku hills. Simmonds. Cachar, more or less hairy, the hairs long and very deciduous. Leaves 2-3 ½ in., pedicels 1 in. Sepals rounded. Corolla1/2 in. diam., cream coloured; lobes short, incurved. Corona-processes short, broadly elliptic, pink concave above, inner angle produced into an erect spur which is shorter than the large broad anther tips. Follicles 12-16 in. long, very slender, cylindric, pericarp thin. — seeds ½ in. long, slender. * Reported here as 1860 should be 1880. In Dictionary of Gardening 2 (1884) 155. George Nicholson. H. globulosa (globular)* fl. pale straw or cream-colour, disposed in globose umbels; corona pink at base. April. l. oblong coriaceous, rounded at the base, acuminate at the apex, more or less hairy. India, 1880. (G.C. n.s., xvii. 741.) In Flore Generale des Indo Chine 4 (1912) 140 J. Constantin (H. M. LeConte) . 21. H. globulosa Hook. in Gard. Chron. 1882, II, p. 732, fig. 115; Floral Magaz., 1860, tab. 106*. Tige glabre ou avec quelques poils. Feuilles elliptiques ou oblongues, arrondies a la base, cuspidees ou acuminees; nervure mediane teres vigoureuse avec quelques glandes et poils a la base; nervures secondaires obliques et arquees. Inflorescence: ombelle multiflore portee sur un pedoncule aile poilu ou glabre, long de 7-8 cm.; pedicelles greles, longs de 1.5-2 cm., couverts de poils ocraces ou jaunatres. — Calice a sepales arrondis, poilus et cilies. Corolle de 10-12 mm.; lobes triangulaires-aigus, courts et incurves, pubescents inrerieurment et cillies fortement aux bords. Coronule a 5 pieces courtes, largement elliptiques, rosees, concaves en dessus, presentant a la partie interne un eperon dresse qui est plus court que l’extremite large des antheres. Fruit: follicules longs, tres greles, cylindriques, longs de 30-40 cm., a pericarpe mince; graines greles (8 mm.). Cochinchine: Mt. La (Pierre). — Indes angl. Pedicelles poilus. Poils des pedicelles jaunatres; feuilles grandes, allongees (13 cm. x 5.5 cm.)........ 21 H. globosa. * The citations here are incorrect. The year is 1880 and the tab. is 406.

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Translation: Glabrous stem or with some polishes. Elliptic or oblong, leaves rounded at the base, cuspidate or acuminees; midrib vigorous teres with some glands at the petiole base; secondary nerves turn off and are arched. Inflorescence: umbulate multiflowered reach on a shaggy or glabrous stalked wing, 7-8 cm. long; spindly pedicels, 1.5-2 cm. long, covered with a few hairs ocraces colored. - Calyx is rounded, shaggy sepals and ciliate. Corolla of 10-12 mm.; triangular-shaped, short lobes and curved, pubescent inwardly and very ciliate at the edges. Corona has 5 short, extensively elliptic pieces, dews, concave on top, presenting has the internal part in a spur that is shorter than the large extremities of the anthers. Fruit: long, very spindly, cylindrical follicles, 30-40 cm. long, has thin pericarpe; spindly seeds (8 mm.). Cochinchine: Mt. La (Pierre). - Indies angl. Shaggy Pedicels. Pedicels glabrous yellow; leaves large (13 cm. x 5.5 cm.)........ 21. H. globosa. In Journal of the Linnean Society 43 (1915) 476. C. C. Lacaita. Hoya globulosa, Hook. fil. Glen S. E. of Kalimpong, c 1900’, 23. iv. (16041). In Standard Cyclopedia of Gardening and Horticulture (1930) 1603. F. H. Bailey. Hoya globulosa, Hook. f. Hairy: lvs. elliptic-oblong or long-oblong, acuminate, rounded at the base, the midrib very stout, the petiole an inch or less long: fls. pale straw-or cream-color, the star like crown-segms. white, with pink at the base, borne in dense, globular umbels: follicles a foot or more long, Sikkim. F. M. 1880:406. G.C. II. 17:741.— A handsome species, requiring a general given to H. carnosa. In The Royal Horticulture Society Dictionary of Gardening (1965) 1016. (1st Ed. 1951). H. globulosa l. oblong, rounded at base, slender-pointed, leathery, more or less hairy. fl. pale straw or cream, in globose umbels; corona pink at base. April. Sikkim. 1880. (G. C. 17 (1882), 741.) Succeeds with the same treatment as H. carnosa. In Journal of the Royal Horticultural Society (1973). A. J. Wills. Hoya globulosa. Scent is over-poweringly heavy and languid, reminiscent of sweet chestnuts. In Illustrated Florae di Cera. Ann. Fac. Agric. 27 (1975) 340. Giancinto Donno. Hoya globulosa Hook. Pianta tomentosa. Foglie elliptico-oblunghe, acuminate, arrotondate alba base; la nervatura principale ben robusta, il picciuolo di circa 3 cm o meno; foglie crema-paglia-pallida; la corolla* stellata bianca con color rosa o rosso alba base, constituisce un’ombrella globulosa compatta, follicoli lunghi 30 cm o piu; * the word here should read “corona” not corolla. Translation: Tomentose plant. Leaves elliptic-oblong, acuminate, rounded at the base; the principle nerves very prominent, the stalk is about 3 cm or less; leaves pale cream-straw; the corona starlike white with rose or reddish white at the base, forming a globose compact umbel, follicles 30 cm or more long. In Exotica III 9th Ed. (1978) 1631. A. B. Graf. Hoya globulosa (Sikkim, Himalayas) , slender vine with thick oval-pointed, spoon-like, lightly hairy leaves, and round clusters of cream colored waxy flowers, the corona marked brown-red at base,

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In Botanical survey of India, Calcutta “Bengal Plants” 2 (1983) 519. D. Prain (R.K. Basak) 269. Hoya globulosa Hooker. f. F. B. I. iv. 60. Chittagong A stout climber. In The New Royal Horticulture Society Dictionary globulosa Hook. f. Climbing with downy stems and puberulent, leathery, base rounded. apex mucronate. lobes cream-yellow or straw-yellow; corona with a summer. N. India, Nepal, Tibet. Z9.

of Gardening 2 (1992) 596. H. branches. Lvs. to 15cm. oblong, Umbels globose, many-fld: cor. pink base, apex cream. Spring,

In Tropica 4 (1992) 1020. A. B. Graf. Hoya globulosa (Sikkim, Himalayas); slender vine with thick oval-pointed, spoon-like, lightly hairy leaves, and round clusters of cream colored waxy flowers, the corona marked brown-red at base. Subtropical.

Hoya golamcoiana Kloppenburg In Fraterna 3 (1991) II. R. D. Kloppenburg. Hoya golamcoiana Kloppenburg n. sp. Type sheet #41930 E1 Nido Rest Area Palawan, Philippines found by Professor Juan V. Pancho 15 August 1988. Holotypus: CAHUP. Suffrutex parum ramosus, dependens gracillimus. Rami filiformes, flexuosi, bene foliati, teretes, glabri. Folia laxa, breviter petiolata, oblongo-elliptica, obtusiuscule acuminate, utrinque glabra, petiolo glabro. Inflorescentiae umbelliformes c. 15 florae, pedicellis filiformibus, sparsum pubescenta. Calycis foliola, lanceolato-ovata, subacute, margine ciliate, quam corolla paulo breviore. Corolla subrotata, usque ad tertiam partem basilarem 5-fida, extus glaberrima, intus sparsum pubescenta, lobus recurvis, acutis. Coronae foliola patentia marginum exterioreum versus leviter gibbo obscuro ornate, apice apiculata, extus subacute. Pollinia oblique oblongoidea, translatoribus brevissimus, retinaculo minutissimo rhomboideo. This species differs from other known Hoya species. Its foliage blades are similar to H. cumingiana Decaisne but it is petiolate and on a dangling plant. It was collected by Professor Juan V. Pancho at El Nido rest Area Northern Palawan Is., Philippines growing from cracks in limestone areas. In its preferred habitat it is also similar to H. cumingiana Decaisne. The leaves are 5.3 cm. long and 2.7 cm. wide at the widest portion, enervis, although in dried specimens the midrib and five pairs of pinnate nerves are visible. It is a densely leaved hanging vine with almost sessile petioles, 0.3 cm. long. with c. 15 white reflexed flowers per umber. In this respect it also differs from H. odorata Schlechter, which has few and larger flowers. Internodes are 3.05.0 cm. long, stems round, 0.3 cm in diameter. Peduncles are 2.7 cm. long, sparsely pubescent. Pedicels are 2.4 cm. long, 0.09 cm. in diameter, sparsely pubescent. Calyx lobes sparsely ciliate, calyx about 0.4 cm. in diameter, ligules present, with dark tips. Corolla reflexed, very finely pubescent, 1.5 cm. in diameter, flat. The species is distinctive and attractive. Other measurements and detail can be seen in the accompanying drawing by Ruel D. Medina, botanical artist, drawn from live material in 1988 at the University of the Philippines, Los Banos, Laguna Province, Philippines.

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This new species is named in honor of Andres S. Golamco Jr. who has devoted much time and love to Philippine plants and who has done a remarkable job in completing a comprehensive publication on the Philippine orchids.

Hoya gonoloboides Regel In Acta Horti Petropolitani 8 (1883) 642. E. Regel. 3. Hoya (Dregia E Meyer) gonoloboides Rgl. Volubilis, caulibus fusco-hispido-villosis. Folia membranacea penninervia, reticulato-venosa, cordato-ovata, acuminata, petiolata, utrinque hirto-pilosa, subtus ad nervos hispida. Petioli pedunculique fusco-hispido-villosi. Calyx 5-sepalus; sepalis ellipticis, acutis, praecipue ad marginem hirto-villosis. Corollae tubo brevi, glabro, quam sepala parum breviore limbo rotato, 5-lobo, intus atrofusco, extus pallido, utrinque breviter hirtulo; lobis ovatis, obtusis. Coronae stamineae foliola tubo corollae adnata supra incurva, subquadrata, angulo interiore in laminam obtusa antherae incumbentem exeunte, Affinis H. lacunae Ham. (Dregia volubilis Benth, gen. pl. 775 Hooker fl ind. tom. X.* p. 46), villosior autem, corollae limbo atro-fusco lobis rotundatis, corona staminea tubo adnata etc.—Haage et Schmidt specimen siccum misit. * Hoyas in Flora of British India are in Volume 4 not 10 as quoted here (Asclepiadaceae 45). Translation: Climbing, stalk dark brown, with hispid shaggy hairs. Leaves membranaceous, penninerved, reticulately veined, cordate ovate, acuminate, petiolate, both sides with long ascending hairs, hairs below on the nerves. Petioles and pedicels also with long hispid hairs. Calyx of 5 sepals; sepals elliptic, acute, with stiff bristly hairs especially on the margines. Tube of the corolla short, glabrous, not as short as sepals, limb rotate, 5 lobed, inside dark brown, outside pale, both sides briefly hirsute; lobes ovate obtuse. Leaflets of the staminal corona adnate to the tube of the corolla incurved above, somewhat four-sided, interior angle blade obtuse, anthers incumbent on the end. Allied to H. lacuna Ham, (Dregia volubilis Benth. plate 775, Hooker in Flora of British India tom. X p.46), villose but nevertheless the limbs of the corolla lobes rounded dark brown, staminal corona adnate to the tube etc. Haage & Schmidt specimen looks different when dried. In Dictionary of Gardening Century Edition (1901) 430 George Nicholson. H. gonoloboides (Gonolobus-like). fl. brownish, rotate, with ovate, obtuse lobes, umbellate; peduncles hispid. l. membranaceous, cordate-ovate, acuminate, hairy on both sides. Stem fulvous-hispid, climbing. India (?) 1884. A distinct plant. In The Royal Horticulture Society Dictionary of Gardening (1965) 1016. (1st Ed. 1951). H. gonoloboides. Climber with reddish-brown bristly hairs. l. cordate-ovate, slenderpointed, rather thin, hairy on both sides. fl. brownish, rotate; lobes ovate, obtuse; umbels n bristly hairy peduncles. India? 1884.

Hoya gracilipes Schlechter In Nachtrage zur Flora der Deutschen Schutzgebeite in der Südsee (1905) 363 K.

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Schumann & K Lauterbach. H. gracilipes Schlechter n. sp. Epiphytica in truncis arborum, alto scandens ramosa; ramis ramulisque filiformibus, flexuosis, teretibus, glabris, laxe foliatis, interdum radicantibus; foliis patentibus patulisve lanceolatis vel el1iptico-lanceolatist acuminatis; utrinque glabris, textura coriaceis, breviter 7.5—10 cm longis, infra medium 2—3 cm latis, petiolo 0.4—0.7 cm longo; pedunculis, gracillimus teretibus vulgo 4.5—5.5 cm 1ongis, glabris, rhachide paulo incrassata; pedicellis filiformibus, glabris, ca. 2 cm longis; umbellis 8—12-floris; calycis segmentis oblongis obtusis, margine minute ciliatis, caeterum glabris, ca. 1.5 mm longis; corolla rotata 1.2 cm diametiente, usque ad medium 5-lobata, extus glabra intus dense et brevissime granuloso-puberula, lobis ovatis subacutis, margine recurvis, 0.5 cm longis; corona carnosa foliolis horizontalibus, ovatis obtusis, apice breviter et acute rostatis, medio longitudinaliter sutbcarinato-incrassatis, subtus (dorso) ovatis, longitudinaliter sulcatis; antheris 1ate trapozoideis, appendice hyalina ovata breviter acuminata; pollinis oblongis dimidio longitudinaliter incrassatis, translatoribus brevibus, retinaculo obtuso, basi exciso, polliniis multo minora; stigmatis capite breviter conico. Kaiser Wilhelmsland; Auf Bäumen in den Wäldern des Torricelli-Gebirges, alt. ca. 1100 m (R. Schlechter, n. 14478, bl. April 1902). Eine sehr characteristische Art, die sich durch den sehr schlanken Habitus, wie durch die dünnledrigen Blätter und die äuszerst schlanken Stiele der Blütendoldea vor allen anderen Arten leicht unterscheiden 1äszt. Translation: same as in “Dr. Schlechter’s Hoya Species” below. In Botanische Jahrbücher (1908) 120-121. R. Schlechter. (Engler’s). 26. H. gracilipes Schltr., in K. Schum. u Lauterb., Nactr. (1905 p. 363). Nordöstl. Neu-Guinea: auf baümen in den Wäldern des Torricelli Geberges, ca 800—1100 m ü M. (Schlechter n. 14478. — Blühend im April 1902; n. 20213.— Blühend im September 1909). Hier liegt weider ein typisches Beispiel dafür vor, wie lokal viel Arten breitet sind. Gerade auf dem Torricelli-Giberge Habe ich viele Arten wiedergefunden, welch ich früher dort in anderen Gegenden gesammelt hatte, die aber auf den übrigen Gebigen zu fehlen scheinen. H. gracilipes Schltr. schlieszt sich am nächsten an H. solaniflora Schltr., ist aber durch die weiszen Blüten und die viel schmäleren, auszen dast spitzen koronaschuppen verschieden. Translation: see entry below. In Hoyas of Northeastern New Guinea (1992) 53-54. R. D. Kloppenburg. (Translation of Die Asclepiadaceen von Deutsch-Neu-Guinea. 26. Hoya gracilipes Schlechter in K. Schumann and Lauterbach, Addendum (1905) p.363. Northeastern New guinea: on trees in the forest of the Torricelli Mountains about 800-1100 m. altitude (Schlechter #14478 - Blooming in April 1902; #20213 - Blooming in September 1909). Here again lies a typical example of how many species are locally disseminated. In the Torricelli Mountains I have again found many species which I earlier had collected in different regions, but which did not appear to occur on the remaining mountain chain. H. gracilipes Schlechter is nearest to H. solaniflora Schlechter but is different in its white blooms and the much diminished corona scales, and the corona

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scales that are much narrower and pointed on the outside (outer apex). In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. gracilipes Schlechter (Schlechter 14478) – 14. In Dr. Schlechter’s Hoya Species (1993) 70-71. R. D. Kloppenburg. Hoya gracilipes Schlechter 1905 Nachtrage zur Flora der Deutschen Schutzgebeite in der Südsee p. 363 (K. Schumann und K. Lauterbach). Epiphytic on the trunks of trees, high climbing, branched, branches rebranched, threadlike, flexible, round, glabrous, loosely leaved, occasionally rooting; leaves spreading outspread, lanceolate or elliptic-lanceolate acuminate, both sides glabrous, texture leathery, shortly petiolate, 7.5-10 cm. long, below the middle 2-3 cm. wide. petiole 0.4-0.7 cm. long; with the pedicels very slender. round, glabrous, about 2 cm. long; with umbels 8-12 flowered; with the segments of the calyx oblong obtuse, margins minutely ciliate, otherwise glabrous, about 1.5 mm. long; corolla rotate 1.2 cm. in diameter, 5 lobed all the way to the middle, outside glabrous, inside densely and briefly granulous-puberulous, lobes ovate somewhat acute, margins recurved, 0.5 cm. long; corona fleshy, scales horizontal, ovate obtuse, apex short and acutely beaked, in the middle longitudinally somewhat keeled thickened, on the back ovate, longitudinally sulcate, with the anthers broadly trapezoidal, with the appendages hyaline ovate briefly acuminate; with the pollinia oblong gradually longitudinally thickened, translators small, retinaculum obtuse, base excised, many times smaller than the pollinia, with the head of the stigma conic. Northeast New Guinea (Kaiser Wilhelmsland): On trees in the forest of the Torricelli Mountains, altitude about 1100 meters (R. Schlechter #14478 flowering in April 1902). A very characteristic species distinguished in its slight appearance from any other species through its very slender habit and through the very thin leaves and narrowly slender peduncles of the umbel.

Hoya gracilis Schlechter In Beiblatt zu den Botanischen Jahrbücher 40 #92 (1908) 14. R. Schlechter. Hoya gracilis Schltr. n. sp.; epiphytica, gracilis, volubilis, alte scandens, ramosa; ramis ramulisque filiformibus elongatis, flexuosis, glabris, laxe foliatis, radicantibus; foliis patulis breviter petiolatis, lanceolato-ellipticis acutis vel obtusiusculis, basi cuneatis, glabris, lucidis, textura crasse coriaceis, 5.5—7.5 cm longis, medio fere 1.8—2.3 cm latis, petiolo crassiusculo 0.5—1 cm longo, glabro; cymis umbelliformibus, graciliter pedunculatis, 20—30-floris, pedunculo tereti glabro, 4—6 cm longo, pedicellis filiformibus 5.9—2 cm longis; floribus illis H. lacunosae Bl. fere aequimagnis; calycis segmentis oblongis obtusis, obtusis, vix 1.5 mm longis; corolla subrotata recurvula c. 0.7 cm diametiente usque infra medium 5-lobata, extus glabra, intus minutissime papilloso-puberula, lobis ovatis breviter acuminatis; phyllis apicem et dorsum versus adscendentibus, superne elliptico-lanceolatis antice breviter rostratis antheris paululo brevioribus, subtus oblongis dorso in lobules 2 adscendentibus obtusis, partem superiore phylli bene superantibus productis; antheris trapezoideis, appendice hyalino ovato;

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polliniis oblique oblongoideis, translatoribus subnullis, retinaculo rhomboideo minuto; stigmatis capite conico; folliculis gracilibus fusiformibus glabris, in rostrum sensim productis, c. 11 cm longis, medio fere 0.3 cm diametientibus. Celebes: bei Talisso (ohne Name des Sammlers [Eingeborene?], blühend im Juni 1886). Ich erhielt die Pflanze als. Hoya lacunosa., von der sie sowohl in der Form der längeren Blätter als auch in der Korona erheblich abweicht. Translation: See below in “Hoya Section Acanthostemma (Blume) Kloppenburg. In Leaflets of Philippine Botany 10 (1938) 3581. A. E. D. Elmer. Hoya Gracilis Schltr. Forming mats upon branches of Quercus pruinosa Blm. growing in damp woods at 4000 feet elevation; stems rigid, numerously branched, green but ultimately turning brown; leaves spreading, rigid, thick, pale or yellowish green beneath, with age becoming softer and wrinkled, deep green and grooved upon the upper side, occasionally turning reddish; inflorescence upon descending terminal stalks; pedicels curvingly deflexed, nearly red; flowers pendant, deeper red, odorless and also rigid. Aputulibung in Bagobo. Number 10482 collected by Elmer at Todaya.—This determination may not be correct when compared with the type of Schlechter’s Celebes plant. In An Enumeration of Philippine Flowering Plants 3 (1922) 352. E. D. Merrill. Hoya gracilis Schltr. in Engl. Bot. Jahrb. 50* (1908) Beibl. 92: 14. Luzon (Sorsogon), Mindoro, Panay, Merrill 56553, 5673, B. S. 32378 McGregor, 15703 Fenix, F. B. 6874 Merrill, Elmer 16719 as H. obscura Elm. In primary forests on ridges extending into the mossy forest, altitude about l000 m. Celebes. Local name: Aputulibung (Bag.). In An Enumeration of Philippine Flowering Plants 4 (1923) 97 E. D. Merrill. Hoya gracilis Schltr. (H. imbricata and H. pseudomaxima Koord., with one Celebesian species form a subgenus confined to Celebese and Philippines). In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. gracilis Schlechter (Type without name & number) – 14. In Dr. Schlechter’s Hoya Species (1993) 72-73. R. D. Kloppenburg: From Botanische Jahrbücher 40 #92 (1908) 14, (Beiblatt zu den). Hoya gracilis Schlechter. Epiphytic, slender, twining, high climbing, branched; branches rebranched, threadlike, elongate, flexible, glabrous, loosely leaved, with adventitious roots (rooting); leaves outspread, briefly petiolate, lanceolate-elliptic acute or somewhat obtuse, with the base wedge-shaped glabrous, lucid, texture thickly leathery, 5.5-7.5 cm. long, near the middle 1.8-2.3 cm. wide, with the petiole moderately thick 0.5-1 cm. long, glabrous; with cymes shaped like umbels, narrowly pedunculate, 20-30 flowered, with the pedicels threadlike 1.2-2 cm. long; flowers similar to H. lacunosa Blume, nearly the same size; with the segments of the calyx oblong obtuse, glabrous, scarcely 1.5 mm. long; corolla somewhat rotate recurved, about 0.7 cm. in diameter, 5 lobed all the way to below the middle, outside glabrous, inside minutely papillose-puberulous, with the lobes ovate shortly acuminate; with the scale of the corona, tip and outer part, turned upward, above elliptic-lanceolate with the tip shortly beaked, with the anthers a little shorter, below

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along the outer side are two ascending lobes, obtuse the superior (upper) part of the scale projected well above; with the anthers trapezoidal, appendages ovate, hyaline; with the pollinia obliquely oblong, translators almost lacking, retinaculum minute rhomboid; with the head of the stigma conic; with the follicles narrowly spindle shaped, glabrous, produced gradually into a beak about 11 cm. long, in the middle almost 0.3 cm. in diameter. Celebes: near Talisso (by a native named Sammlers) blooming in June 1886. I present this plant in the character of (comparison) of Hoya lacunosa. It is divergent in the form of the longer leaves as well as in comparison somewhat of the corona. In Hoya Section Acanthostemma (Blume) Kloppenburg (1994) 48-55. R. D. Kloppenburg. Hoya gracilis Schlechter in Engler's Botanische Jahrbücher 40 Beibl. 92 (1908) 14. Type: Sammlers s.n. June 1886, near Talisso, Celebes (B). Epiphytic, gracilis, volubilis, alte scandens, ramosa; ramis ramulisque filiformibus elongatis, flexuosis, glabris, laxe foliatis, radicantibus; foliis patulis breviter petiolatis, lanceolato-ellipticis acutis vel obtusiusculis. basi cuneatis, glabris, lucidis, textura crasse coriaceis, 5.5-7.5 cm. longis, medio fere 1.8-2.3 cm. latis, petiolo crassiusculo 0.5-1 cm. longo, glabro; cymis umbelliformibus, graciliter pedunculatis, 20-30 floris, pedunculo tereti glabro, 4-6 cm. longo, pedicellis filiformibus 1.2-2 cm. longis; floribus illis H. lacunosae Bl. fere aequimagnis; calycis segmentis oblongis obtusis glabris, vix 1.5 mm. longis; corolla subrotata recurvula c. 0.7 cm. diametiente usque infra medium 5-lobata, extus glabra, intus minutissime papilloso-puberula, lobis ovatus breviter acuminatis; coronae phyllis apicem et dorsum versus adscendentibus, superne elliptico-lanceolatis antice breviter rostratis, antheris paululo brevioribus, subtus oblongis dorso in lobulos 2 adscendentibus obtusis, partem superiorem phylli bene superantibus productis; antheris trapezoideis, appendice hyalino ovato; polliniis oblique oblongoideis, translatoribus subnullis, retinaculo rhomboideo minuto; stigmatis capite conico; folliculis gracillibus filiformibus glabris, in rostrum sensim productis, c. 11 cm. longis, medio fere 0.3 cm. diametientibus. Celebes: bei Talisso (ohne Name des Sammlers (Eingeborene ?), blühend im June 1886). Ich erhiet die Pflanze als Hoya lacunosa von der sie sowohl in der Form der längeren Blätter als auch in der Korona eheblich abweicht. Translation: of Latin: Epiphytic, slender, twining, lofty climber, branched, branches rebranched, threadlike elongated, flexible, glabrous, loosely leaved, rooting; leaves shortly petioled spreading, lanceolate-elliptical acute or somewhat obtuse, base cuneate, glabrous, lucid, texture thick leathery, 5.5-7.5 cm long, in the middle almost 1.8-2.3 cm. wide. petiole somewhat thicker 0.5-1 cm. long, glabrous; cymes shaped like umbels, slenderly pedunculate, 20-30 flowered, peduncle round, glabrous, 4-6 cm. long, pedicels threadlike 1.2-2 cm. long; flowers similar to Hoya lacunosa Blume,* almost the same size; segments of the calyx oblong obtuse, glabrous, barely 1.5 mm. long; corolla somewhat rotate recurved, about 0.7 cm. in diameter, 5 lobed all the way to the middle, outside glabrous, inside minutely papillose-puberulous, lobes ovate briefly acuminate; leaflets (scales) of the corona with the apexes and back ascending , above ellipticlanceolate with the front shortly beaked, anthers a little shorter, below oblong, below with 2 lobes obtusely ascending, the part of the superior leaf (scale) well overtopping (longer

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than the anther); anthers trapezoid, appendages ovate, hyaline; pollinia obliquely oblong, with the translators almost lacking, retinaculum rhomboid very small; with the stigma head conical; follicles (seed pods) narrow, fusiform (narrowed toward both ends) glabrous, produced into a visible beak, about 11 cm. long, in the middle about 0.3 cm. in diameter. Celebes: near Talisso (without name from Sammlers (A native?) blooming in June 1886. German: I received this plant as Hoya lacunosa, from which it differs considerably both in the shape of the longer leaves as well as in the corona. References: Elmer in Leaflets of Philippine Botany 1 May 1938. 10 Art. 131:3581. Merrill in Enumeration of Philippine Flowering Plants 3 (1923) 352. Schlechter in Engler's Botanische Jahrbücher 40 (1908) beibl. 92:14. Merrill in Enumeration of Philippine Flowering Plants 4: 97. Kloppenburg in Philippine Hoya Species (1991) 49. Herbarium Sheets: Felix #15703 1903 Philippines; Wenzel #2p 1928 (UC) Mt. Bulusan, Luzon, Philippines; Mendoza #41905 1961 (PNH) Tengao, So. Mindanao. McGregor #32378, #52372 1918 (BO) Antigua, Panay, Philippines; Elmer #10482 1904 (BO) Todaya Is, Panay, Philippines; Sulit #9894 1949 (PNH) Mt. Katanglad, Mindanao, Philippines; Merrill #5679, #5673, 1906 (B) Panay, Philippines, #5653 1906 (B) Mt. Halcon 900 m., Mindoro, Philippines; Edano #7404 1946 (PNH) Cuerenos De Negros, Negros, Philippines; #333 1947(PNH) Bacungan, Palawan, Philippines; #14203 1951 (PNH) Victoria Mts., Palawan, Philippines; Ramos & Edano #45478, #45477 #45570 1925 (UC) Casiguran, Luzon, Philippines; Britton #19519 1953 (PNH) Badoc Calopan, Midanao, Philippines; Allan #150190 1981 (PNH) Bigio Cove, Philippines; Fox #5023 (PNH) 1948 Dibutunan, Quezon, Luzon, Philippines; Ramos & Convocar #83890 (PNH) Dinaget Is., Mindanao; Merritt #6847 Philippines. *Discussion: The flowers of this section do, in general, look like those of Hoya lacunosa Blume (Type of the Section Otostemma (Blume) Miquel), as Schlechter notes. Schlechter also rightfully noted in German that the corona is considerably different. The corona is boat shaped and thin and not sulcate below as in this section. The most distinctive feature of the Section Otostemma (Bl.) Miquel is the annulus-like skirt below the crown surrounding the short column. Varieties and subspecies: A. D. E. Elmer in Leaflets of Philippine Botany 10 (1938) 358 describes a Philippine specimen as follows: Hoya gracilis Schlechter. Forming mats upon branches of Quercus pruinosa Bl. growing in damp woods at 4000 feet elevation; stems rigid, numerously branched, green but ultimately turning brown; leaves spreading, rigid, thick, pale or yellowish green beneath, with age becoming softer and wrinkled, deep green and grooved upon the upper side, occasionally turning reddish; inflorescence upon descending terminal stalks; pedicels curvingly deflexed, nearly red; flowers pendent, deeper red, odorless and also rigid. Apuntulibung in Bagobo. Number 10482 collected by Elmer at Tadaya.-- This determination may not be correct when compared with the type of Schlechter's Celebes plant. Comments: Elmer’s sheet #10482 at (BO) has relatively narrow long leaves, certainly not conforming to Schlechter's Celebes type for H. gracilis. In addition there is a sheet #32378 collected by McGregor in 1918 at Antigua Province, Panay, Philippines, labeled H. gracilis var. philippinensis with foliage narrowly ovate, most with rounded bases and acuminate apices. The foliage is very different from Elmer #10482 and also different

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from Schlechter's H. gracilis. I have not found a single herbarium sheet of Philippine hoya species that appear to be a variety of H. gracilis Schlechter. Many sheets labeled as varieties of this species are not even in the Section Acanthostemma (Bl) Kloppenburg. For example sheet #14203 (1951) PNH from Victoria Mts. Palawan; Ramos & Edano #45478, #455477 (UC): Britton #19519 (1953) PNH; Fox #5023 PNH; Ramos & Convocar #93890 PNH Dinaget Is. Mindoro all fall into this category. It is doubtful in my opinion that any of the herbarium material collected in the Philippines is this species or even a variety of this species. I have yet to examine a fertile Philippine herbarium sheet, including all those housed at PNH, CAHP, BO or UC, that is even in the same section. All the sheets with flowers can be determined but until DNA typing can be carried out on sterile sheets we will be able to identify these with certainty. There may be more than one species involved in this Philippine collected material. Most of what I have examined belongs in the section Otostemma (Bl.) Miquel, and is an undescribed species. David Liddle in Hoyan 13 (1991) pt. 2 3 published a Hoya gracilis ssp. philippinensis which was collected by David M. Cumming (DMC #8283) 1.5 km from the Lake Bulusan turn off towards the lake at the edge of the rainforest, when coming from Irosin. I judge the elevation here to be about 500'. Described as follows: Slender, twining, much branched climber. Stems: filiform, flexible and glabrous, frequently rooting at the leaf nodes. Leaves: spreading, shortly petiolate, lanceolate-elliptic, cuneate, glabrous, thick and coriaceous, up to 7.5 cm. long and 2.5 cm. wide. Petiole: .5 - 1.2 cm. long, glabrous. Petioles tend to be brownish, whereas the leaves are green. Inflorescence: 10-30 flowers in an umbel. Pedicel: glabrous 3-6 cm.. Calyx lobes: oblong obtuse, glabrous. 1.5 mm. long. Corolla: strongly recurved at the tips 1.2 cm. in diameter, apexes ascending, elliptic-lanceolate, at the outer end two ascending lobes. Color: Corolla pink, corona scales yellow with inner end red. This species is in commerce as IML 892. Discrepancies with Hoya gracilis Schlechter: Schlechter's flower about 0.7 cm. in diameter; Liddle's 1.2 cm in diameter. The former inner corona lobes "well overtopping the anthers" not true of IML 892. The greatest difference I find is in the pollinarium. Schlechter says "pollinia obliquely oblong with translators almost lacking" "retinaculum rhomboid , very small" (as he has drawn them). This would be very unusual for a species of this section. IML 892 has a very noticeably different pollinarium with very pronounced translator arms, bowl shaped with large caudicles. The pollinia are bulky, wide and somewhat contorted. The retinaculum also is large, bifid and spreading at the outer apex and covered with a clear membrane. (see drawing on following pages). I have photographed this type of pollinarium on other unidentified (and undescribed) Philippine herbarium sheets. It is slightly similar to the commercial Hoya sp. Ceram, sold as Hoya gracilis Schlechter, but certainly not identical to it. At this time I do not feel this is a subspecies of Hoya gracilis Schlechter. I believe it should be written up by David Cuming or David Liddle as a new Philippine species. Although there are unidentified Philippine herbarium sheets with pollinarium like IML 892 none are labeled as a variety or subspecies of the Celebes species. One additional fact should be noted. This taxa has a thin spatulate inner coronal lobe which places it near the species H. rizaliana and H. loheri. Like the later on drying these lobes turn outward from the axis at 90°.

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Hoya grandiflora Sprengel In Sprengel System Vegetabilum 1 (1825) 843. H. grandiflora* 3. H. foliis cordatoovatis acutis ramisque, pubercentibus, umbellis subsessilibus paucifloris. Nov. Holland. Translation: Hoya leaves cordate-ovate acute branches very pubescent, Umbels almost sessile, with few flowers. From New Hollandia. In A General System of Gardening and Botany 4 (1837) 126. G. Don. 1. T. grandiflora (R. Br. prod. p. 406.) Umbels nearly sessile, simple, few-flowered; pedicels glabrous; leaves cordate-ovate, acute, and are, as well the branches, downy. Woody perennial shrub. Native of New South Wales, about port Jackson. Hoya grandiflora, Spreng. syst. 1. p, 843. Great-flowered Tylophora. Fl. Ju. Jul. Cult. 1822. Sh. tw. In Flora Australiensis 4 (1867) 334. G. Bentham. 3 T. grandiflora, R. Br. 460. A rather slender twiner, the stems and foliage pubescent. Leaves on slender petioles, ovate to ovate-lanceolate, shortly and acutely acuminate, more or less cordate, 1 to 2 in. long. Flowers large for the genus, purple, on filiform pedicels of ½ to 1 in., usually only 1 or 2 rarely 3 on a short interpetiolar peduncle. Calyx-segments ovate, about 1 line long. Corolla spreading to about 1 in. diameter, the lobes ovate-oblong obtuse. Gynostegium small. Corona-segments thick, rounded, horizontally spreading and united in a ring at the base. Follicles acuminate, about 3 in. long. Seeds broad.—Dcne. in Prod. viii 612; H. grandiflora, Spring. Syst. i. 843. N. S. Wales Hunter’s River, R. Brown, also Herb. F. Mueller, but the collector not indicated. In Queensland Flora (1900) 1006. F. M. Bailey. 2. T. grandiflora (Large flowered), R. Br. Prod. 460: Benth Fl. Aust. iv. 334. A rather slender twiner, the stems and foliage pubescent. Leaves on slender petioles, ovate to ovate-lanceolate, shortly and acutely acuminate more or less cordate, 1 to 2 in. long. Flowers large for the genus, purple, on filiform pedicels of ½ to 1 in., usually only 1 or 2 rarely 3 on a short interpetiolar peduncle. Calyx-segments ovate, about 1 line long. Corolla spreading to about 1 in. diameter, the lobes ovate-oblong obtuse. Gynostegium small. Corona-segments thick, rounded, horizontally spreading and united in a ring at the base. Follicles acuminate, about 3 in. long. Seeds broad.—Dcne. in Prod. viii 612; H. grandiflora, Spring. Syst. i. 843. Hoya grandiflora Blume ex Decaisne In Decandolle, Prodramus System Vegetabilum 8 (1844) 635. Decaisne. 2. H. grandiflora (Bl. mass.), scandens, foliis oblongis v. elliptico-oblongis basi rotundatis apice abrupte breviterque acuminatis supra pube molli raraque inspersis subtus dense tomentoso-velutinis rufis (an siccitate?). Affinis H. coronariae Bl bijdr. quae a corona Ariadnes Rumph. differt foliis subtus velutinis coriaceis nec carnosis glabrisque. climbing shrub in insula Java. (v.s. sine flor. h. M. p.) Translation: climbing, leaves oblong or elliptic-oblong base rounded apexes abruptly and shortly acuminate pubescent above somewhat soft and scattered below densely tomentose velvety reddish (When dried?). Allied to H. coronaria Blume in Bijdragen which differs

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from Corona Ariadnes Rumphius by having the leaves velvety below, coriaceous not fleshy glabrous. Climbing shrub on the island of Java. (I have seen it without flowers in the Paris Museum Herbarium.)

Hoya graveolens Kerr In Kew Bulletin (1939) 461 A. F. G. Kerr. Hoya graveolens Kerr (AsclepiadaceaeMarsdenieae); H. ovalifoliae Wight et Arn. affinis, foliis crassioribus, nervis obscuris distinguenda. Suffrutex volubilis; rami subquadrati, cortice lenticellis rotundis elevatis asperso obtecti. Folia oblongo-elliptica vel oblongo-oblanceolata, basi late cuneata vel rotundata, apice breviter acuminata, margine plana, 5-8.3 cm. longa, 2-3.8 cm. lata, coriacea glabra, costa supra impressa subtus subplana, nervis lateralibus 4-5 paribus angulo 60° ascendentibus saepius obscuris rarius aegre aspectabilibus; petiolus 4-6 mm. longus, crassus, supra concavus, glaber. Inflorescentia lateralis, umbelliformis, pedunculata, 812-flora; pedunculus sat robustus, 1.8-3.8 cm. longus, glaber; rhachis crassa, accrescens, usque 5 cm. longa; pedicelli graciles, glabri, vel apicem versus pilis paucis obsessi, l.4-l.8 cm longi. Flores graveolentes. Calyx 5-lobatus, extra parce hirsutus; intus glandulis minutis praeditus; lobi anguste ovati, ciliati, circiter 2 mm. longi, basi 1 mm. lati. Corolla alba rotata, extra glabra, intus puberula; tubus explanatus, 3 mm. longus; lobi ovati, acuminati, 5.5 mm. longi, basin versus 5.5 mm. lati. Coronae segmenta rosea, distaliter paullum subrecta, lateraliter leviter compressa, dorso convexa, 3 mm. longa, angulo externo obtusa, angulo interno in dentem acutum erectum producta. Antherae appendix coronae aequilonga; pollinia oblonga, corpusculo apice acuto multo longiora. Stigmatis caput disciforme, apice apiculatum, apiculo minute bifido. Folliculi 8-10 cm. longi, apicem versus sensim attenuati, siccitate leviter striati, vivo flavi nigro-purpureomaculati; semina circiter 5.5 mm. longa, 1 mm. lata, compressa minute punctata, apice comosa, basin versus pilis longis paucis praedita. Sriracha, climbing on bushes near sea, flowers with an offensive small, Kerr 4245 (type); Sriracha, on trees by shore, Kerr 4138. The description of the fruit and seeds has been drawn up from Kerr 4138. Translation: Twining subshrub, branches almost 4 sided, cortical conceals appeased elevates lenticels. Leaves oblong-elliptic or oblong oblanceolate, base broad wedge shaped or rounded, apex briefly acuminate, margins flat, 5 to 8.3 cm. long, 2 to 4.8 cm. wide, leathery, glabrous, midrib above sunken below almost flat, lateral nerves 4 to 5 pairs often ascending with an angle of 60° difficult to observe; petioles 4 to 6 mm. long, thick, above concave, glabrous. Inflorescence lateral, in the shape of an umbel pedunculate 8 to 12 flowered; peduncles moderately robust, 1.8 to 3.8 cm. long; rachis thick, increasing up to 5 cm. long; pedicels narrow, glabrous, or the apex with a few hairs. , 1.4 to 1.8 cm long. Flowers strong smelling. Calyx 5 lobed, outside sparsely hirsute, inside provided with minute glands; lobes narrowly ovate, ciliate, nearly 2 mm. long, base 1 mm. wide. Corolla white, rotate, outside glabrous, inside puberulous; tube flattened, 3 mm. long; lobes ovate, acuminate, 5.5 mm. long, base nearly 5.5 mm. wide. Segments of the corona rose colored, a little away form the attachment almost straight , laterally lightly compressed, , convex above, 3 mm. long, exterior angle obtuse, interior angle acute erect tooth-like. Anther appendages of equal length of the corona.; pollinia

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oblong, corpuscle apex acute much longer. Stigma head disk shaped, apex apiculate, apiculum minutely bifid. Follicles 8 to 10 cm. long, toward the apex gradually attenuate, dried lightly lined, alive yellow with black purple spots; seeds about 5.5 mm. long, 1 mm wide, compressed, minutely punctate, apex comose, base with a few long hairs. In Florae Siamensis Enumeratio (1951) 36. W. G. Craib & A. F. G. Kerr. Hoya graveolens Kerr in Kew Bull. 1939. 461. Prachinburi. Sriracha, on rocks near the beach, Mrs. D. J. Collins 136 B ! 4245 (type !). A species near Hoya obtusifolia Wight et Arn., distinguished by its thicker leaves with obscure nervation. In The Taxonomy and Phytochemistry of the Asclepiadaceae in Tropical Asia (1995) 88. “The Genus Hoya in Thailand” O. Thaithong. 12. H. graveolens Kerr, Kew Bull. (1939) 418.* (Note: should be 416). Fl. Siam. Enum. 3 (1951) 36-37. Occurrence: )SE): Chon Buri. (SW): Prachuab Kiri Khan. Ecology: Climbing on bushes, trees and rocks near sea. Notes: Leaves carnosus, midrib grooved on upper surface, venation indistinct; umbel lax with 6-12 flowers; flower 10 mm dia. when fully expanded; corona white, rotatecampanulate, margin of lobe reflexed; outer end of corolla pale purplish pink, inside reddish purple.

Hoya greenii Kloppenburg In Fraterna 2nd Quarter (1995) 12-13 R. D. Kloppenburg. Hoya greenii Kloppenburg sp. nova. Affinis Hoya darwinii Loher a qua folium nervis perpendiculars cum coronae lobis interiore spatulatae, et pollinia margine exteriore differt. Cultivated at Green: Plant Research, Kaaawa Hawaii (ex clone collected by Ted Green), TG93132, Mt. Apo, Mindanao, Philippines at 4000’ elevation, on trail to hot springs lake, 30 October 1993. Holotype BISH. #7. Suffrutex, epiphyticus parum ramosus, scandens. Rami filiformibus, flexuosi, laxe foliati, teretes, glabri. Folia erecto-patentia vel patula, elliptica oblonga 15-20 cm. longa; 5-6 cm. lata, glabra, coriacea, nervis non conspicuis, breviter petiolata, umbellae multiflorae, flores magni 2 cm. diametere; corolla profunde lobia, collar prebrevi, lobi cordata-ovati, reflexi; coronae foliola erecta, magma triquetro-conica, subtus caniculata; styli mutico incumbentes, affinis Hoya darwinii Loher. An ornamental climbing vine with leaves similar to Hoya darwinii Loher but darker green (almost blackish) in color; more leathery and of only one type. In addition the veins of the leaves of this species are perpendicular to the midrib and finely reticulate, whereas those of Hoya darwinii Loher are at 45 degrees to the midrib, venation in neither is prominent. The flower size here is slightly smaller, the inner lobe is more spatulate than dentate (as Hoya darwinii’s is). The pollinia on this new species has a full sterile pellucid edge, differing markedly from H. darwinii’s rather short rudimentary sterile edge. The persistent peduncle bears approximately 50 buff pink with red centered flowers on equal pedicels; creating a hemispherical umbel. The sap is milky white. Stem: Terete, glamorous, 0.4-0.5 cm. in diameter, green lenticeled, becoming woodycorky; internodes ca. 3-4 cm. long but varying widely in length.

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Leaf blade: 12-15 cm. long x 3-4 cm. wide, narrowly elliptic, both surfaces glabrous, dull, very dark green, above slightly channeled along the midrib area, lighter green below. 10 +/- pairs of veins perpendicular to the midrib, somewhat obscure, with fine secondary reticulate venation with the base tapered acuminate, outer apex curved under acuminate, rigid. Petiole: Lighter green ca. 1.5 cm. long with definite groove above, otherwise terete. Peduncle: Axillary, persistent, 4 cm. long x 0.2 cm. wide. Pedicel: Straight, glabrous, terete, 2.7 cm. long x 0.10 cm. diameter, glabrous with fine granular lenticels, orange colored. Calyx: Ciliate (as opposed to H. darwinii Loher) cupped inward with slightly overlapping sepals, 0.12 cm. long widest 0.11 cm. , apex rounded, with distinct ligules present. Ovaries: Somewhat bottle shaped 0.22 cm. tall, rose apex otherwise pale lemon colored, base of pair 0,12 cm. wide. Corolla: Sharply reflexed with apex revolute, outside glossy glabrous, inside with a very fine pubescence, with long hairs under the corona around the collar area. Deeply lobed. Sinus- sinus 0.40 cm., sinus- apex 0.85 cm., apex- center 1.15 cm. thus making the flattened flower 2.30 cm in diam., widest 0.60 cm just outward from the sinus. Collar (tube) 0.05 cm. tall, opening 0.15 cm. Corona: Inner lobe narrowly spatulate touching in the center, reddish with very fine sulcations, outer lobe much raised, a very columnar corona. Outer lobe narrow but thick, fully channeled below, glossy waxy appearance except for inner lobes. Apex-apex (not flattened) 0.43 cm.; outer apex slightly hooked. Outer apex of anther wings protrude, anther wing- anther wing 0.23 cm.; retinaculum-retinaculum 0.12 cm; center to anther wing inner apex 0.14 cm.; retinaculum outer apex to anther wing apex 0.12 cm. (anther wing length). Stylar columnar with head subcapitate, with pentagonal table edges raised (thickened); fused stigmas distinct above. Pollinarium: Pollinia 0.074 cm. long rounded apices with long pellucid sterile edge and extending to outer curve where attached to caudicle; with a prominent vacuole. Translators narrow and long 0.18 cm x 0.028 cm., granular. Caudicle subclear, comma shaped with bulb 0.09 cm diam.. Retinaculum with distinct inner horns on a broad head 0.15 mm. wide; waist ca. 0.095 mm., 0.19 mm. long with 0.07 mm.; hip area at attachment of Translators and caudicle 0.09 mm. wide. Follicle: Linear lanceolate with a truncate outer apex, 9 x 0.8 cm, which bear approximately 150 seeds. I take great pleasure in naming this hoya species for its collector Ted green. Ted has collected extensively over a long period of time. He is responsible for the introduction of more hoya species and clones into commerce than any other single collector. His collecting has been carried out in the Fiji islands, Vanuatu (when it was called The New Hebrides), the Solomon islands, Indonesia, Singapore, Malaysia, Thailand, Borneo and the Philippines. Ted and I have been personal friends since childhood, a long fruitful relationship of more than sixty two years. Translation: Like Hoya darwinii Loher, from which it differs by the leaf nerves being perpendicular, with the inner coronal lobes spatulate, and pollinia exterior margins larger. Perennial epiphyte not very branched, climbing. Stems threadlike, flexible, loosely leaved, round, glabrous. Leaves erect spreading or outspread, elliptic-oblong 15 to 20 cm. long; 5 to 6 wide, glabrous, leathery nerves not conspicuous, shortly petioled,

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umbels of many flowers, flowers large 2 cm. in diameter, corolla definitely lobed, collar short, lobes cordate ovate reflexed; leaflets of the corona erect, large 3 angled-conic, below grooved; style without a point, incumbent, like Hoya darwinii Loher. In Philippine Hoya Species 3rd Ed. (1996) 76-77. R. D. Kloppenburg. Hoya greenii Kloppenburg in Fraterna 2nd quarter 1995. Type #7 BISH collected by Ted Green (TG #93123) Mt. Apo, Mindanao, Philippines 4000’ elevation, along trail to hot springs lake 30 October 1993. This is a ornamental climbing vine with leaves narrowly elliptic 12-15 cm. long x 3-4 cm. wide, narrowly elliptic, leathery in substance and very dark green, lighter green below. The leaf is slightly channeled along the upper midrib surface, glabrous, nerves almost perpendicular to the midrib, somewhat obscure, with fine secondary reticulations, the base tapered acuminate, outer apex curved under acuminate, rigid. Petiole lighter green ca. 1.5 cm. long with a distinct groove above otherwise terete. Peduncle axillary, persistent, 4 cm. long x 0.2 cm wide. Pedicels straight, glabrous, terete 2.7 cm. long x 0.10 cm. diameter, with fine granular lenticels, orange colored. Calyx ciliate, cupped inward with slightly overlapping sepals 0.12 cm. long, widest portion 0.11 cm., apex rounded, distinct ligules present at the base sinus areas. Ovaries somewhat bottle shaped 0.22 cm. tall, with rose apex otherwise pale lemon colored, base pair 0.12 cm. wide. Corolla sharply reflexed with apex revolute, outside glabrous, inside with a very fine pubescence, with long hairs under the corona around the collar, deeply lobed 2.30 cm. in diameter flat. Corona with an spatulate inner lobe which touch in the center, reddish with fine sulcations, outer lobe much raised, columnar in shape. Outer lobe narrow but thick, fully channeled below, glossy waxy in appearance except for the inner lobes 0.43 cm long. Pollinia 0.074 cm. long with rounded apices with long germinal mouth (pellucid edge), Caudicle comma shaped semi-clear, Retinaculum with distinct inner horns. Follicles linear lanceolate with truncate outer apex 9 x 08 cm. with ca. 150 seeds. Meaning: greenii — Named in honor of the collator Ted Green of Green Plant Research Kaaawa Hawaii.

Hoya griffithiana Decaisne In Flora of British India (1883) 53. J. D. Hooker. (Only reference here is under H. acuminata as H. Griffithiana, Dcne. mss.

Hoya griffithii Hooker In Flora of British India 4 (1883) 59 J. D. Hooker. 28. H. griffithii, Hook f.; glabrous, leaves 6-9 in. narrowly oblanceolate or linear-oblong acute or acuminate very thick, nerves nearly horizontal, peduncle long stout, sepals large coriaceous, corolla 1½ in. diam. glabrous within. Khasia Mts., Griffith; Nowgong, alt. 2-4000 ft. J. D. H. & T. T. A stout climber. Leaves distant, 1-2 in. diam., midrib stout, nerves quite obscure in some, in others distinct and forming large areoles; petiole ½ - 1 in.. very thick. Peduncle 2-4 in., as thick as the branch; pedicles 1-1 ½ in., very stout. Sepals ½- 1/3 in., oblong, obtuse, coriaceous, Corolla purple ? g1abrous within, 1obes acute. Coronal-

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processes produced upwards from a suborbicular base into an erect stiff obtuse spur as long as the anther-tip, the orbicular base is 2-lmellate below.— In the large calyx this approaches H. coronaria. It has been flowered at Fallowfield (Manchester) by Mr. Swan. In The Gardener’s Chronicle (1885) 334. N. E. Brown. Hoya Griffithii, Hook. f.* This very fine species was sent to me (through the Editor of the Gardeners’ Chronicle), in July, 1879. by Mr. W. Swan, of Fallowfield, Manchester, for a name. At that time it was undescribed, and it was intended to figure it in these columns, but a good specimen was not forthcoming, and now a specimen has been again received from the same source, and will be figured in the Botanical Magazine; but the plate may not appear for some time, I take the opportunity of calling attention to it here, as it is one of the finest species in the genus. It is a native of Khasia, in Northeastern India, where it was originally discovered by Griffith, and afterwards collected there at an attitude of 2000 feet, by Sir Joseph Hooker. Concerning this plant in cultivation, Mr. Swan, writing in 1879, states that it was sent with a quantity of Orchids, &c., about three years ago, to Mr. Leach, from " Northern India." Hoya Griffithii is a large climbing species, with oblong leaves 3 ½ —6 inches long, ¾—1 ¼ inch broad, narrowed to an acuminate apex and an acute base, glabrous, as is the entire plant, bright green above, paler beneath. peduncle stout, 2 inches long; pedicels 1 ½ inch long, stout. Calyx segments spreading, green, ½ inch long, 2—2 ½ lines broad, oblong obtuse, concave from the margins, being strongly incurved. Corolla when expanded about 1 ½ inch in diameter, pale reddish outside, cream colour, tinted with the same reddish hue inside, the lobes are broadly ovate, acute. Corona waxy-white, the segments elliptic in outline as seen from above, with ascending subulate apices, the sides are perpendicular with the upper edge overhanging at the back, and beneath at the base they have A- shaped notch. This species is at once well distinguished by its unusually large calyx segments. The rather delicate red tint of the corolla is somewhat difficult to describe intelligibly, but perhaps the nearest well known colour-tint is that of red blotting-paper diluted. N. E. Brown. * Hoya Griffithii, Hook. f. in Hook Florae Indica, vol., iv. In Botanical Magazine (Curtis’s) (1886) 6877. J. D. Hooker. Hoya (Euhoya) Griffithii; frutex glaberrimus, scandens, ramis elongatis flexuosis teretibus, foliis 9-9-pollicaribus per paria distantibus breviter petiolatibus oblongo-lanceolatis oblanceolatisve acutis acuminatisve crasse coriaceis, costa crassiuscula, nervis tenuissimis patentibus obscuris, pedunculis axillaribus crassis elongatis, umbellis multifloris, pedicellis pollicaribus crassis, floribus 1— l 1/4 poll. diam., sepalis lineari-oblongis obtusis crassis, corolla, sordide roseae lobis crassis triangulari-ovatis dorso concavis intus glabris, coronae processubus magnis ambitu hemisphericis faciebus planis in spinas obtusas stigma superantes productis. H. griffithii Hook. f. Fl. Brit. Ind. vol. iv. p. 59; N. E. Brown in Gard. Chron. 1885 vol. ii. p. 334, fig. 74 This noble species of Hoya was discovered by the indefatigable Indian botanist and collector, Dr. W. Griffith, in the Khasia Mountains of Eastern Bengal, between Assam and Silhet, upwards of half a century ago; and it was; again gathered there by Dr. Thomson and myself in 1860 near Nowgong, at an elevation between 2000 and 4000 feet

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above the sea. It is a near ally of the still nobler H. imperialis, Lindl., of Malacca, figured at Tab. 4397 of this work. A still nearer ally is the beautiful H. coronaria, Blume, a hitherto unfigured species of Malacca and Penang, which has pubescent leaves and flowers. Flowering specimens of H. Griffithii were sent to Kew by Mr. Swan, of Oakley, Fallowfield, near Manchester, in July but with the information that the plant had been at Oakley about ten years. Mr. Swan states that it came from Northern India, and was originally sent to hits employer, W. Leach, Esq. It is, of course, a stove plant, and as such well worthy of cultivation. Descr, A tall quite glabrous climber. Stem slender, terete flexuous. Leaves in distant pairs, four to ten inches long very shortly petioled, elliptic- or oblong-lanceolato. or oblanceolate, acute or acuminate, thickly coriaceous bright green above, paler beneath; midrib rather stout nerves slender, very obscure, horizontally spreading and loosely anastomosing; petiole stout, one-sixth to one third of an inch long. Flowers one to one: and a half inch in diameter, numerous, umbelled on a stout axiallary peduncle two to three inches long; pedicels long, stout, one to one and a half inches long. Sepals much larger in proportion to the corolla than usual in the genus, oblong, obtuse thickly coriaceous. Corolla quite glabrous, pale rather dull rose-red (the colour of blotting-paper externally with yellowish edges; paler and yellowish within, with three faint pink stripes on each segment. Segments triangular acute, thickly coriaceous. concave behind. Corona large with five hemispheric lobes; these lobes have the convex surface downwards and longitudinally cleft. the flat surface upwards and producing inwards a short ascending spine which reaches and rises above the stigma. J. D. H. Translation: a glabrous shrub, climbing, stems elongated flexible round, leaves 6 to 9 inches between the pairs briefly petiolate oblong-lanceolate oblanceolate acute acuminate thickly leathery, midrib thick, nerves very thin spreading obscure, peduncles axillary thick elongate, many-flowered umbels, many times thicker than the pedicles, flowers 1 to 1 ¼ inches in diameter, sepals linear-oblong obtuse thick, lobes of the corolla a dusty rose lobes thick triangular-ovate on the back concave inside glabrous, with the coronal processes large outline hemispherical general appearance flat with spines obtuse overtopping the stigma. In The Dictionary of Gardening Century Edition (1901) 430. G. Nicholson. H. Griffithii (Dr. W. Griffith’s).* fl. externally pale and rather dull rose red, with yellowish edges, paler and yellowish with three faint stripes on each segment, 1 in. to 1 ½ in. long. July. l. in distant pairs, 4in. to 10in. long, very shortly petiolate, elliptic or oblong-lanceolato or oblanceolate. Stem flexuous, climbing. Eastern Bengal, 1885. (B. M. 6877.) A fine species. In The Royal Horticulture Society Dictionary of Gardening (1965) 1016. (1st Ed. 1951). H. griffithii. Stem climbing, flexuose. l. distant, elliptic to oblanceolate, 4 to 10 in. long; stalk very short. fl. dull rose-red with yellowish edges without, paler and yellowish within with 3 faint pink stripes on each seg., 1 to 1 ½ in. long. July. E. Bengal. 1875. (B.M. 6877.) In The New Royal Horticulture Society Dictionary of Gardening 2 (1992) 596-597. H. griffithii Hook. f. Climber, stems twining. Lvs. 10-25cm. short-petiolate, oblanceolate to

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elliptic. Umbels many-fld; peduncles stout 2.5- 4.5cm; fls. 2.5-4cm.; cor. lobes dark rosered. with faint pink stripes; corona waxy-white. Summer, NE India to China Z9.

Hoya guppyi Oliver In Icones Plantarum 23 (1892) 2247. W. Hooker. A.—H. Guppyi, Oliv. in Guppy, 'Solomon Islands,' p. 298; ramulis ultiimis parce hirtellis deinde glabratis foliis petiolatis coriaceis late ellipticis breviter acuminatis cuspidutisve basi late rotundatis subcordatisve supra glabris v. fere glabris subtus plus minus hirtellis 1-costatis nervis lateralibus primiariis subtus utrinque 7-9, umbellis pedunculatis pedunculis pedicellisque glabris, calycis parvi corollae tubo 2-4-plo brevioris carnosuli 5-partiti lobis ovatis obtusis ciliolatis, corollae rotatae lobis patentibus ovatis v. late ovato-lanceolatis acutatis intus hirtellis extus glabris sinubus reflexis, coronae foliolis caltilagineo-incrassitis disco ovato-lanceolatis (sicco) concavis obtusis basi angustatis dorso profunde excavatis marginibus lateralibus utrinque carinatis, folliculis subteretibus longitudinaliter striatis parce hirtellis. Hab. Solomon Islands, Fauro Island, Bougainville Straits, H. B. Guppy (No. 188). Folia 3½-4 ½ poll. longa, 2 ¼ - 2 ½ poll. lata; petiolus hirtellus ½ - 1 poll. longus. Umbella 10-14- Flora; pednuculus 2 poll. longis; pedicelli 1-1 ½ poll. longi. Corolla 1-1 ½ poll. diam. rubro-purpurea. Follicula 8-9 poll. longa. Translation: ultimate branches moderately hairy thereafter becoming glabrous leaves petiolate leathery broadly elliptic briefly acuminate with sharp rigid point base rounded somewhat cordate above glabrous or nearly glabrous below more or less hairy leaf midrib with 7 to 9 lateral primary nerves prominent on the lower side, umbels pedunculate with peduncles and pedicels glabrous, calyx small 2 to 4 times smaller than the tube of the corolla fleshy 5 parted lobes ovate obtuse ciliate, lobes of the corolla rotate flat ovate or ovate lanceolate acute inside hairy outside glabrous sinus reflexed, leaflets of the corolla polished and cartilage-like disk ovate-lanceolate (when dry) concave obtuse base narrow back definitely hollowed out lateral margines on both sides keeled, follicles somewhat round longitudinally lined sparsely hirsute. Leaves 3 ½ to 4 ½ inches long, 2 ¼ to 2 ½ inches wide; petioles hairy ½ to 1 inch long. Umbels 10 to 14 flowered; peduncles 2 inches long; pedicels 1 to 1 ½ inches long. Corolla 1 to 1 ½ inches in diameter red-purple. Follicles 8 to 10 inches long. In Flora de Deutschen Schutzgebiete in der Südsee (1901). K. Schumann. H. Guppyi Oliver in Guppy, Solom. Isl. 298. Solomon-Inseln: Insel S. Cristoval (Comins). 1st endemisch.

Hoya gymnanthera Wight In Contributions to the Botany of India (1834) 37 R. Wight. 12. H. gymnanthera Wight; aphylla? caulis epidermide laevi nitida; umbellis multifloris parvifloris; cor. st. foliolis ovatis, antherarum membranaceis apicibus pellucidis angustissimis pollinis massas vix tegentibus. — Wall. ! Asclep. n. 142 (absque nomine). —.......; herb. Finlayson. The specimen of this plant which I have seen are leafless, but whether from

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accident or otherwise, I have not at present the means of ascertaining.- (R. W.) Translation: stalk epidermis smooth, shining, umbels many flowered. Flowers small’ corona scales ovate, anthers membranaceous, with the apexes of the pollinia masses transparent scarcely covered. In General System of Gardening and Botany 4 (1837) 126. G. Don. 14 H. gymnanthera (Wight, l. c.) leaf; epidermis of stem smooth and shining; umbels many-flowered, on short peduncles; flowers small; leaflets of corona ovate; tops of anthers membranous, pellucid, very narrow, hardly touching the pollen masses. woody shrub Native of the East Indies. Wall. ascl. no. l45, without a name. Naked-anthered Hoya. Shrub. tw. In Synopsis Plantarum 6 (1840) 891-892. D. N. F. Dietrich. 12. H. gymnanthera Wight; aphylla? caulis epidermide laevi nitida; umbellis multifloris parvifloris; cor. st. foliolis ovatis, antherarum ,membranaceis apicibus pellucidis angustissimis. In Ind. or. Perennial shrub. Translation: stalk epidermis smooth, shining, umbels many flowered. Flowers small’ corona scales ovate, anthers membranaceous. In Prodromus Syst. Veg. 8 (1844) 638. DeCandolle. 30. H. gymnanthera (Wight contr. p. 37). aphylla? caulis epidermide laevi nitido, pedunculis brevibus multifloris, floribus parvis, umbellatis, coronae stem. foliol. ovatis antherarum apicibus membranaceis pellucidis' angustissimis pollin, massas vix tegentibus. Woody Perennial In India orientali, loco uatali proprio ign. ex herb. Finlayson. Translation: Leafless? Stalk epidermis smooth and shining, peduncles short multiflowered, flowers small, umbellate, leaflets of the staminal corona ovate anther apexes membranaceous, transparent narrow pollinia masses, scarcely covered, Woody perennial in Eastern India location or origin not known. From Finlayson’s Herbarium. In Flora of British India 4 (1883) 62. J. D. Hooker. (Under doubtful or excluded species). H. gymnanthera, Wight. Contrib. 37; Wallich’s sheet of this consists of a leafless twig of a hoya with buds and a good specimen of Parsonsia spiralis.

Hoya hainanensis Merrill In Philippine Journal of Science 23 (1923) 263. E. D. Merrill. Hoya hainanensis sp. nov.. Herbacea, scandens, patulis intus parce pubescentibus exceptis glabri, ramis teretibus. 1.5 ad 2 mm diametro; foliis in siccitate chartaceis. ovato-ellipticis, penninerviis, oppositis, 6 ad 8 cm longis, 2.5 ad 4 cm latis, utrinque subaequaliter angustatis, apice acuminatis, basi acutis, nervis utrinque circiter 4, tenuibus, obscuris, subobliquis, reticulis laxis, inconspicuis; petiolo 8 ad 10 mm longo; umbellis circiter 20floris, pedunculo circiter 2 cm longo, pedicellis tenuibus, pedunculo subaequantibus; floribus albis, 8 ad 10 mm diametro, calycis lobis oblongo-ovatis, rotundatis, 1 mm longis, glabris, petalis oblongo-ovatis, acutis vel acuminatis, intus puberulis. Ng Chi Leng, McClure 9759, May 1922, on tree trunks, south slope of the mountain. A species belonging in the same group with an apparently closely allied to Hoya

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oblanceolata Cost., of Indo-China. from which it can be distinguished by its somewhat smaller flowers, the sepals rounded and glabrous, not acute and ciliate. Translation: Herbaceous, climbing, petals inside a little pubescent otherwise glabrous, branches round, 1.5 to 2 mm in diameter; leaves when dry papery, ovate-elliptic, penninerved, opposite, 6 to 8 cm long, 2.5 to 4 cm broad, both ends somewhat equally narrow, apex acuminate, base acute, nerves about 4 on both sides slender, obscure, almost oblique, reticulations loosely arranged, inconspicuous; petiole 8 to 10 mm long, pedicels thin, peduncle somewhat equal; flowers white, 1 mm in diameter, calyx lobes oblong-ovate, rounded, 1 mm long, glabrous, petals oblong-ovate, acute or acuminate, inside puberulous. In Sunyatsenia 3 #2/3 (1936) 172. Tsiang. Hoya hainanensis Merrill in Philip. Journ. Sc. 23: 263, May 1923; in Lingnan Sc. Journ. 5: 153. 1927. Hainan: Hung-mo Shan, Tsiang. Tang & Fung 166, May 13, 1929. Endemic. This is distinct from all other species by its penninerved leaves with characteristic narrow cuneate base. The base of the retinaculum is acute, not common in this genus. A Hainan plant with a showy pendulous inflorescence and ovate-elliptic leaves of which we have a beautiful photograph taken by Professor Chun in Hainan in 1920 might also belong this species. In Flora Kainantensis (1943) 267. Genkei Masamune. Hoya hainanensis Merr., in Philipp. Jopurn, Sci. XXIII. p. 263 (1923) et in Lingn.-J. V. p. 153 (1927); Groff, Ding & Groff, in Lingn.-R. II. p. 131 (1924); Tsiang, in Sunyt. III p. 175 (1936); Takaka et Odashima, in Trop. X. p. 379 (1938) .......... Script. In Flora Republicae Popularis Siamicae 63 (1977) 480. Hoya hainanensis Merr. in Philip. Journ. Sci. 23: 263. 1923, et Lingnan Sci. Jour. 5: 153. 1927; Tsiang in Sunyatsenia 3: 174, 1936. .... script 3: 172. 1974. .................paragraph of script.

Hoya halconensis Kloppenburg In Fraterna 3rd Quarter (1990) III. R. D. Kloppenburg. Publication of the International Hoya Association. Hoya halconensis Kloppenburg sp. nov. Type sheet #5674 collected by Elmer D. Merrill Nov. 1906 on Mt. Halcon, Mindoro, Philippines in a mossy forest at 900 m altitude. Flowers with faint odor, purplish outside, yellowish within. Berlin (B) Holotypus. This sheet is Schlechter's unpublished Hoya halconensis. ex. spec. Ramulis crassis 0.5-0.6 cm. diametro, glabris, internodiis elongatis 18 cm.. Foliis 9-12.5 x 4-6 cm. plerumque parvis, glabris, ovatis-ellipticis, apice breviter acute acuminatis, basi late rotundatis. Pedunculo 3.0 cm. longis, rachi incrassatis cir. 0.5 cm. diametro. Pedicellis hirsutis, filiformibus, teretibus. Calycis segmentis ovatus, hirsutis extis, 0.3 cm. longis, ligulea presentia. Corolla lobis late ovatus acutis ad apiculatis, profunde lobatus cir. 2 cm. diametro. Coronae foliolis, obovatus apice interne aliquantum breviter rostratis, externe subobtusa, medio longitudinaliter, inter apices carina donatis, subtus longitudinaliter foveolatis. Anthera apicem folioli excedente. Poliniis oblongoidiis, translatoribus linearibus, retinaculo rhomboides linealiter compressa. The foliage of this species is similar to that of H. mindorensis Schlechter although

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more broadly rounded at the base much like H. cardiophylla Merrill. It has the large pencil sized stems like H. angustisepala Burton. Its corona scales are elevated on the outer lobe more than H. cagayanensis Burton, but like this species it's outer lobes are blunt although the very apex of this species is somewhat more narrowly rounded. Like H. benguetensis Schlechter and has a similar longitudinal ridge on the upper surface of the coronal scale, extending from apex to apex.. It is, however, distinct from any of these species. I have retained Schlechter's area name. Translation: see below under Dr. Schlechter’s Hoya Species. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. halconensis Schlechter ex Kloppenburg (Merrill 5674) - +. In Dr. Schlechter’s Hoya Species (1993) 74-75. R. D. Kloppenburg. Hoya halconensis Kloppenburg. H. halconensis Schlechter (unpublished). Branches thick 0.5-0.6 cm. in diameter, glabrous, with elongated internodes 18 cm. (long). Leaves 9-12.5 x 4-6 cm, many smaller, glabrous, ovate-elliptic apex briefly acute, acuminate, with the base widely rounded, Peduncle 3.0 cm. long, rachis enlarged about 2.0 cm. long. Pedicels hirsute,, threadlike, round. With segments of the calyx ovate, hirsute, outside 0.3 cm. long with ligules present. Corolla lobes widely ovate, acute, nearly apiculate, definitely lobed about 2 cm. in diameter. With scales of the corona obtuse, inner apex somewhat briefly beaked, outer somewhat obtuse, longitudinally ridged down the center, inner apex with a keel, below longitudinally channeled. Anther apex extended. Pollinia oblong, translators linear, retinaculum rhomboid, laterally compressed. The foliage of this species is similar to that of H. mindorensis Schlechter, although more broadly rounded at the base much like H. cardiophylla Merrill. It has the large pencil sized stems like H. angustisepala Burton, like this species its outer coronal lobes are blunt although the very apex of this species is somewhat more narrowly rounded. Like H. bordenii Schlechter it has a similar longitudinal ridge on the upper surface of the corona scale extending from apex to apex. It is , however, distinct from any of these species. Mt. Halcon, Mindoro, Philippines in a mossy forest at 900 meters altitude (Merrill #5674) November 1906. Flowers with faint odor, purplish outside, yellowish within. In Philippine Hoya Species 3rd Ed. (1996) 78-79. R. D. Kloppenburg. Hoya halconensis Kloppenburg in Fraterna 1 #1 fall 1990. Type #5674 (B). Collected by Elmer D. Merrill Nov. 1906 on Mt. Halcon, Minoro, Philippines in a mossy forest at 900 m. altitude. Flowers with faint odor, purplish outside yellowish within. Scandent, glabrous with stems to lead pencil diameter to 0.7 cm., long internodes, 18 cm.. Foliage 9-12.5 cm. x 4-6 cm. with most small, spreading, glabrous, base broadly rounded, rather rectangular tapering slightly to the mucronate apex. Midrib visible above and below, lateral nerves obscure. Peduncle 3.0 cm. long, rachis perennial, enlarged about 0.5 cm. in diameter, rough and scaly. Pedicels long about 3.5 cm. Calyx lobes 0.3 cm. long hirsute, broadly overlapping at the bases, ligule present. Corolla broadly ovate with a drawn out sharply acute apex, glabrous inside and out, cut to below the middle, flower flattened about 2 cm. in diameter. Coronal scale outer lobe raised, saddle shaped,

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with a longitudinal ridge the full length on top, beneath fully channeled, 1.0 cm., crown 1.0 cm. diameter, outer apex of scale narrowly rounded, inner lobe short, drawn narrowly out, anther apex exceeding inner lobe. Staminal column short. Pollinaria with long narrow pollinia, long translators, broad at pollinia attachment, long narrow retinaculum, The collecting locality of this species is presumed to be the same as for the species Hoya paziae. Meaning: halconensis — named for the location of discovery, Mt. Halcon, Mindanao, Philippines.

Hoya halophila Schlechter In Botanische Jahrbücher 50 (1913) 107 R. Schlechter “Die Asclepiadaceen von Deutsch Neu Guinea”. 1. H. halophila Schltr. n. sp. — Suffrutex,, epiphyticus, ramosus, volubilis. Rami filiformes, flexuosi, bene foliata, teretes, glabri. Folia patentis vel patula., petiolata, late ovata vel suborbiculari-ovata, breviter acuminata, basi obtusissima, utrinque glabra, superne lucida, subtis opaca, carnosa. Cymae longius pedunculatae umbelliformibus, folia vulgo distincte superantia, 15—25 flora, pedicellis gracilibus glabris. Calycis segmenta ovata, obtusa, glabra, margine subinconspicue ciliolata, corolla multo breviora. Corolla revoluta usque ad tertiam partem basilarem 5-fida, lobis ovalis, acuminatis, extus glabris, intus apice glabrato excepto papulis reversis subtis cristalinis obtectis. Coronae foliola extus decurva, oblanceolato-elliptica, antice acuminata, postice obtusa subtus in auriculas 2 oblongas foliolum superantes extensa, antheris subaequilonga. Pollinia oblique oblongoideo-subfalcata, translatoribus brevibus triangulis, obliquis, retinaculo rhomboideo, minuto. Ein zierlicher Schlinger mit fadenförmigen Zweigen. Blätter 3.5—5 cm lang, unterhalb de Mitte 2.3—3.4 cm breit, Blattstiele bis 3—4 mm lang, fleshig. Blütenstände doldenartig auf 5—6 cm langen Stielen. Blütenstiele bis 1.3 cm lang. kelchzipfel ca 1.5 mm lang. Korolla ausegebrietet gegen 9 mm im durchmesser, Koronaschuppen mit dem Gynostegium einen brieten Kegel bildend, 3 mm lang. Nordöstl. Neu-Guinea: auf Bäumen in den Wäldern am Strande von Eitape, ca. 10 m ü. M. (Schlechter n. 19968. — Blühend im August 1909). Soweit man sich zur Zeit ein Urteil erlauben, ist die Art durch die sehr breiten Blätter von H. litoralis Schltr. sowohl wie auch von H. inconspicua Hemsley verscheiden. Ich habe allerdings bei Arten derselben Sektion von den Philippinen die Erfahrung gemacht, dasz die Blätter in irher Grösze und Breit stark variieren, doch sind hier die Unterschiede doch zu stark hervospringend. Es scheint mir auszerdem dasz das Gynostegium bei H. litoralis Schltr. Höher ist die Koronaschuppen Schmäler sind. Auszerdem sind die rötlichen Blüten heirr heller. Translation: See next below. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. halophila Schlechter (Schlechter 19968) – 15. In Dr. Schlechter’s Hoya Species (1993) 76-77. R. D. Kloppenburg. Hoya halophila Schlechter. An epiphytic branched, twining half-shrub. Branches threadlike, flexible, well leaved, round glabrous. Leaves spreading or outspread, petiolate, broadly ovate or

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somewhat round-ovate, briefly acuminate, base very obtuse, both sides glabrous, lucid above, opaque below, fleshy. Cymes long, peduncled, shaped like an umbel, generally longer than the leaves, 15-25 flowered, pedicels slender, glabrous. Segments of the calyx ovate, obtuse, glabrous, margins somewhat inconspicuously ciliate, much smaller than from the corolla. Corolla revolute, 5 parted all the way to one third from the base, lobes ovate, acuminate, outside glabrous, inside papillose, except the concealed wedge shaped terminal Scales of the corona decurved, outside oblanceolate-elliptic, inner (anterior) portion acuminate, outer (posterior) portion obtuse, two oblong ears extending beyond the outer apex, anthers almost equal. Anthers small somewhat acute. Pollinia obliquely oblong, almost sickle shaped, with the translators shortly triangular, oblique, retinaculum rhomboid, minute. An ornamental clinging vine with threadlike branches. Leaves 3.5- 5 cm. long, below the middle 2.3-3.4 cm. wide, petiole 3-4 mm. long, fleshy. Inflorescences well umbelled, with the peduncle 5-6 cm. long. Pedicels up to 1.3 cm. long. Calyx lobes about 1.5 mm. long. Corolla outspread about 9 mm. in diameter. Corona scales with gynostegium forming a broad cone, 3 mm. long. Northeastern New Guinea: on trees in the forest along the beach at Eitape, about 10 m. altitude (Schlechter #19968 - Blooming in August 1909). So far as one is allowed to make a judgment at the present time, this species with very wide leaves is different from Hoya litoralis Schlechter and from Hoya inconspicua Hemsley. I had the experience to be sure, with the species of the same section from the Philippines, in which the leaves vary greatly in their size and width, but here the dissimilarities are too vividly apparent. It seam to me moreover, that the gynostegium of Hoya litoralis Schlechter is higher and the corona scales are narrower. Moreover, the reddish blooms are brighter. Published by Dr. R. Schlechter in: 1913 Botanische Jahrbücher V.50 p.107, "Die Asclepiadaceen von Deutsch Neu Guinea ". In Hoyas of Northeastern New Guinea (1992) 96. R. D. Kloppenburg. (Translation from Die Asclepiadaceen von Deutsch Neu Guinea by R. Schlechter. Hoya halophila Schlechter. An epiphytic branched, twining half-shrub. Branches threadlike, flexible, well leaved, round glabrous. Leaves spreading or outspread, petiolate, broadly ovate or somewhat round-ovate, briefly acuminate, base very obtuse, both sides glabrous, lucid above, opaque below, fleshy. Cymes long, peduncled, shaped like an umbel, generally longer than the leaves, 15-25 flowered, pedicels slender, glabrous. Segments of the calyx ovate, obtuse, glabrous, margins somewhat inconspicuously ciliate, much smaller than from the corolla. Corolla revolute, 5 parted all the way to one third from the base, lobes ovate, acuminate, outside glabrous, inside papillose, except the concealed wedge shaped terminal scales of the corona decurved, outside oblanceolate-elliptic, inner (anterior) portion acuminate, outer (posterior) portion obtuse, two oblong ears extending beyond the outer apex, anthers almost equal. Anthers small somewhat acute. Pollinia obliquely oblong, almost sickle shaped, with the translators shortly triangular, oblique, retinaculum rhomboid, minute. An ornamental clinging vine with threadlike branches. Leaves 3.5- 5 cm. long, below the middle 2.3-3.4 cm. wide, petiole 3-4 mm. long, fleshy. Inflorescences well umbelled, with the peduncle 5-6 cm. long. Pedicels up to 1.3 cm. long. Calyx lobes about

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1.5 mm. long. Corolla outspread about 9 mm. in diameter. Corona scales with gynostegium forming a broad cone, 3 mm. long. Northeastern New Guinea: on trees in the forest along the beach at Eitape, about 10 m. altitude (Schlechter #19968 - Blooming in August 1909). So far as one is allowed to make a judgment at the present time, this species with very wide leaves is different from Hoya litoralis Schlechter and from Hoya inconspicua Hemsley. I had the experience to be sure, with the species of the same section from the Philippines, in which the leaves vary greatly in their size and width, but here the dissimilarities are too vividly apparent. It seam to me moreover, that the gynostegium of Hoya litoralis Schlechter is higher and the corona scales are narrower. Moreover, the reddish blooms are brighter. Published by Dr. R. Schlechter in: 1913 Botanische Jahrbücher V.50 p.107, "Die Asclepiadaceen von Deutsch Neu Guinea ". In Hoya Section Acanthostemma (Blume) Kloppenburg (1994) 55-58. R. D. Kloppenburg. Hoya halophila Schlechter in Botanische Jahrbücher 50 (1913) 107. Hoya halophila Schltr. n. sp. - Suffrutex, epiphyticus, ramosus, volubilis. Rami filiformes, flexuosi, bene foliati, teretes, glabri. Folia patentia vel patula, petiolata. late ovata vel suborbiculati-ovata, breviter acuminata, basi obtusissima, utrinque glabra, superne lucida, subtus opaca, carnosa. Cymae longius pedunculatae umbelliformes, folia vulgo distincte superantia, 15-25 florae, pedicellis gracilibus glabris. Calycis segmenta ovata, obtusa, glabra, margine subinconspicue ciliolata, corolla multo breviori. Corolla revoluta usque ad tertiam partem basilarem 5-fida, lobis ovatis, acuminatis, extus glabris, intus apice glabrato excepto papillis reversis subulatis cristalinis obtectus. Coronae foliola extus decurva, oblanceolato-elliptica, antice acuminata, postice obtusa subtus in auriculas 2 oblongas foliolum superantes extensa, antheris subaequilonga. Antherae parvulae subacutae. Pollinia oblique oblongoideo-subfalcata, translatoribus brevibus triangulis, obliquis, retinaculo rhomboideo, minuto. Ein sierlicher Schlinger mit fadenförmigen Zweigen. Blätter 3.5-5 cm lang, unterhalb der Mitte 2.3-3.4 cm breit, Blattstiele 3-4 mm lang. Blütenstände doldenartig auf 5-6 cm langen Stielen. Blütenstiele bis 1.3 cm lang. Kelchzipfel ca. 1.5 mm lang. Korolla ausgebreitet gegen 9 mm Durchmesser. Koronaschuppen mit dem Gynostegium einen Kegel bildend, 3 mm lang. Nordöstl. Neu-Guinea: auf Bäumen in den Wäldern am Strande von Eitape, ca. 10 m ü. M. (Schlechter n. 19968. - Blühend im August 1909). Soweit man sich zur ein Urteil erlauben kann, ist die Art durch die sehr breiten Blätter von H. litoralis Schltr. sowohl wie auch von H. inconspicua Hemsl. verschieden. Ich habe allerdings bei Arten derselben Sektion von den Philippinen die Erfahrung gemacht, dasz die Blätter in ihrer Grösze und Breit stark variieren, doch sind hier die Unterschiede doch zu stark hervorspringenend. Es scheint mir auszerdem dasz das Gynostegium bei H. litoralis Schltr. höher ist und die Koronaschuppen schmäler sind. Auszerdem sind die rötlichen Blüten hier heller. Translation: An epiphytic branched, twining half-shrub. Branches threadlike, flexible, well leaved, round glabrous. Leaves spreading or outspread, petiolate, broadly ovate or somewhat round-ovate, briefly acuminate, base very obtuse, both sides glabrous, lucid above, opaque below, fleshy. Cymes long, peduncled, shaped like an umbel, generally longer than the leaves, 15-25 flowered, pedicels slender, glabrous. Segments of the calyx

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ovate, obtuse, glabrous, margins somewhat inconspicuously ciliate, much smaller than from the corolla. Corolla revolute, 5 parted all the way to one third from the base, lobes ovate, acuminate, outside glabrous, inside papillose, except the concealed wedge shaped terminal scales of the corona decurved, outside oblanceolate-elliptic, inner (anterior) portion acuminate, outer (posterior) portion obtuse, two oblong ears extending beyond the outer apex, anthers almost equal. Anthers small somewhat acute. Pollinia obliquely oblong, almost sickle shaped, with the translators shortly triangular, oblique, retinaculum rhomboid, minute. An ornamental clinging vine with threadlike branches. Leaves 3.5- 5 cm. long, below the middle 2.3-3.4 cm. wide, petiole 3-4 mm. long, fleshy. Inflorescences well umbelled, with the peduncle 5-6 cm. long. Pedicels up to 1.3 cm. long. Calyx lobes about 1.5 mm. long. Corolla outspread about 9 mm. in diameter. Corona scales with gynostegium forming a broad cone, 3 mm. long. Northeastern New Guinea: on trees in the forest along the beach at Eitape, about 10 m. altitude (Schlechter #19968 - Blooming in August 1909). So far as one is allowed to make a judgment at the present time, this species with very wide leaves is different from Hoya litoralis Schlechter and from Hoya inconspicua Hemsley. I had the experience to be sure, with the species of the same section from the Philippines, in which the leaves vary greatly in their size and width, but here the dissimilarities are too vividly apparent. It seams to me moreover, that the gynostegium of Hoya litoralis Schlechter is higher and the corona scales are narrower. Moreover, the reddish blooms are brighter. References: Kloppenburg in Hoyas of Northeast New Guinea (1992) 13-14. A. Nicholas in The Asclepiadaceous Works of Schlechter (1992) 30. Kloppenburg in Schlechter's Hoya Species (1993) 76-77.

Hoya hasseltii Miquel In Rumphia 4 (1849) 29. C. L. Blume. 4. A. Hasseltii: foliis ovatis acutis v. acuminatis basi subcordatis margine subrepandis glabris subtus albido-punctatis; racemis umbelliformibus longiuscule pedunculatis multifloris; corolla ....... — Habit. In province javanica Bantam a divo Van Hasselt observata. Translation: leaves ovate acute or acuminate base somewhat cordate margines somewhat wavy glabris below white-punctate; racemes shaped like umbels on long peduncles many flowered, corolla. Habitat observed by Van Hasselt in the javan province of Bantam. In Museum Botanicum Lugduno-Batavorum 1 (1849) 58. C. L. Blume. 145. Acanthostemma Hasseltii Bl.: foliis ovatis acutis v. acuminatis basi subcordatis margine subrepandis glabris subtus albido-punctatis; racemis umbelliformibus longiuscule pedunculatis multifloris; corolla ....... Bl. in Rumphia l. c. — In provincia javanica Bantam a divio Van Hasselt observata. Translation: Same as above. In Annales Botanicis Systematicae 3 (1852) 64. G. C. Walpers. 4. A. Hasseltii Blume l. c. — Foliis ovatis acutis vel acuminata, basi subcordatis, margine subrepandis glabris, subtus albido-punctatis; racemis umbelliformibus longiuscule pedunculatis multifloris;

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corolla ...... ? — Crescit in insula Java. Translation: Leaves ovate acute or acuminate, base somewhat cordate. margines somewhat wavy glabris, below white-punctate; racemes like umbels on long peduncles many-flowered; corolla ..... ? — Growing in the island of Java. In Flora Indiae Batavia 2 (1857) 524. Zollinger & F. A. W. Miquel. 26 Hoya Hasseltii Miq. Rami tenues glabri, radicellis plerisque papillosis; folia distantia brevi-petiolata, e basi subcordata ovata acuta vel acuminata glabra, subrepanda, subtus albido-punctata 2 ¾ — 2 pol. longa, racemi longiter pedunculati, breves, multiflori, corolla ...... Acanthostemma Hasseltii Blume Rumphia IV, p. 28 Mus. l. c. Java. in Bantam (van Hasselt). Translation: Branches thin glabrous, covered with rootlets very frequently papillose; leaves far apart briefly petiolate, with the base subcordate ovate acute or acuminate glabrous, margine somewhat wavy, below white dotted, 2 ¾ to 2 inches long, racemes long pedunculate, small, many flowered, corolla ...... Acanthostemma Hasseltii Blume in Rumphia IV page 28 Mus. as above Java, in Bantam (van Hasselt). In Exkersionsflora von Java 3 (1912) 1103. S. H. Koorders. (key). 13b. Blätter entternt stehend, eiförmig, kahl, am Grunde herzförmig, oben spitz oder zugespitzt, unterseits weisz punktiert, (5—7 cm lang, mit (ausgeschweistem Rande. Pedunculi vielblütig. Korolle und Frucht unbekannt. Miq. l. c. 524; Acanthostemma Hasseltii Miq. Java: In der Res. Banten ohne nähere Standortsangabe (van Hasselt); bisher have ich diese Art noch nicht im Buitenzorger Herbar wiedergefunden......... H. Hasseltii Miq.* Translation of German: 13 b. The leaves are standing outspread, egg-shaped, bare, at base heart shaped, apex acute or pointed, lower side spotted white, +/- 5-7 cm. long, with more or less wavy edges. Peduncles multi flowered. Corolla and fruit unknown. Miquel at the place cited page 524; Acanthostemma hasseltii Miquel. Java: In the residency of Banten* without any detailed description of location (Van Hasselt); so far I was unable to find this species in the herbarium at Buitenzorg. * Note: His spelling differs from above Bantam. In Hoya Section Acanthostemma (Blume) Kloppenburg (1994) 59. R. D. Kloppenburg. Hoya hasseltii Miquel in Flora Indiae Bataviae 2 (1857) 524. Rami tenues glabri, radicellis plerisque papillosis; folia distantia brevi-petiolata, e basi subcordata ovata acuta vel acuminata glabra, subrepanda, subtus albido-punctata, 2 3/4-2 poll. longa, racemi longiter pedunculati, breves, multiflori, corolla .... Acanthostemma Hasseltii Blume Rumphia IV:28 Mus. l.c. Java, in Bantam (Van Hasselt). Translation: of the Latin: Branches thin glabrous, covered with papilla and rootlets; leaves far apart with short petioles, from the base somewhat cordate ovate acute or acuminate, glabrous, somewhat repand (having a slightly uneven and waved margin) (undulant) with white punctations on the lower surface (white dots) 2.75-2 inches long (he probably meant 2.75 to 3 inches since Koorders's key says 5-7 cm.), racemes on long peduncles, small many flowers, corolla ....

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The older reference to this species is found in Rumphia 4 (1848) 28 as follows. 4. A. Hasseltii: foliis ovatis acutis v. acuminatis basi subcordatis margine subrepandis glabris subtus albido-punctatis; racemis umbelliformibus longiuscule pedunculatis multifloris; corolla .........- Habit. In province javanica Bantam a divo Hasselt observata. Translation: Leaves ovate acute or acuminate with the base somewhat cordate, margins undulant glabrous with white dots below; racemes shaped like umbels with long pedicels many flowered: corolla...........- habitat in the province of Bantam, Java observed by the late Hasselt. The description found in Museum Botanicum Lugd.- Bat. 1 (1849) 58 is the same as the above. Koorders in Exkursionsflora von Java, 2 (1912) 1103 offers more detail. 13 b. Blätter entternt stehend, eiförmig, kahl, am Grunde herzförmig, oben spitz oder zugespitzt, unterseits weisz punktiert, ± 5-7 cm. lang. mit ± ausgeschweiftem Rande. Pedunculi vielblütig. Korolle und Frucht unbekannt. Miq. l.c. 524; Acanthostemma Hasseltii Miq. Java: In der Res. Banten ohne nähere Standortsangabe (van Hasselt); bisher habe ich diese Art noch nicht im Buitenzorger Herbar wiedergefunden. German Translation by Dieter Paul: 13 b. The leaves are standing outspread, egg-shaped, bare, at base heart shaped, apex acute or pointed, lower side spotted white, ± 5-7 cm. long, with more or less wavy edges. Peduncles multi flowered. Corolla and fruit unknown. Miquel at the place cited page 524; Acanthostemma hasseltii Miquel. Java: In the residency of Banten without any detailed description of location (Van Hasselt); so far I was unable to find this species in the herbarium at Buitenzorg. Note: see comb. nov. under H. kuhlii (Bl.) Kds..

Hoya hellwigiana Warburg In Repertorium Specierum Novarum 3 (1907) 341. O. Warburg & R. Schlechter. 27. Hoya hellwigiana Warb., nov. spec. Ramulis glabris 3 mm crassis flavidis lenticellis elevatis rotundatis parce inspersis, petiolis 10—12 mm longis 3 mm crassis flavidis, foliis cuneato-lanceolatis vel oblongis 9—15 cm longis, apice et basi acutis crasse coriaceis, venis utrinque ca. 2 ascendentibus fere omnino inconspicuis. Pedunculo exstante 5.5 cm. longis glabris, sepalis 1—1.5 mm longis ovatis obtusis glabris, corolla ca. 4 mm in diametro utrinque glabra, lobis late triangularibus, coronae staminae lobis patentibus fere planis late ovatis externo obtusis, apice interno sensim in antheras transeuntibus, antherarium appendiculo oblongo obtuso. Kaiser Wilhelms-Land: Bussum bei Finschhafen (Warburg no. 21313). Eine durch ausgebreiten, förmlich das untere Ende der Antheren bildenden Coronalappen ausgezeichnete Art, die ich zu Ehern meins leider kurze Zeit später verstorbenen Reisegefährten auf jener Tour, des Botanikers Dr. Hellwig, benannt habe. Translation: Branches glabrous 3 mm thick yellow with round elevated lenticels moderately interspersed, petioles 10 to 12 mm long 3 mm thick yellow, leaves cuneatelanceolate or oblong 9 to 15 cm long, apex and base acute thickly leathery, veins on both sides about 2 ascending almost entirely inconspicuous. Peduncles extended 5,5 cm long 1 mm thick apex flower bearing end brief barely thickened glabrous; pedicles 0.25 mm thick 1 mm long glabrous, sepals 1 to1.5 mm long ovate obtuse glabrous, corolla about 4

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mm in diameter both sides glabrous, lobes broad triangular, lobes of the staminal corona almost flat ovate external apex obtuse, internal apex slightly beyond the anthers, the anthers appendages oblong obtuse. Kaiser Wilhelms-land: at Bussum near Finschharbor (Warburg number 21313). One through the wide spread, formally the low end of the anthers of forming Corona scales excellent species, that I have named mine unfortunately a short time later the botanist's Dr. Hellwig in a very brazen vehicle trip was killed from that tour. In Botanische Jahrbücher 50 (1913) 116. R. Schlechter. 16. H. hellwigiana Warburg in Fedde, Repertor. III (1907) p.342. H. hellwigii Warb. ex K. Schum. u. Lauterb. Fl. Deutsch Schutzgeb. Südsee (1901) p.512 (nomen). Nordöstl. Neu-Guinea: auf Bäumen in den Wälden des Torricelli-Geberges, ca. 500 m ü. M. (Schlechter n. 20149. Blühend im September 1909); auf Bäumen in den Urwäldern von Wobbe, im Minjemtale ca. 300 m ü. M. (Schlechter n. 16265. Blühend im Juli 1907); auf Bäumen in den Wäldern des Kani-Geberges, ca. 600 m ü. M. (Schlechter n. 17631.- Blühend im April 1908); auf Bäumen an den Ufern des Wabe, ca. 100 m ü, M. (Schlechter n. 18073. Blühend im July 1908) auf Bäumen in den Wäldern des Finisterre-Gebirges, ca. 500-700 m ü. M. (Schlechter n. 17939, 17995. -Blühend im Juli 1908); am Ramuflusz (K. Tappenbeck n. 37.- Blühend im Mai 1898); auf Bäumen in den Wäldern auf Flusze des Bismarck-Gebirges, ca. 150-300 m ü. M. (Schlechter) n. 18447, 18640. - Blühend im October -November 1908); bei Bussum, unweit Finschafen (O. Warburg n. 21343); auf Bäumen am Waria bei Pema, ca. 100 m ü. M. (Schlechter n. 17457. -Blühend im März 1908); auf Bäumen inden Wäldern des Gomadjidji, am Waria, ca. 450 m ü. M. Schlechter n. 19388. (Blühend im Mai 1909). Ich halte die sämtlichen hier aufgefürten Exemplare für Formen einer etwas variablen Art. Möglich ist allerdings, dasz weitere Studien an lebendem Material doch och Veranlassung geben werden zu einer weiteren Aufteilung. Zur Zeit ist selbst durch as mir vorleigende gute Material nicht ausreichend festzustellen, ob das, was ich heir für Formen halte, beständige Arten sind oder nicht, So sehr ich mich denn gezwungen, hier die Art ungleich weiter zu fassen als bei den übrigen. Besonders grosz sind die Neränderungen an der Blättern. Stutzig macht mich aber der Umstand, dasz einige Exemplare in der Behaaruing der Innenseite der Korolla von dem Typus abweichen. Diese werden doch wohl noch weiterer Begutachtung bedürfen, Die Blüten sind gewöhnlich weisz, selten blasz-rosa überlaufen mit weiszer Korona. Translation: see next below. In Hoyas of Northeastern New Guinea (1992) 41. R. D. Kloppenburg (Translation of Die Asclepiadaceen von Deutsch Neu-Guinea). 16. H. hellwigiana Warburg - in Fedde, Repertor. III (1907) p. 342. H. hellwigii Warburg in K. Schumann and Lauterbach Flora of the German South Seas protectorate (1901) p. 512 (name). Northeastern New Guinea: on trees in the forest of the Torricelli Mountains about 500 m. altitude (Schlechter #20149 - blooming in September 1909); on trees in the primary forest of Wobbe in the Minjem Valley about 300 m. altitude (Schlechter #16265 - blooming in July 1907); on trees in the forest of the Kani Mountains about 600 m. altitude (Schlechter #17631 - blooming in July 1908); on trees on the banks of the Wabe about 100 m. altitude (Schlechter #18073 - blooming in April 1908); on trees in the

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forest of the Finisterre Mountains about 500-700 m. altitude (Schlechter #17939, 17995 - blooming in July 1908); on the Rama river (K. Tappenbeck #37 - blooming in may 1898); on trees in the forest at the foot of the Bismarck Mountains about 150-300 m. altitude (Schlechter #18447, #18640 - blooming in October - November 1908) at Bussum, not far from Finschhafen (O. Warburg #21313); on trees at Wari, near Pema about 100 m. altitude (Schlechter #17457 - blooming in March 1908); on trees in the forest of Gomaljidji at Wari about 450 m. altitude (Schlechter #19388 - blooming in May 1909). I regard the specimens here as forms of one highly variable species. It is certainly possible, with more in depth study of living material there may turn out to be nevertheless, a further division. At the present time even with good material one cannot ascertain for certain weather the ones I have, have a standing as species or not. Thus, this being so, I am forced to consider as being superfluous, the dissimilar species presented here, especially the variation of the leaves are large. It is startling to me under the circumstances, as an example, that the pubescence on the inside of the corolla differ from the type. These in turn require, nevertheless, further opinions. The blooms are generally white, rarely light coppery rose, overtopped with a white corona. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. hellwigiana Warb. – 15, 33, 36.

Hoya hellwigii Warburg In Nachtrage zur Flora der Deutschen Schutzgebeite in der Südsee (1901) 512. K. Schumann & K. Lauterbach. H. Hellwigii Warb. Ms, in Mons. Ined. Kaiser Wilhelmsland: Finschhafen, bei Bussum (Warburg); Ramufluss (Tappenbeck n. 37, blühend am 30. Mai 1898). Ist endemisch. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. hellwigii Warb. ex Schumann – 15.

Hoya heuschkeliana Kloppenburg In the Hoyan 11 #1 pt. 2 (1989) i. R. D. Kloppenburg. Hoya heuschkeliana Kloppenburg nov. spec. Epiphyticis vel terrestris, ramosa, puberulis, ramis filiformibus, flexuosis. Folia enervia, petiolatis, obovata-ellipticis, obtusis, basi cuneata, carnosis, dorso puberulis utrinque-punctata, (or pusticulata vel granulata) 2-5 cm longis, medio 0.5-2 cm latis sapra convexis. Petiolo curvis pubescentes circ. 0.2 cm longis, carnosa. Pedunculo robusto circ. 0.2 cm longis. Pedicellis 0.4 cm longis. Cymis umbelliformibus 2-7 floris. Calyx parvus fere ad basin in lobos 5 anguste triangulares partitus, lobis circ. 0.15 cm , punctata, auriclis 1 ab inter se basi lobis. Corolla urceolato-campanulata recurva apace revoluta, extus glabra et punctata, lobis ovatus acutis intus apice excepto velutinus-puberulis, recurva apace glabra, usque ad 1/3 fere 5 lobata, 0.23 cm longis, cum complaratis circ 0.8 cm diam. Coronae lobis compressis apiculata, apice externo bifidis, obtuso, apice intus acutissimus, inferiore concavo superiore, basi ad apicem 0.2 cm longa, angulo dorso 45 degrees. Anthera appendiculatus non excedente. Pollinius

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oblique-oblongodius, translatoribum brevissimis, retinaculo minuto anguste rhomboideo. Stigmatis columno capite 5 bifid basis. Distribution and habitat: Luzon (northern portion of Lake Bulusan, Sorsogon, Philippines), epiphytic on trees at ca. 50 m alt. Endemic. Now cultivated in Prof. J.V. Pancho's garden at Libis, Los Banos, Laguna, Philippines. Specimens Examined: Luzon (Sorsogon), Pancho 2175 (holotypus CAHP; isotypus UC), Kloppenburg #89008 (CAHP). This species probably belongs in a new section (?) and is closely related to the unpublished Hoya intermedia of Schlechter's, collected by W. I. Hutchinson in 1906 in the province of Zamboanga, Mindanao, Philippines #5177 from which it differs in having acute calyx lobes and shorter outer bilobed appendages on the corona scales. Being urceolate it is different than other hoyas published to date. The sides of the outer corona lobe are extended as in Hoya bilobata, Schltr. and other Acanthostemma species, but are not incurved at their tips as in H. bilobata, Schltr. The pollinia is typically Hoya with an outer keel as opposed to Dischidia's keelless pollinia. This species is named for Mr. Dexter Heuschkel, Vice President, Manila Memorial Park, Paranaque, Metro Manila, Philippines, who maintains a collection of Philippine endemic ferns and hoyas among others. Through his kindness I was able to study some of his hoya collections otherwise now difficult to find and collect. Translation: Epiphytic or terrestrial, branched, puberulous, stems threadlike, flexible. Leaves without nerves, petiolate, ovate-elliptic obtuse, base wedge-shaped, fleshy, on the back puberulous both sides punctate (pustulous or granulate) 2 to 5 mm long, in the middle 0.5 to 2 cm. wide, above convex. Petioles curved about 0.2 cm. long. Pedicels 0.4 cm. long. Cymes shaped like an umbel 2 to 7 flowered. Calyx small divided to nearly the base with 5 narrow triangular lobes, lobes about 0.15 cm. punctate, at the base of the lobes on the inside is one ear. Corolla urceolate-campanulate recurved, apex revolute, outside glabrous and punctate, lobes ovate acute, inside except for the apical area velvety-puberlous, apex recurved glabrous, at least nearly 1/3 5 lobed, 0.23 cm. long, being relatively about 0.8 cm. in diameter. Lobes of the corona compressed apiculate, external apex bilobed obtuse, apex inside very acute, below concave, above at the base apex 0.02 cm. long, above angle 45 º. Anther a small appendage nit exceeding. Pollinia oblique-oblong translators very small, retinaculum small, narrow rhomboid. Stigma column head base of 5 divisions. In Philippine Hoya Species 3rd Ed. (1996) 80-81. R. D. Kloppenburg. Hoya heuschkeliana Kloppenburg. Type: #2175 (CAHUP) collected by Professor Juan V. Pancho at the northern portion of Lake Bulusan, Sorsogon Prov., Luzon, Philippines. Epiphytic on trees at about 50 m. altitude, endemic. Section Skenostemma Kloppenburg. An epiphytic or terrestrial small vine, twining and creeping. Its stem is puberulous, filiform with adventitious roots at the nodes. Foliage is enervis, petiolate, ovate-elliptic, with a obtuse base cuneate, carnose with numerous surface granules or punctations, 2.5 cm. long x 0.5-2 cm. wide at the middle, undersides lighter green and concave. The leaf color is dark green. Petioles are curved, pubescent about 0.4 cm. long. Umbels of 2-7 pale cream to rose tinted flowers, urceolate, but flattened 0.8 cm. in diameter. Peduncles stiff, terminal and axillary 0.5 cm. long. Pedicels also rigid 0.4 cm. long, expanding as they approach the calyx. Calyx lobes nearly free at the base, narrowly

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triangular, punctate 0.15 cm. long with one ligule at each lobe intersection. Corolla urceolate cut about 1/3 with the tips reflexed and glabrous inside, outside surface glabrous and punctate, inside except for tip velvety puberulous ovate with acute apex 0.23 cm long. Corona lobes thin, compressed, tapering outward from the centers, inner apex extended upward and narrowly elongated with a sharply acute apex, outer lobe broad, rounded with apex bilobed extensions which are typical of the Acanthostemma section. Extensions are broad and flat tapering inward at a 45 degree angle, ends are rounded, 0.2 cm. beyond outer scale apex. Anther appendages not exceeding apex of inner lobe. Stigma base 5 rotate and bifid at each rotate edges, columnar and capitate with central extensions. Ovaries are urn shaped 0.18 cm. tall. Pollinia 0.05 cm long, relatively short and broad, outer apex tapering inward at a 45 degree angle, keeled. no vacuole present. Keel not extending to pollinia attachment point, narrow at translator. Retinaculum with broad apex, split at lower outer apex. Meaning: heuschkeliana — Named to honor Dexter Heuschkel, director Manila Memorial gardens, who has been responsible for introducing a vast number of Hoya species into commerce. In Asklepios 82 (2001) with permission of the publication and the author Ulrich Meve. HOYA HEUSCHKELIANA, A NEO-ENDEMIC OF MT. BULUSON (PHILIPPINES, LUZON), AND REMARKS ON THE URCEOLATE FLOWER TYPE IN ASCLEPIADOIDEAE Introduction The southernmost tip of Luzon, the main island of the Philippines, is occupied by the Sorsogon Province, which is topographically dominated by the Mt. Bulusan Volcano. From the forested areas around Lake Bulusan at the foot of Mt. Bulusan, and only from this very restricted area, an unusual small-flowered Hoya was originally brought to our attention by Dale Kloppenburg, who described it as Hoya heuschkeliana (Kloppenburg 1989). It was named after Dexter Heuschkel, vice president of the Manila Memorial Park. Recently, J. Schneidt recollected Hoya heuschkeliana while collecting Asciepiadaceae on the Philippines and in China in 1996. Living material was brought into cultivation in Germany. Cuttings made from Schneidt’s two different collections have been distributed by the author to German and British Hoya collectors. Hoya heuschkeliana Kloppenburg This interesting and unusual Philippine endemic possesses globose-urceolate flowers, which only open slightly in the center by recurving the tips of the corolla lobes (Figs. 1, 2). It is the only Hoya species that does so, and for this reason has been placed in a section of its own by Kloppenburg (1993), Hoya sect. Skenostemma. Such flowers could be easily mistaken for those belonging to the genus Dischidia (Fig. 3. pitcher plants; cf. Walker 1985) where urceolate tubular flowers are the rule, while typical Hoya possesses radiate (sometimes with revolute corolla lobes) to broadly campanulate corollas (cf. Forster et al. 1998). Since the leaves are rather small, rounded and succulent, the vegetative appearance of H. heuschkeliana is also more similar to Dischidia rather than to most species of Hoya. However, the flowers of’ H. heuschkeliana must be regarded as “falsely urceolate” (or pseudo-urceolate), since the corolla tube, the fused part of the corolla, is shortly campanulate rather then truly urceolate and the corolla lobes are longer than

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the tube. Its globose-urceoate appearance is due to the erectocnivent corolla lobes, which do not separate except for the tips but with their margins adhering for most of their length. From the evolutionary point of view this is a derived character in Hoya, although, seemingly it makes a very short step. In contrast, Dischidia and also the monotypic genera related to Hoya such as Madangia (cf. Forster et al. 1997) and Micholitzia (Goyder & Kent 1994) have more completely fused corollas with only small free lobes (Fig. 3). Considering the corona and pollinaria also, there cannot be left any doubt as to the generic affiliation of Hoya heuschkeliana. The corona lobes are lanceolate and star-like arranged around the gynostegium—never found in Dischidia. The corona lobes are grooved on the lower side, which typically characterizes Hoya (but which is also present in Micholitzia and Absolmsia (the third monotypic genera in the complex, e.g., Omlor (1 997)). Finally, each pollinium bears a germination crest along the outer margin as in so many hoyas (also Absolmsia, Madangia and Micholitzia) but unlike Dischidia. Nevertheless, there is a structural peculiarity of the corona otherwise rare in the genus. These are the wings formed by the basal margins of the corona lobes, and which extend dorsally into tooth-like projections (see illustrations in Kloppenburg 1989). This character is only known from a group of species with characteristically revolute corolla lobes such as Hoya bilobata Schltr. and H. tsangii C. Burton, both endemics of Mindanao, or the Indonesian H. picta Miqu. And H. revoluta Wight.

Hoya heuschkeliana Kloppenburg, Hoyan 11(1.2): 1-3 (1989). Type: Philippines, Luzon, Sorsogon, Lake Bulusan, alt. 50 m, Pancho 2175 [CAHP, holo; UC, iso].

Description: Plants epiphytic, climbing on tree trunks, with white latex: stems procumbentcreeping (stem tips twining by chance), many-branched, being attached to the undergrowth by wiry adventitious roots appearing along the nodes and internodes at the lower side of stems. Stems green, cylindrical, wiry, 1—2 mm in diam., with scattered, translucent, decumbent hairs; leaves stalked for 3—5 mm, lamina broadly ovate to elliptic, 2—5 cm long, 1- 2.2. cm broad, succulent to leathery, upper-side dark green (purplish when stressed by sun or drought), underside brighter, usually slightly convex, basally obtuse to cuneate, scattered hairs on the margins; inflorescences globoid, 2—7-flowered, inserting extra-axillary on lower face of stem; peduncle 2—3 mm long, scatteredly hairy with appressed, translucent hairs: pedicels 2—4 mm long: sepals deltate, ± 1.2 mm long. Flowers nectariferous, emitting a dull-sweetish scent mainly in the late afternoon; corolla flattened globose-urceolate, ca. 3.5—45 mm long, 5—7 mm broad, outside glabrous, inside finely papillose, corolla tube cream-coloured, corolla lobes deltoid to deltoidcucullate, ca. 2.5 mm long, 2.5 mm broad at base, acute, erect-connivent, tips completely recurved, pinkish, margins slightly recurved, pinkish. Corona uni-seriate, staminal corona lobes lanceolate, ca. 3 mm long, 1.2 mm broad, ascending in a 45° spiral. Yellowish, lower side canaliculate (grooved), dorsal part of corona lobe apically aproned, basally winged and bifid (wings extending into a dorsal tooth-like projection), inner parts of corona lobes acute, erect, tips connivent above the stylar head. Pollinaria: pollinia rounded rectangular, ca. 0.25 mm long, 0.14 mm broad, corpuscle rhomboid, ca. 0.25 mm long, 0.1 mm broad, caudicles short, cylindrical. Carpels glabrous; fruits and seeds unknown. Chromosome number: 2n=22 (Voucher: Schneidt 96-95 [MO, UBT]).

Habitat: Only known from lowland to submontane forest surrounding Lake Bulusan (50—600 m). Material investigated: PHILIPPINES: Luzon, Sorsogon, Prov., Mt. Bulusan Volcano, Lake Bulusan, “Lover’s Lane” footpath around lake, alt. ca. 600 m, 4th of June 1996, J. Schneidt 96-95 (MO, UBT) et 96-96 (MS UN, UBT).*

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Cultivation Hoya heuschkeliana is a lovely and easily pleased plant in cultivation. It roots easily, branches generously and flowers nearly the whole year round if the conditions are suitable (avoid too dry room air). Originally an epiphytic species it can be grown in a pot with ordinary soil, in a hanging basket, or (initially) fixed onto trunks (fern trunks). It is only the somewhat hidden and small inflorescences, which are usually formed on the lower side of stems and often partly hidden between the dense foliage, which might diminish growers delight. But since the emission of a distinct sweetish-dull scent (similar to some plastics) is considerable, predominantly in the late afternoon, the flowers cannot be overlooked in the greenhouse.

On the distribution of urceolate flowers in Asclepiadoideae Urceolate-shaped flowers represent a wide-spread floral variation in Asclepiadoideae, which shows up in a limited number of genera, sometimes even as singular exceptions as in Hoya. Within the species-rich Old World genus Tvlophora (ca. 100 species from West Africa to New Caledonia). T. urceolata Meve is the single globose-urceolate-flowered species in a genus of otherwise radiate flowers (Meve 1996). Even in size, the flowers of H. heuschkeliana and T. urceolata are of similar dimensions. In the stapeliad genus Huernia (over 60 species in Africa and Arabia), with its large and fleshy flowers, we find another of these exceptions in H. urceolata L.C. Leach (cf. Leach 1988). In general, urceolate-shaped flowers are quite regularly distributed in the stapeliads, in particular in Eehidnopsis (Fig. 4). Pseudolithos, Stapelianthusi, Stapeliopsis and Pectinaria. In these genera this flower structure is more or less dominant or it even serves as genetic marker (e.g., Stapeliopsis). Additional examples from the Old World are to be found in ceropegiads and marsdeniads such as Brachystelma (e.g., B. oianthum Schltr.), Heterostemma (H. sect. Oianthus), Sisyranthus, Gunnessia, Gyrnnerna and Marsdenia (e.g.. M. australis R.Br. Druce, M. microstoma Schltr., M. tinctoria R.Br.). Quite exceptional is the corona-less Asclcpiadeae genus Microloma (e.g., M. tenuifoliurn (L.) K. Schum.) in southern Africa (cf. Bruyns 1991). though, the flowers of most species of this genus are rather elongated to tubular. With Stenostelma (S. capense Schltr., Fig. 5) and some small-flowered species of the tuberous relatives of Asciepias found in southern Africa, there are also some representatives of the tribe Asclepiadeae that have developed urceolateshaped flowers (or at least globose pseudourceolate flowers as in Stenostelma capense, where the general construction of the corolla is comparable to that of Hoya heuschkeliana). In the New World, which houses more Asclepiadoideae species hut much fewer genera than the Old World, the urceolate flower type is quite frequently spread over some groups, typically represented by small-flowered Asclepiadeae — Metastelmatinae such as Metastelma (e.g., Metastelma sepicola Pittier), Cynanchum (C. nubicola Morillo), Ditassa (D. franciscoi (Morillo) Liede med.), Tassadia (T. guianensis Decne., T. obovata Decne.) and Orthosia (0. urceolata Fourn.). It occurs also in the Hemipogon "Asiephanus” and Gonioanthela (e.g., G. bradeana Fontella Pereira & A. Schwarz) lineages. In Asclepiadeae----Oxypetalinae flowers with deep (cylindrical) corolla tubes are wide-spread, urceolate-shaped corollas, however, are quite rare (e.g., Amblystigma cionophoruin E. Foum., Morrenia herzogii Schltr.). Finally, urceolate flower structures are wide-spread in New World Marsdenia (e.g., M. altissima (Jacq.) Dugand, M. amazonica Morillo or M. sprucei Rothe).

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Typically, the advantage of urceolate flower structures lies in the restriction of access for most of the flower-visiting organisms to the interior of the flower (and its nectar). This kind of specialization can finally lead to highly specific flower-pollinator relationships. Such systems are known for Microloma (pollinated by sunbirds of the genus Necctarinia, Pauw 1998) or Ditassa franciscoi (pollinated by wasps as photographed by Zarucchi: http://mobot.mobot.org/cgibin/search_pick?w3till=02610433_001). In the stapeliads, the urceolate flowers obviously do not differ in being fly-pollinated as the open flowers (cf. Meve & Liede 1994). Only for these three examples the pollinators (pollinating groups) are known but they might demonstrate already that the urceolate flower type is involved in several fairly different pollination systems. Acknowledgements

My thanks go the Dr. Josefine Schneidt, Edinburgh, who abandoned the whole living material collected on the Philippines to me.

* MO = Mossouri Bot. Gdn. UTB =Univ. Bereuth, Gremany MS = Univ. of Messia, Italy. References BRUYNS, P.V. (1991): Notes on Microlonia. Asklepios 52: 71—74. FORSTER, P.I., LIDDLE, D.J. & LIDDLE, E.M. (1997): Madangia inflata (Asclepiadaceae: Marsdenieae), a new genus and species from Papua New Guinea. -1 Astrobaileya 5(l): 53 57. FORSTER, P.!., LIDDLL, D.J. & LIDDLE, E.M. (1998): Diversity in the genus I-Iota (Aselepiadaceac Marsdenieae). Aloe 35(2): 44-48. GOYDER, D.J. & KENT, D.H. (1994): Micholitzia obcordata N.E. Br. (Asclepiadaceae-— Marsdenieae) reinstated. Asklepios 62: 1 3—l 9. KLOPPENBURG, D. (1989): Hoya heuschkeliana. Hoyan 11(1.2): 1—3. KLOPPEBURG, D. (1993): Hoya sections, a complete study with, modifications and additions. D. Kloppenburg: Fresno, California, USA. LEACH, L.C. (1988): A revision of Huernia R. Br. (Asclepiadaceae). Excelsa Tax. Sci: 4: l--197. MEVE, U. (1996): A new species of Tvlophora (Asclepiadaceae) from West Tropical Africa. Kew Bull. 51(3): 585—589. MEVE, U. & LIEDE, S. (1994): Floral biology and pollination in stapeliads—new results and a literature review. Pl. Syst. Evol. 192: 99—I 16. OMLOR, R. (1998): Generische Revision der Marsdenieae (Asclepiadaceae). Shaker Verlag: Aachen, Germany. PAUW A. (1998): Pollen transfer on birds tongues. Nature 394: 73 l—732. WALKER, C.C. (1985): Dischidia—--—An introduction to asclepiad alit plants. Asklepios 35: 3

Hoya hirsuta Moon In Moon Cataloge 21. Cited under Tylophora asthmatica Wt. & Arnott. (Handbook of the Flora of Ceylon).

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Hoya hollrungii Warburg In Nachtrage zur Flora der Deutschen Schutzgebeite in der Südsee (1901) 512. K. Schumann & K. Lauterbach. H. Hollrungii Warb. Ms. In Mons. Ined. ( H. purpurea K. Sch. Fl. Kais. Wilhelmsland. 110, non Fl. Deutsch-ostas. Schutzgeb. 216, non Bl. Kaiser Wilhelmsland: Augustafluss, erste Station (Hollrung n. 661, blühend im August 1889). Ist endemisch. In Nachtrage zur Flora der Deutschen Schutzgebeite in der Südsee (1905). K. Schumann & K. Lauterbach. H. Hollrungii Warbg. ex Fl. d. S. (l901), p. 512. Kaiser-Wilhelmsland: Zwischen Bäumen und Sträuchern windend im Walde bei Konstautinhafen, alt. ca. 20-50 m (R Schlechter n. 14297, bl. März 1902). Bei den von mir gesammelten Exemplaren sind die Blüten etwas gröszer als bei dem von Hollrung entdeckten, sonst stimmen beide recht gut überein. Sehr charakteristisch ist die hohe dicke Säule, auf der die Korona aufsitzt; dieselbe ist sehr fein behaart. Die Blüten sind gelbgrün gafärbt, nach der Mitte zu rötlich angehaucht. Die ganze Pflanze erinnert etwas an H. coronaria Bl. und ist wohl a1s ihre nächste Verwandte zu betrachten. Translation: Kaiser-Wilhelmsland: Between trees and shrubs winding in the forest with Konstautinhafen, altitude approximately 20-50 m (R. Schlechter n. 14297, Bl. March 1902). With the copies collected by me, the blooms of some larger are as with that of Hollrung discovered, otherwise they agree both quite well. The high fat column, on which the corona mounts, is very characteristic; the same is very finely hairy. The blooms are colored yellow-green, after the middle too reddish hue. The whole plant reminds something of H. Coronaria Bl. and is about where to look at its next relative. In Repertorium Specierum Novarum 3 (1907) 342. O. Warburg & R. Schlechter. 28. Hoya Hollrungii Warb., nov. spec. Ramulis in sicco luteis 3 mm crassis ad nodos haud incrassatis; petiolis 1— 2 cm longis, 1—1.5 mm crassis glabris, foliis subcoriaceis glabris ovatis 5—7 cm longis 3—3.5 cm latis, apice obtuse breviter acuminatis saepe obliquis basi rotundis vel fere subpeltatis, in costae basi supra haud glandulosis, venis utrinque 5 patulis longe ante marginem arcuate conjunctis vix conspicuis. Pedunculis juxtapetiolaribus 1—3 cm longis 1.5—2 mm crassis, junioribus griseo-puberulis, parte apicali florigera seape usque 1.5 cm longa cylindrica cicatricibus florum obtecta, pedicellis usque 2 cm longis 0.5 mm latis puberulis, sepalis ovatis apice rotundatis 2- 3 mm longis extus puberulis, corolla fere 2 cm in diametro in sicco nigra extus griseo-puberula intus glabra, coronae stamineae lobis late naviculiformibus supra concavis subtus subcarinatis 1.5 mm latis 2 mm longis corollae haud incumbentibus, apice exteriore haud recurvato obtuso, apice interiore elevato demum, retuso, gynostegii stipite brevissimo dense piloso, antherarum appendiculis brevibus rotundatis. Fructu longe (2—3 cm) pedicellato basi sepalis subincresentibus recurvis circumdato 12 cm longo. 2.5 cm lato, semnibus planis 6 mm longis 3 mm latis longe comosis. Kaiser Wilhelms-land: 1. Augusta-Station (Hollrung no. 661). Von H. purpurea, als welche sie in der Flora von Kaiser Wilhelms-Land figuriert, vor allem durch die aussen behaarten Corollen und kelchzipfel, sowie durch die zugespitzten

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kleineren Blätter, übrigens auch durch die anders gebaute Corona verschieden. Translation: Branches when in the dry state yellowish 3 mm thick not thickened at the nodes, petioles 1-2 cm long, 1-1.5 mm thick glabrous, leaves somewhat leathery glabrous ovate 5-7 cm long 3-3.5 cm wide, apex obtuse briefly acuminate often oblique with the base rounded or almost peltate, at the base above the midrib not at all glandular 5 conspicuous veins on both sides long spreading curving and joining near the margines. Peduncles approaching the petioles 1-3 cm long 1.5-2 mm thick, when young graypuberulous, the apical portion bearing flowers often as long as 1.5 cm cylindrical covered with scars of fallen flowers, pedicels often 2 cm long 0.5 mm wide puberulous , sepals ovate with apexes rounded 2-3 mm long outside puberulous, corolla nearly 2 cm in diameter when dried black outside gray-puberulous inside glabrous, with the lobes of the corona broadly boat shaped concave above somewhat channeled below 1.5 mm wide 2 mm long not at all incumbent upon the corolla, exterior apex not at all recurved obtuse, interior apex elevated at length retuse, gynostegium stypes briefly densely hairy, with the anther appendages short rounded. Fruit on a long peduncle (2-3 cm) base of sepals somewhat increased recurved long, 2.5 cm wide, seeds flat 6 mm long 3 mm wide, with long comas. Kaiser Wilhelmsland (German New Guinea) at the Augusta Station (Hollrung #661). Near Hoya purpurea, which is figured in Flora of Kaiser Wilhelmsland, it differs from others through the shape of the corolla and sepals also the smaller leaves. Moreover through the distinct corona. In Botanische Jahrbücher 50 (1913) 136 R. Schlechter. 47. H. Hollrungii Warb. In Fedde, Repertor. III (1907) p. 342. H. purpurea K. Schum. in K. Schum. u. Hollrung, Fl. Kaiser-Wilhelmsland (8189) p. 110 (nec. Bl.). Nordöstl.. Neu-Guinea: bei der ersten Kaiserin-Augustaflusz-Station (M. Hollrung n. 661. (Blühend im August 1889); zwischen Bäumen und Sträuchern windend im Walde bei Konstantinhafen, ca. 100 m ü. M. (Schlechter n.l4297. —Blühend im März 1902); an Bäumen an Abhängen des Minjem-Tores, ca. 100 m ü. M. (Schlechter n. 16086. — Blühend im Mai 1907); Liane an Waldrändern bei der Kaulo-Etappe, ca. 250 m ü. M. (Schlechter n. 17530.— Blühend im April 1908.) Die Unterschiede zwischen dieser Art und H. purpurea Bl. sind von Warburg richtig auseinandergesetzt worden, ich habe hier nur hinzuzufügen, dasz auch die Blütenfärbung recht verschieden ist. H. Guppyi Hemsl. und H. affinis Hemsl. sind, nahe mit dieser Art verwandt. Die Blüten sind grünlich gelb, inner nach dem Grunde rötlich die Kronen oben rot. Translation: See below under Hoyas Of Northeastern New Guinea Kloppenburg.

In Hoyas of Northeastern New Guinea (1992) 96. R. D. Kloppenburg. (Translation from Die Asclepiadaceen von Deutsch Neu Guinea by R. Schlechter. 47. Hoya hollrungii Warburg in Fedde Repertor. III (1907) p. 342 - H. purpurea K. Schumann in K. Schumann and Hollrung, Fl. Kaiser Wilhelmsland (1898) p. 110 (not of Blume). Northeastern New Guinea: at the first Empress Augusta River Station (M. Hollrung #661 - Blooming in August 1889); climbing among the trees and shrubs in the forests at

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Constantine Harbor about 20-50 meters altitude (Schlechter #14297 - Blooming in May 1902); on trees and slopes of the Minjem Gate, about 100 meters altitude (Schlechter #16086 - Blooming in May 1907); vines at the forest edge of the Kaulo-Etappe about 250 meters altitude (Schlechter #17530 - Blooming in April 1908). The differences between this species and H. purpurea Blume were correctly set forth (explained) by Warburg, I have only here to add, that the bloom color is variable. H. guppyi Hemsley and H. affinis Hemsley are closely related to this species. The blooms are greenish yellow, inside near the base reddish, the crowns are red on top. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. hollrungii Warb. – 15, 33, 36.

Hoya hookeriana Wight In Contributions to the Botany of India (1834) 37. Robert Wight & Wallich. 8. H. Hookeriana (Wight) volubilis glabra, foliis carnosis oblongo-lanceolatis 3-nervis (6— 10 uncias longis 1—1 ½ latis), pedunculis brevibus multifloris corolla (parvula) glabra segmentis obtusis, cor. St. foliolis ovatis acutis angulo interiore brevi, stigmate obtuso. ( Hoya, Wall.! Asclep. N. 28, 39 (ex parte). (Chittagong; Silhet; Wallich. (R. W.)) Translation: Twining glabrous, leaves fleshy oblong-lanceolate 2 nerved (6 to 10 inches long 1 to 1 ½ wide), peduncles short many-flowered, corolla (small) glabrous segments obtuse, with the lobes of the staminal corona ovate acute with the interior angle short, stigma obtuse, (I have seen the Hoya, Wallich Asclepias # 28, 39 (in part). In General System of gardening 4 (1837) 126. G. Don. 8. H. Hookeriana (Wight, contrib. ind. bot. p. 37.) twining, glabrous; leaves fleshy, oblong-lanceolate, 3-nerved; peduncles short, many-flowered; corollas small, glabrous, with obtuse segments; leaflets of corona ovate, acute, with the inner angles short; stamina obtuse. Perennial twining shrub. Native of Chittagong and Silhet. H. Nicobarica, R. Br. mss. is this or very similar species. Leaves 6-10 inches long, and 1 or 1 ½ broad. Hooker's Hoya. Shrub tw. In Synopsis Plantarum 6 (1840) 891. D. N. F. Dietrich 8. H. Hookeriana Wight; volubilis glabra; fol. Carnosis oblong-lanceolatis 3- nerviis; pedunc. Multifloris; cor. Segmentis obtusis. In Chittagong, Silhet. Translation: twining glabrous, leaves fleshy oblong-lanceolate 3 nerved; peduncles manyflowered; corona segments obtuse. In Prodromus Syst. Veg. 8 (1844) 636. Decandolle. 16. H. Hookeriana ( Wight ! contrib. p. 37). Volubilis glabra foliis. carnosis oblongo-v. elliptico-lanceolatis basi rotundatis v. subattenuatis apice acuminatis acutis marginibus reflexis, obscure 3-nerviis, pedunculatis petiolos superantibus multifloribus, pedicellis gracilibus, corolla glabra, laciniis obtusis, coron. Stam. Foliolis ovatis acutis angulo interiore brevi, stigmate obtuso perennial shrub in India orientali Silhet, Chittagong (Wallich). (v. s. h. Mus. Par.) Translation: Twining glabrous, leaves fleshy oblong or elliptic-lanceolate with base rounded or somewhat attenuate apexes acuminate acute margines reflexed, with 3

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obscure nerves, peduncles longer then the pedicels many-flowered , pedicels glabrous, corolla glabrous, leaflets obtuse, with the leaflets of the staminal corona ovate acute interior angle short, stigma obtuse. Perennial shrub in Eastern India Silheit, Chittagong (Wallich) (seen in part at the Museum.) In Handbook of Indian Flora 2 (1866) 241. W. D. Wight (6) H. Hookeriana (R. W.) Ident. Wight’s p. 37.—Dec. prod. VIII. p. 636. Spec. Char. Twining: leaves fleshy oblong-lanceolate, 3-neved: peduncles short, many flowered: corolla glabrous, segments obtuse: leaflets of the staminal crown ovate, acute, inner angle short: stigma obtuse. Chittagong. Silhet.

Hoya hypolasia Schlechter In Botanische Jahrbücher 50 (1913) 123-124. R. Schlechter “Die Asclepiadaceen von Deutsch Neu Guinea”. 31. H. Hypolasia Schltr. n. sp. — Suffrutex parum ramosus, alte scandens. Rami filiformes, flexuosi, laxe foliati, teretes, primum minute et molliter puberuli, mox glabrati, lenticellis verruciformibus numerosis obsessi. Folia patentia vel patula, petiolata, lanceolata, acuminata, basi subcordata, textura coriacea, superne glabra, lucida, subtus brevissime et dense puberula, petiolo superne sulcato, brevissime et molliter puberulo. Inflorescentiae umbelliformes, ca. 10-florae, longius pedunculatae, pedunculo puberulo, pedicellis gracillimis glabris. Calycis foliola parvula ovata, obtusa, margine breviter ciliata quam corolla multi minora. Corolla recurva usque ad tertiam partem basilarem 5-fida, extus glabra intus minutissime papillosa, lobis late rhombeoovatis, acutis. Coronae foliola compressa superne carinato-incrassata a latere visa oblique quadrata, apice brevi acuta, antheris paululo breviora. Pollinia obovoideo-oblongoidea, trabslatoribus prebrevibus, retinarulo oblongoideo minuto. Ein hochkletternder Schlinger mit dünnen schnurartigen locker beblätterten Zweigen. Blätter 12 (20 cm lang. Unterhalb der Mitte 3.3—5.3 cm breit, Blattstiel sehr kurz, weichaarig, etwa 10-blütig auf 5 cm langen kurz behaartem Stiel. Blütenstiel kahl, etwa 5 cm lang. Blüten denen der vorigen Art ähnlich. Kelch klein, ca. 2 mm lang. Korolla ausgebreitet ca. 2.1 cm im Durchmesser. Koronaschuppen von der Spitz bis sum äuszern Rande 3 mm lang, vorn 3.5 mm hoch. Nördl. Neu-Guinea: auf Bäumen in den Wäldern der Hügel am Wabe-Flusz, am Flusz des Finisterre-Gebirges ca. 400 m. ü. M. Schlechter n. 18075. ( Blühend im Juli 1908). Schon äuszerlich ist die Art vor H. piestolepis Schltr. Durch die behaarten später mit vielen Lenticellen überdeckten Zweige und die langen schmalen, unterseits dicht und kurz-weichaarigen Blätter vor dickerer Konsistenz zu unterscheiden. In der Korolla sind biede Arten einander äbnlich, doch in den Koronaschuppen durchaus verschieden, denn bie der vorliegenden sind diese bie weitem nicht so stark zusammengedrückt und viel kleiner als bei H. piestolepis Schltr, Die Blüten sind gelblich-weisz, auszen leicht rötlich überlaufen. Translation: See next below. In Hoyas of Northeastern New Guinea (1992) 96. R. D. Kloppenburg. (Translation from

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Die Asclepiadaceen von Deutsch Neu Guinea by R. Schlechter. 31. Hoya hypolasia Schlechter n. sp. - A high climbing half shrub with few branches. Branches threadlike, flexible, loosely leaved, round, at first minutely and softly puberulous, soon glabrous, occupied with numerous wart shaped lenticels. Leaves spreading or outspread, petiolate, lanceolate, acuminate, base somewhat cordate, leathery texture, glabrous above, lucid, shortly and densely puberulous below, petiole grooved above, very shortly and softly puberulous. Inflorescences shaped like an umbel, about 10 flowered, long pedunculate, peduncle glabrous, with the pedicels very slender. Leaflets (lobes) of the calyx small, ovate, obtuse, margins shortly ciliate, much shorter than from the corolla. Corolla recurved, 5 parted all the way to one third from the basal area, glabrous outside, very minutely papillose inside, lobes broadly rhomboid-ovate, acute. Leaflets (scales) of the corolla compressed above, thickly keeled above, scales visibly obliquely squared, apex shortly acute, anthers somewhat short. Pollinia obovoid-oblong like, with the translators short over-all, retinaculum oblong like. A high climbing clinging vine with slender cordlike, loosely leaved branches. Leaves 12-20 cm. wide. Petiole very short, softly pubescent, about 1 cm. long. Inflorescences umbellate, about 10 flowered, on a 5 cm. long, shortly hirsute peduncle. Pedicels bare, about 5 cm. long. Flowers like those of the previous species. Calyx small about 2 mm. long. Corolla spread out, about 2.1 cm. in diameter. Corona scales, from the tip to the outer edge 3 mm. long, at the front 3.5 mm. high. Northeastern New Guinea: on trees in the forest of the hills of the Wabe river at the foot of the Finisterre Mountains, about 400 m. altitude (Schlechter #18075 Blooming in July 1908). Indeed the species is distinguished from H. piestolepis Schlechter through the pubescence, branches subsequently covered with many lenticels and by the long narrow thickened leaves, whose undersides are thickly covered with short hairs. In the corolla, both species are similar but in the corona scales completely different, for in the ones in hand, they are by far not so strongly compressed and are much smaller than those of H. piestolepis Schlechter. The flowers are yellowish-white, outside lightly overlaid with red. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. hypolasia Schlechter (Schlechter 20451) – 15. In Dr. Schlechter’s Hoya Species (1993) 78-79. R. D. Kloppenburg. (Translation). Hoya hypolasia Schlechter. A high climbing half shrub with few branches. Branches threadlike, flexible, loosely leaved, round, at first minutely and softly puberulous, soon glabrous, occupied with numerous wart shaped lenticels. Leaves spreading or outspread, petiolate, lanceolate, acuminate, base somewhat cordate, leathery texture, glabrous above, lucid, shortly and densely puberulous below, petiole grooved above, very shortly and softly puberulous. Inflorescences shaped like an umbel, about 10 flowered, long pedunculate, peduncle glabrous, with the pedicels very slender. Lobes of the calyx small, ovate, obtuse, margins shortly ciliate, much shorter than from the corolla. Corolla recurved, 5 parted all the way to one third from the basal area, glabrous outside, very minutely papillose inside, lobes broadly rhomboid-ovate, acute. Scales of the corolla compressed

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above, thickly keeled above, scales visibly obliquely squared, apex shortly acute, anthers somewhat short. Pollinia obovoid-oblong like, with the translators short over- all, retinaculum oblong like. A high climbing clinging vine with slender cord like, loosely leaved branches. Leaves 12-20 cm. wide. Petiole very short, softly pubescent, about 1 cm. long. Inflorescences umbellate, about 10 flowered, on a 5 cm. long, shortly hirsute peduncle. Pedicels bare, about 5 cm. long. Flowers like those of the previous species. Calyx small about 2 mm. long. Corolla spread out, about 2.1 cm. in diameter. Corona scales, from the tip to the outer edge 3 mm. long, at the front 3.5 mm. high. Northeastern New Guinea: on trees in the forest of the hills of the Wabe river at the foot of the Finisterre Mountains, about 400 m. altitude (Schlechter #18075 Blooming in July 1908). Indeed the species is distinguished from H. piestolepis Schlechter through the pubescence, branches subsequently covered with many lenticels and by the long narrow thickened leaves, whose undersides are thickly covered with short hairs. In the corolla, both species are similar but in the corona scales completely different, for in the ones in hand, they are by far not so strongly compressed and are much smaller than those of H. piestolepis Schlechter. The flowers are yellowish-white, outside lightly overlaid with red. Published by Dr. R. Schlechter in: 1913 Botanische Jahrbücher 50:123-124, "Die Asclepiadaceen von Deutsch Neu Guinea".

Hoya imbricata Decaisne In Prodromus Syst. Veg. 8 (1844) 637. Decandolle. ** Folia orbicularia imbricata alterna. H. imbricata, scandens radicans, foliis abortu alternis unilateralibus, imbricatis orbicularilis supra conrexis aveniis subtus venois marginibus reflexis, pedunculis multifloris, pedicellis gracilibus glabris, calyce brevi, corollae laciniis triangularibus acutis, coronae stam. fol. supra convexiusculis marginibus revolutis angulo inter, porrecto antherarumque membrana attenuata stigmati apice: lato incumbentibus, folliculis laevibus; frutex scandens arbor, truncis conjunctissime affixus. Perennial Shrub in sylvis montium Calawan insula Philippines. Hoya imbricata Callery mss. H. Mus. Par. Deless. Icon. Select. Vol. 5 (v.s.h. Mus. Par.) Translation: climbing rooting, leaves aborted alternately unilaterally, imbricate orbicular above convex veinless below veined with reflexed margines, peduncles many-flowered, pedicels glabrous slender, calyx small, leaflets of the corolla triangular acute, leaflets of the staminal corona above convex with revolute margins interior angle produced outward and forward with the membrane attenuate to the stigma apex; broadly incumbent, follicles smooth; shrub climbing on tree trunks, tightly appressed. In Icones Selectorum 5 (1846) 37. Delessert. Hoya imbricata. Dcne in DC. Prodr. 8, p. 637. H. scandens radicans, foliis abortu alternis unilateralibus imbricatis orbicularibus supra convexis aveniis subtus venosis marginibus reflexis, pedunculis multifloris, pedicellis gracilibus glabris, calyce brevi, corollae laciniis triangularibus acutis, coronae stam. fol. supra convexiusculis marginibus revolutis angulo inter. porrecto antherarumque membrana attenuata stigmati apiculato incumbentibus, folliculis laevibus. Frutex

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scandens arbor. truncis conjunctissime affixus. Dne l.c. Crescit in sylvis montium Calawan insula Philippinis. Translation: same as above. In Flora Indiae Bataviae 2 (1857) 520. F. A. W. Miquel & Zollinger.. Folia abortu alterna. 13. Hoya imbricata Callery mss. Decaisn. In D.C. Prodr. VIII. P. 637. Scandens, radicans, folia abortu alterna uilateralia, imbricata, orbicularia, supra convexa avenia, subtus venosa, marginibus reflexis, pedunculi multiflori, pedicelli graciles glabri, calyx brevis, corollae laciniae triangulares acutae, coronae stamineae phylla supra convexiuscule marginibus revoluta, angulo interiore porrecto antherarumque membrana attenuata stigmati apiculato incumbentibus, folliculi laeves. — Frutex scandens arborum truncis conjunctissime infixus. Deless. Icon. Vol. V. p. 37, tab.90. ( Philipp. eil. In de wouden op het gebergte Calawan (Callery). Translation: same as two above. In Linnean Society of London Translations Botany (1888-1894) 495. CXXVI. (3501). Hoya R. Br. (Dcne. In DC prodr. VIII 634.) 1. H. imbricata Dcne. L.c. 637, no. 23 ( Deless. Icon. Select. Pl. V. 37, t. 90. In Fragmenta of the Florae of the Philippineae 1 (1904). (Perkins) R. Schlechter & O. Warburg. Hoya imbricata Dcne. Luzon Isl. Prov., Rezal, Tanay (Merrill no. 2363). In Philippine Journal of Science 15 (1919) 263. S. H. Koorders. Hoya imbricata Callery ex Decaisne und Hoya pseudomaxima Kds. In den Filipien auf Grund von Einigen Herbar-exemplaren des Bureau of Science in Manila. Von S. H. Koorders. Buitenzorg, Java. Mit vier tafeln. 1. Hoya imbricata Callery ex Decaisne, forma Typica Kds. Taf. I. Decaisne giebt in DC. Prodromus 8 (1844) 637 folgende von ihm in Delessert, Icones selectae plant. 5 (1846) 37 und später von Miquel, Flora Indiae Batav. 2 (1857) 520, unverändert übergenommene Spezies-Beschreibung von Hoya imbricata: Scandens radicans, foliis abortu alternis unilateralibus imbricatis orbicularibus supra convexis aveniis subtus venosis marginibus reflexis, pedunculis multifloris, pedicellis gracilibus glabris, callyce brevi, corollae laciniis triangularibus acutis, coronae stam. fol. supra convexiusculis marginibus revolutis angulo int. porrecto antherumque membrana attenuata stigmati apiculato incumbentibus, folliculis laevibus Frutex scandens arbor, truncis conjunctissime affixus. (Decne. l. c.) Von dieser characteristischen, bisher im Buitenzorger Herbar fehlenden Art, erhielt ich leihweise aus dem Herbarium des Bureau of Science in Manila, von Herrn E. D. Merrill, folgendes von ihm in den Filipinen gesammeltes Herbar-Exemplar: Luzon, Rizal Province, Tanay, Merrill 2363, ausgeblüht im Mai, 1903. Dieses Exemplar ist von Merrill etikettiert: Hoya imbricata Dcne., und von ihm mit folgendel Bemerkung versehen: "This should be typical H. imbricata. Leaves mottled, green and purple." Das Exemplar wurde nach seiner Angabe von ibm gesammelt ziemlich nahe bei dem Fundort von den authentischen Exemplar von Hoya imbricata des Pariser Herbar. Es besteht aus einem Blütenzweig mit 4 sich dachziegelig deckenden Blättern und einem

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ausgeblühten Blütenstand, und einer Papierkapsel mit einem einzehlen Laubblatt. Von allen mir aus dem Herbarium des Bureau of Science in Manila, unter dem Namen Hoya imbricata leihweise zugeschickten Herbarexemplaren, stimmt kein einziges so gut überein mit der Beschreibung und Abbildung von Hoya imbricata in DeCandolle Prodromus, und in Delessert, wie obengenanntes Spezimen (Merrill 2363), dass es als "Topotype" (im Merrillschen Sinne) von Hoya imbricata betrachtet werden darf. Über dieses bemerkenswerte Herbarexemplar (Merrill 2363) lasse ich folgende Beschreibung, folgen: Blätter kreisrund, ± ganzrandig oder eckig-auageschweift am Grunde abgerundet-abgestutzt, oben abgerundet oder emarginat; oberseits convex, ohne deutliche Nerven, völlig kahl und glatt (auch ohne Cuticular-Höcker); unterseits concav (auch ohne Cuticular-Höcker); ± in der Mitte dem sehr kurzen, fast fehlenden Blattstiel peltat inseriert und handnervig, mit undeutlichen, gabelig verzweigten Nerven; ± 9 cm lang und 10 cm breit. Epidermis des Blatt-Oberseite ohne Haare, mit ± glatter oder schwachwelliger etwa 20 µ dicker Cuticula, fast ohne Cuticular-Höcker und dadurch aussen ± glatt (nicht körnigrauh) aussehend. Epidermis Zellen des Blattes Unterseite je mit einem wagerecht abstehendem, einzelligem, unverzweigtem, fadenförmigem oder schmalkegelförmigem, spitzem, glattem, inhaltlosem, farblosem, gerade oder ± gekrümmtem, 30-50 µ langem Haar versehen und mit nur 5 µ dicker Cutieula. Stengel stieltund, ± 4 mm Durchmesser, kahl (auch ohne Papillen Cuticular-Höcker), mit zahlreichen, dichtstehenden Haftwurzeln, welche meistens ganz oder fast ganz von den dichig deckenden Laubblättern geschützt sind. Pedunculus axillar (in sicco abstehend), ± 10 cm lang, stielrund kahl; rhachis ± 9 cm lang, kahl, dicht bedeckt mit den Narben der abgefallenen Pedicelli, spindelförmig, in der Mitte fast 1 cm breit. Dieses Exemplar (Merrill 2363) ist von mir Hoya imbricata Callery ex Decaisne, forma typica Kds. (msc. 1918) etikettiert worden. 2. Hoya imbricata Callery forma basi-subcordata Kds. forma nov. Taf. II, III. A typo recedit foliis basi subcordatis. Blätter kreisrund, ± ganzrandig, oben abgerundet, am Grunde bis 1 cm (oft nur 0.5 cm) untief-herzförmig, ± in der Mitte peltat, dachig deckend, bisweilen (Copeland 399) auseinander entfernt. Blüten ± wie der Typus. Früchte unbekannt. Die folgenden 3 Exemplare des Manila-herbers halte ich für diese neue Form (foliis basi-subcodatis) von Hoya imbricata: Biliran, Bur. Sci. l8893 McGregor, blühend im Juni, 1914. Mindanao, Davao District, bei Davao, Copeland 399, blühend im März, 1904. Samar, Bur. Sci. 24910 Edano. Diese Exemrplare stimmen Beide im Blütenbau mit der Beschreibung und Abbildung von Hoya imbricata in Decaisne gut überein, und besitzen obenso wie auch Merrill 2363, beiderseits völlig kahle Blätter (die auch keine Cuticular-Höcker haben), jedoch ist der Blattfuss deutlich "subcordat". Die Tiefe des herzförmigen Blattfuss-Einschnittes beträgt ± 0.5-1 cm. Die Blüten von Bur. Sci. 18893, McGregor sind noch nicht völlig ausgebidet und kleiner als von Copeland 399. Bei beiden Exemplaren liegen keine Früchte vor. Mit obiger forma basi-subcordata zeigt folgendes Filipinen-Exemplar einige Ähnlichkeit, jedoch ist die Korolla auf innen auffallend dicht behaalt: Samar, Catubig River, Bur. Sci. 24910 Edano. Blühend im Februar-März, 1916. Diesem Exemplar ist von Merrill folgende Bemerkung hinzugefügt: "Flowers appear to be different from Hoya imbricata."

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Auch bei diesem Exemplar sind keine Früchte vorhanden. Vielleicht liegt hier eine von Hoya imbricata neue Spezie vor. 3. Hoya pseudomaxima Kds. sp. nov. Taf. IV. A Hoya imbricata Callery foliis basi profunde cordatis et a Hoya maxima Kds. foliis supra glaberrimis et apice haud raro emarginatis differt. Luzon, Rizal Province, Bosoboso, Bur. Sci. 22089 Ramos, blühend am 7ten December, 1913. Blätter kreisrund, am Grunde bis 3 cm tief-herzförmig, oberseits völlig kahl (nicht nur unbehaart, sondern auch ohne Cuticular-Höcker) und glatt, unterseits kahl und auch ohne Cuticular-Höcker, bis 8 cm lang und 9 cm breit. Blüten ± wie Hoya imbricata, Früchte unbekannt. Bemerkung: Während bei allen (12) Einsammlungs-Nummern meiner Hoya maxima aus Nord-Ost-Celebes (Kds no. 16204, ß U.S.W.) die Blatt-Oberseite stets mit Cuticular-Höcker dicht bedeckt ist, welche dauernd oder seltener nur in der Jugend je ein Flaumhaar tragen, bei der nur aus den Filipinen (Mount Isarog) bekannten Hoya pseudomaxima ist die Blatt-Oberseite völlig glatt (g1aberrima) ohne Cuticular-Höcker und ohne Flaumhaare. Translation: In the Philippines because of some sharp-copies in the office of Science in Manila. From S. H. Koorders. Buitenzorg, Java. With four sheets. 1. Hoya imbricata Callery ex Decaisne, forma Typica Kds. Taf. I. Decaisne from DC. Prodromus 8 (1844) 637 following from him in Delessert, Icones selectae plants. 5 (1846) 37 and later from Miquel, Flora Indiae Bataviae. 2 (1857) 520, unchanged from the species-description of Hoya imbricata: Climbing rooting, leaves aborted alternate and unilateral imbricate and orbicular above convexis with no veins below veined margins reflexed, pedoncules multi-flowered, pedicellis narrow glabrous, calyx small, leaflets of the corolla triangula acutis, leaflets of the staminal corona above slightly convex with revolute margines, interior angle incumbent upon the apiculate stigma, anther membrane attenuate, follicles smooth. A twining climber in the trees. Affixed to the trunks. (Decne. l. c.) From these characteristics, lacking of the type until now in the Buitenzorger Herbar, I obtained from the herbarium of the office of Science in Manila, from Mr. E. D. Merrill, following from him on loan in the Philippines, collected a sharp copy: Labeled Luzon, Rizal Province, Tanay, Merrill 2363, in the May, 1903. This copy is labeled by Merrill: Hoya imbricata Dcne., and from him provided with following remark: “This should be typically H. imbricata. Leaves mottled, green and purple”. The example was collected after his statement from near by, rather approaches Hoya imbricata of the Paris Herbar at the place of discovery of the authentic specimen of this Hoya. It consists of a bloom-branch with 4 roof tile-like covering leaves and a faded inflorescence, and a fruit-capsule with one detailed foliage page. All for me from the herbarium of the office of Science in Manila, Herbarium examples sent under the name Hoya imbricata on loan, no single agreement of Hoya imbricata with the description and illustration of Hoya in DeCandolle Prodromus, and in Delessert, so well as the afore mentioned Specimen (Merrill 2363), that is the Topotype (in the Merrill’s sense) may be viewed as Hoya imbricata. Over this remarkable Herbar example (Merrill 2363) I give the following pages of description: Leaves circular-round, ± with entire edges or angular-indentation at the base without rounding-truncate, rounded off above or emarginate; otherwise convex, without clear

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nerves, completely glabrous and smooth (also without cuticular-protuberances); undersides concave (also without cuticular-protuberances); ± in the middle very short, almost lacking petioles peltate inserted and palmately-nerved, with un-clear nerves branching forked; ± 9 cm long and 10 cm wide. Epidermis of the leaf upper side without hair, with ± smooth or weakly wavy, approximately 20 µ thick cuticle, almost without cuticular-protuberances and through it outside ± smoothly (not granulated-roughly) looking. Epidermis cells of the under-side of the blade ever with a single-cell horizontally sticking out, unbranched, threadlike or narrowconical, pointed, smooth, content-less, colorless, exactly or ± bent, provided with 30-50 µ long hairs and with only 5 µ thick cuticle. Stem peduncle, ± 4 mm diameters, glabrous (also without Pa-pillen cuticular-protuberances), with numerous, dense-standing customary rooted, which is protected by the close quite or almost completely mostly covering foliage-blade. Pedunculus axillar (when dry sticking out), ± 10 cm long, stalk-approximately bald; rhachis ± 9 cm long, glabrous, pedicels dense, spindle-shaped, almost covered with the scars of the abgefallen-en in the middle 1 cm wide. This copy (Merrill 2363) has been labeled by me Hoya imbricata Callery ex Decaisne, forma typica Kds. (msc. 1918). 2. Hoya imbricata Callery forma basi-subcordata Kds, forma nov. Taf. II, III. A typical receding foliage at the base subcordate. Leaves round-circular, ± entirely complete, rounded off above, at the base with 1 cm. (often only 0.5 cm) shallowly heart shaped, ± in the middle peltate, roof covering, from time to time (copeland 399) some removed. Blooms ± like the figure. Fruits unknown. I consider the following 3 copies from the Manila Herbarium as this new form (formis basi-subcordata) of Hoya imbricata: Biliran, Bur. Sci. 18893 McGregor, blooming in June 1914. Mindanao, Davao District, near Davao, Copeland 399, blooming in March 1904. Samar, Bur. Sci. 24910 Edano. These examples agree both in the bloom-construction with the description and illustration of Hoya imbricata with good agreement, and are also like Merrill 2363 mentioned above on both sides of the leaves completely bald (that also have no circular indentations) however the leaf base is clearly subcordate, the depth of the heart-shaped leaf-foot is ± 0.5-1 cm. The blooms of McGregor's Bureau of Science 18893 are not yet completely fully formed and smaller then Copeland 399. No fruits are available for either specimen. With the above forma basi-subcordata shows some similarity to the following Filipine example, however the corolla internally is notably open: Samar, Catubig River, Bureau of Science 24910. Edano. Blooming in February-March., 1916. The following remark is added by Merrill: Flowers appear to be different from Hoya imbricata, also with this copy there is no existing fruit. Maybe a new species differing from Hoya imbricata is available here. 4. Hoya pseudo-maxima Kds. sp. nov. Taf. IV. From Hoya imbricata Callery leaf base definitely cordate, and from Hoya maxima Kds. Leaf above glabrous at the apex not at all emarginate, differing. Luzon, Rezal Province, Bosoboso, Bureau of Science 22089 Ramos, blooming on the 7th of December, 1913. Leaves circular round at the base 3 cm. Deep, heart shaped, otherwise entirely glabrous (not only hairless but also without cuticular bumps) and smooth, undersides

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glabrous and also without cuticular bumps, up to 8 cm long and 9 cm. Wide. Blooms ± like Hoya imbricata, fruits unknown. Remarks: while all (12) of the numbered collections of my Hoya maxima from the north-Eastern Celebes (Kds. Number 16204  and so on) the upper side of the leaf always has cuticular bumps which is constant or more rarely in youth with a little fuzzy-hair, which is known in the Philippines only from (Mount Isarog) Hoya pseudo-maxima with the with the upper side of the leaf completely smooth (glabrous) and without cuticular bumps and without fuzzy-hairs. In Bulletin du Jardin Botanique Series III 2 (1920) 247-248. S. H. Koorders. Untergattung Paraconchopllum Koord.—Blätter mit der unteren Seite ausgehöhlt, dachziegelig deckend und dem Substrat (ameisten’s Baumstämmen) dicht anliegend. Von jedem Blattpaar geht in ….hester Jugend stets ein Blatt zu Grunde und kommt nur je ein Blatt zur Sntwicklung. Die erwachsenen, grosser, kreisrunden Blätter sind daher Scheinbar abweschselnd, und nur iu frühester Jugend (in der Knospenlage) gegenstäbdig. Hieher 3 oder 4 nahe verwandle Arten: 1. Hoya imbricata Ca11ery ex Decaisne DC. Prodr. VIII (1844) p.637; Decaisne in Delessert: Icones selectae V. (1846) p. 37. Tab. 90. Filippinen.— Zu dieser Art gehören: forma typica Kds. (leg. Merrill und forma basisubcordata Kds. msc. in Herb. Bureau of Science in Manila (leg. Edano, McGregor). 2. Hoya maxima (Karsten) Koord. Nordost-Celebes. Durch vergleichung von meinen Minahasa-Exemplaren mit einigen mir freundlichst von Herrn E. D. Merrill, Botanist of the Bureau of Science in Manila, aus dem Herbarium of the Bureau of Science in Manila geliehenen Filippinen Exemplaren konnte ich feststellen dass bei H. imbricata aus den Filippinen der Blattfuss sehr untief-buchtig-ausgeschweift oder ± abgestutzt ist und niemals, wie bei Hoya maxima, tief (herzförmig) und dass Hoya maxima im Filippinen-Herbar fehlt. 3. Hoya pseudomaxima Kds. (msc. Juni 1918 in Herb. Bureau of Science in Manila). Hoya imbricata. Cailery fo!iis basi profunde cordatis et Hoya maxima Kds. foliis supra glaberrimis et apice haud raro emarginatis differt.- Filippinen (leg. M. Ra mos). wähend bei Hoya maxima die Blätter stets einen tief-herzförmigen Fuss besitzen mit Fusslappen; welche ± einander berühren oder ± übereinander greiten. In Engler und Prantl, Pflzfam. IV. 2 (1895) p. 289 sagt Schumann folgendes: “Streng der Priorität nach hat O. Kuntze Recht wenner Sperlingia Vahl bevorzugt. Da die Gattung (Hoya) aber eine Menge cultivierten, darunter eine aligemein verbreitete Pflanze umsasst so ziehe ich es vor, Hoya, den gewohnten Namen zu belassen." Jedoch geben Dalla Torre et Harms, Genera siphonog. (1900-1907) folgende Angaben: Hoya R. Br. (in: Mem. Werner Soc. 1 (1809) 26 ) und Sperlingia Vahl. (in: Skrirt. naturh. Selsk. Kjöbenhava VI (1810) 113.) Aus den Angaben von Harms und Dallal Torre muss geschlossen werden, dass Hoya als Gattung die Priorität hat oberhalb Sperlingia. Es scheint desshalb besser Hoya den gewohuten Namen der Gatiang zu belassen. Die von Schumann, Bentham et Hooker und von anderen eingezogene Gattung Sperlingia wähle ich zur Bildung der neuen Untergattung “Eusperlingia." Morphologische und systematische Notizen. (Bei Herb. Kds. n. 16199  wulde

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folgendes über die Blätter notiet: Blattoberseite unter der Lupe gesehen ± dicht-flaumhaarig und papillös, unterseits unter der Lupe ± kahl. Unter dem Mikroskop ist die Epidermis der Blattoberseite ohne Spaltöffnungen und-mit ± lockerstehenden ± 1/3 mm (300µ) langen, fadenförmigen, unverzweigten, dünnwandigen, oben spitzen; geraden; oder gekrümmten, 2—4 zelligen, farblosen Haaren bedeckt, welche je einzelin auf einer höckerartig-papillösen Erhebung der Cuticula inseriert sind. Ausserdem befinden sich auf der Blatt-Oberseite sehr zahiretche 20—25 µ.1ange, schief aufwärts gerichtete oder ± anliegende dickwandige, höckerartig hervorragende Cuticular-warzen und zwar je eine auf jeder Epiderlmis Zelle, die kein Flaumhaar trägt. Die Epidermis der Blattunterseite ist mit dichtstehenden, sehr kurzen, mit Luft erfüllten, kegelförmigen, 20-50 µ langen Haaren bedeckt, welche je einzeln einer Epidermiszelle aufsitzen,sehr dick wandig und hyalin sind. Auf der Blattunterseite befinden sich im Gegensatz zu Blattoberseite (vergl. oben) eine sehr grosse Anzahl Spaltöffnungen. Bei Herb. Kds.(n.16199 , 16204 , usw.) wurde u. a. folgendes notierl. Mit Haflwurzein epiphytisch-kletterndes Kraut, oft 10 Meter oder mehr hoch. Blälter dachziegelig deckend, durch Abort abwechselnd, wie kleine Teller dem Substrat flach anliegend, oberseits matt oder schwach glänzend schmutzig-grün, bisweilen oberseits purpurn und blassgrün gefleckt oder oberseits fast ganz dunkelpurpurn, unterseits stefs matt-dunkelpurpurnbraun oder dunkelpurpurn. Blüten Abends stark-süsslich-duftend, jedoch während des Tages nach einigen Notizen fast geruchlos. Pedicelli und Aussenseite des Kelches schmutzig purpurn oder blassbräunlich. Korolle derb, mit zurückgeschfagenen Zipfeln, welche beiderseits gelblich-weiss sind und oberseits (innen) mit dichtstehenden, sammetartigen, langen, weissen Haaren bedeckt sind. Corona aussen violettpurpurn, innen weisslich. Antheren schön gelb. Die Stigmata der geöffneten Blüte sind “mutica". (und nicht apiculat), jedoch besitzen sehr junge Blütenknospen ein, “Stigma apiculatum''. Translation: Leaves with the undersides hollowed out, above imbricate and the substratum (ameisten's tree-trunks) densely covered. From each leaf-pair when in the youth always a leaf aborts and only one leaf remains at a node. The mature, larger, round leaves alternate, and only in of very earliest youth are therefore seemingly (in the budsituation) somewhat shrub like. There are 3 or 4 near transformed types: 1. Hoya imbricata Ca11ery ex Decaisne DC. Prodr. VIII (1844) p.637; Decaisne in Delessert: Icones selectae V. (1846) P. 37. Tab. 90. Filippinen. - to this type belongs: forma typica Kds. (leg. Merrill and forma basisubcordata Kds. msc. in sharp. Office of Science in Manila (leg. Edano, Mc Gregor). 2. Hoya maxima (Karsten) Koord. Nordost-Celebes. Through comparison of my Minahasa-Examples with some I have found of Mister E. D. Merrill, Botanist of the office of Science in Manila, from the herbarium of the office of Science in Manila lent Filippinen copies I could determine I that near with H. imbricata from the Philippines the leaf-base very much differed from the shallow sinate or ± truncate is and never, as with Hoya maxima, profoundly (heart-shaped) and that Hoya is missing maxima in the Filippinen-Herbar. 3. Hoya pseudomaxima Kds. (msc. June 1918 in sharp. Office of Science in Manila). Hoya imbricata. Cailery leaf base profoundly cordate and Hoya maxima Kds. leaves above glabrous and the apex not at all rarely emarginate are different. - Filippinen

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(leg. M. Ramos). As with Hoya maxima the leaves always a deep-heart-shaped bases possess with a foot tab; which ± touches each other or ± grasps one on the other. In Engler and Prantl, Pflzfam. IV. 2 (1895) page 289 Schumann says the following: Severely according to the priority O. Kuntze has preferred rightly Sperlingia Vahl. Since the type (Hoya) however has been used in quantity, under it a universally plant usage is embraced so I prefer it, Hoya, the accustomed name to leave as it is. However Dallas give Torre and Harms, Genera siphonog. (1900-1907) following statements: Hoya R. Br. (in: Mem. Werner Soc. 1 (1809) 26) and Sperlingia Vahl. (in: Skrirt. naturh. Selsk. Kjöbenhava VI (1810) 113.) From the statements of Harms and Dalla Torre must be concluded that Hoya as type has the priority above Sperlingia. It seems on that account that Hoya the accustomed names of the genera is better to leave as it is. That of Schumann, Bentham et Hooker and from others drafted the genera Sperlingia chooses Eusperlingia which I propose the name of the new subspecies. “Eusperlingia”. Morphological and systematic notes.—With herbarium Koorders. number 16199  would following over the leaf notations: leaf-upper side under the magnifying glass seems to be ± dense-fuzz-hairy and papillös, the undersides under the magnifying glass ± glabrous. Under the microscope, the epidermis of the leaf-upper side has slit orifice and with ± relaxed-standing ± 1/3 mm (300µ) longs, threadlike, undeniably, thin-walled, above sharpened; straight; or bent, 2-4 celled, which with detail on one fine humppapillose colorless hairs, which are inserted on the cuticle covers. Furthermore one finds very much on the leaf-upper side very numerous 20-25 µ.1ange hairs, crooked in an upward directional or ± to be done thick-walled, fine humps outstanding cuticle-warts in fact one per each Epidermal cell, that carries no fuzz-hair. The epidermis of the leafunderside is with dense-standing, very short, with air filled, conical, 20-50 µS long hair covers, which always are separately placed on a epidermal-cell, which are very thick walled and hyaline. On the leaf-underside is in contrast to leaf-upper side (as above) a very big number of split-openings. With herbarium Kds. (n.16199 , 16204 , etc) became and the following notation. With clamping roots an epiphytisch-climbing herb, often 10 meters or more high. Leaf imbricate, through alternating abortion, affixed as small plates flat to the substratum, upper sides weak or weakly brilliantly dirty-green, from time to time the above side purple and pale-green stained or outsides almost completely dark-purple, undersides constant flat-dark-purple-brown or dark-purple. Blooms of evening strong-fragrant-scent, however during the day after some notes almost odorless. Pedicelli and outside of the cup dirty purple or faintly somewhat brown. Corolla rough, with corner flaps reflexed, which on both sides is short and topsides (centrally) with dense-standing, concentration, which is covered with long white hairs. Corona outside violet-purple, internally whitish. Anthers beautifully yellow. The Stigmas of the opened bloom are without points. (and not apiculate), however in very young bloombudding possess one, “stigma apiculatum''. In An Enumeration of Philippine Flowering Plants 3 (1922) 352. E. D. Merrill. Hoya imbricata Dcne, in DC Prodr. 8 (1844) 637, Deless Ic. 5 (1846) 37, t. 90; Miq. Fl. Ind.

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Bat. 2 (1857) 520; F-Vill. Novis. App. (1880) 135; Schltr. In Perk. Frag. Fl. Philip. (1904) 130; Koord. In Bull. Jard. Bot. Buitenz. III 2 (1920) 247. Collyris major Naves in Blanco Fl. Filip. Ed. 3 (1877-83) t. 212, non Vahl. Luzon (Bontoc, Rizal, Laguna), Busuanga, Merrill 2363, Vanoverbergh 2161, B. S. 41265 Ramos. On tree trunks at low and medium elevations. In Philippine Journal of Science 23 (1923) 263. E. D. Merrill. Hoya imbricata Deene. in DC. Prodr. 8 (1844) 637, Deless. Ic. 5 (1846) 37, t. 90; Miq. FI. Ind. Bat. 2 (1857) 520; F.-Vill. Novis. App. (1880) 135; Schltr. in Perk. Frag. Fl. Philip. (1904) 130; Koord. in Bull. Jard. Bot. Buitenz. III 2 (1920) 247. Collyris Major Naves in Blanco Fl. Filip. ed. 3 (1877-83) t. 212, non Vahl. Luzon (Bontoc, Rizal, Laguna), Busuanga, Merrill 2363, Vanoverbergh, 2161, B. S. 41265 Ramos. On tree trunks at low and medium altitudes. Var. basicordata* Koord. In Philip. Journ. Sci. 15 (1919) 264, t. 212. Luzon (Sorsogon), Biloiran, Samar, Mindanao (Davao), B. S. 18893 McGregor. 24910 Edano, Elmer 15802, Copeland 399, Warburg 14388. On trees at low altitudes. Endemic. * Note: this is var. basisubcordata Kds. In Leaflets of Philippine Botany 10 (1938-39) 3581-3582. A. E. D. Elmer. Hoya imbricata basicordata Koord. Tightly climbing or cleaving to smooth surfaced medians sized trees of the hemp region at 1000 feet elevation; stem wiry, occasionally branched and forming rootlets; blades very thick and rigid, plate tilde, atropurpureus on the lower concave side which is provided with a rigid root system and among which ants live in a symbiotic relation with the plant, the upper outer convex side is mostly of a dull glaucous green; pedicels also green, very flexible; the odorless flowers with much recurred petals which are on their inner sides at least. Number 15802 collected by Elmer at Irosin.—Rarely collected but not rare nor inaccessible. In Plant Life of the Pacific World (1945) 98. E. D. Merrill. Perhaps the most peculiar of all, however, are those leaf adaptations noted in such groups as Hoya, Dischidia, and Conchophyllum, all vines with abundant milky sap belonging in the milkweed family (Asclepiadaceae). Some of these hang free as epiphytes or semi-epiphytes, but in Conchophyllum and in some species of Hoya the vines may be closely appressed to the tree trunks or branches, while the circular leaves, one row on each side of the slender stem, are convex with their margins very closely appressed to the bark (Hoya imbricata, fig. 91). Under each leaf many roots are produced from the leaf axil which often quite cover that part of the bark protected by the leaf, serving to hold the plant in place and to absorb moisture and nourishment for the needs of the plant; each of these ready-made homes under each leaf is occupied by colonies of small ants. In Medicinal Plants of the Philippines (1978) 750 Quisumbing. Hoya imbricata Decne. Collyris major Naves. Local name: Paui-pauikan (Bis.). Paui-pauikan is an endemic species found on tree trunks at low altitudes in Bontoc, Rizal, and Laguna Provinces in Luzon; and in Busuanga.

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This is a smooth, woody vine and climbing or cleaving to tree trunks. The stems are wiry and form rootlets. The leaves are broadly ovate or rounded, 6 to 12 centimeters wide and heart-shaped at the base, rounded at the tip, somewhat leathery and rigid, and reddish, purple; the upper surface is convex and smooth and the lower surface concave, hairy, and inhabited by ants. The inflorescences are borne on leafless stems, are almost stalkless, and have many small, odorless, velvety, and creamy-white flowers 8 to 10 millimeters in diameter. The corolla-lobes are very slender, pointed, reflexed, smooth on the outside, and hairy within. The leaves are applied externally as a poultice to ripen boils. Sulit 567 reports the following) use: a number of the leaves are burned until charred. Then they are triturated, mixed with coconut oil, and stirred thoroughly. The mixture is applied to old wounds or varicose ulcers "bangkukang” in Tagalog for rapid cicatrization. In The Asclepiadaceous Works of Friedrich Richard Rudolf Schlechter (1992) 30. A. Nicholas. H. imbricata (Decaisne) – 32. In Philippine Hoya Species 3rd Ed. (1996) 83-84. R. D. Kloppenburg. Hoya imbricata Decaisne in D. C. Prod. 8 (1844) 637. Type: specimen at the Paris Museum Herbarium. Section Peltostemma Schlechter. Epiphytic, climbing, rooting especially at the nodes, one leaf aborting or not formed, the remaining leaf convex on upper surface and appressing closely to tree trunks, imbricate, orbicular, 8-10 cm. in diameter or more, upper surface rough, sparse ciliation, channeled slightly on the variety "basi-subcordata" , with a acute mucronate tip, the under surface purple in color, concave, finely granulose, sparse cilia mostly along the almost enclosed midrib and around the point of attachment to the stem, veinless, a mat of fine rootlets attaching themselves to the tree trunks. Peduncles axillary 10 cm. long, rachis enlarged and scarred. Umbel geotropic, concave, many fuzzy cream flowers. Pedicels glabrous, curved, variable length, flexible, green. Calyx lobes narrow elliptic, apex very acute, reaching corolla sinus, glabrous. Corolla, 0.8-1.0 cm. in diameter, reflexed, slender, pointed, glabrous outside, inside surface densely ciliate. Corona lobes convex, above margins revolute, inner apex stretched upward and forward at a steep angle, outer portion broad and rounded. Retinacula visible in open flowers. Anther appendages very long overtopping inner coronal apex noticeably, the 5 apices fanning out. Seed brown narrow, 0.7 cm. long outer apex sharply acute, base attached to long silky coma about 1.4 cm. long. Pods linear tapering to both ends, glabrous, straight to slightly curved, distal apex slenderly acute. Collected at Luzon (Botoc, Rizal, Laguna) Busuanga. On trunks of trees at low altitudes. References: Delessert, Icones Selectorum 5 (1846) 37; Zollinger and Miquel, Flora Von Nederlandsch Indie 2 (1856) 520; Naves in Blanco Flora Philippines (1877-83) t. 212; F.Vill. Novis. App. (1880) 135; Linnean Society of London, Translations Botany (1888-94) 495; De Nuttige Plantae Von Nederlandsch Indie 248; Bulletin des Jardin Botanique Series 3, 2 (1920) 247-8; Quisumbing, Medicinal Plants of the Philippines (1951) 22; Perkins, Fragmentia Florae Philippines (1904) 130; Philippine Journal of Science #3, 15 (1919) 262-7; Enumeration of Philippine Plants 3 (1923) 252; E.D. Merrill, Plant life of the Pacific World (1944), 98 fig.91; Leaflets of Philippine Botany 10 (1938) Art. 131, 3581-2.

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Herbarium Sheets: Merrill #2363 1903 (A); Vanoverbergh #2161; Ramos #41265 1907 (pseudomaxima), #41223, #22089 1913, #970; Elmer #15802 1916 (A); Copeland #399 (1904); McGregor #18893 (var. basi.); Edano #24910 1916 (var. basi.), #34523 1956 (PNH,A), #11015 1949 (BO, PNH); Mendoza #91296 1964 PNH), #91238 1964 PNH); Steiner #40033 1959 (PNH); McVittie & Carran #13835 1950 (PNH); Paniza #9395 1949 (PNH); Wayet #41559 1986 (var. basi.) (CAHUP); Vichanco #2239 1951 (CAHUP), #2241 1949 (CAHUP); Villaraza #2242 1913 (CAHUP); Loher No # 1909 (UC), No # 1912 (UC); Piper #133 1911; Koorders #16119. #16204; Warburg #14388 (var. basi.), #15848 (maxima); Sarasin #360 1894. Schlechter #20405 1909 (maxima). Meaning: imbricata — L. imbrco, to cover, with leaves shingle fashioned.

Hoya imperialis Lindley In Botanical Register sub t. 68 (1846) Lindley. Malaya (Edward’s Ornamental Flower Garden and Shrubbery). New Garden Plant. Hoya imperialis. H. imperialis (Hoyae verae); caule volibili, petiolis costaque tomentosis, foliis coriaceis anguste oblongis ciliatis glabris aveniis apiculatis apice recurvis basi rotundatis v. leviter cordatis, pedunculis tomentosis pendulis foliis brevioribus, umbellis multifloris, sepalis ovatis obtusis tomentosis, corollae maximae lobis triangularibus stellatim patulis ciliatis intus laevibus fauce tomentosa, coronae stamineae foliolis compressis bilobis lobo interiore acuto subdentato postico ovato obtuso. This is the most noble climbing plant we have ever seen. Beautiful specimens in flower have for some months been in our possession, sent from Borneo by Mr. Lowe, Junr.; but we have refrained from publishing an account of them, under the supposition that no living plant had reached England. We are now, however, able to state, that the plant is in the possession of Mr. Lowe of Clapton, who has already begun to put it into the trade. Imagine, then, a true Hoya, with wooly stems, leaves six inches long, and clusters of the most magnificent flowers, forming a diadem of ten rays; each flower fully three inches in diameter, and with the delicate texture of the common Hoya carnosa, and he will have some notion of this superb species. In Mr. Lowe’s letter from Sarawak, dated January 12, 1846, we have the following account of its discovery. “On the next day, when in the territory of the Gumbang Dayaks, I found another curious plant, belonging to Asclepiads; it is epiphytic climber; there was but one individual, growing from the decayed part of a tree, also overhanging the river. The flowers are large and in umbels; the leaves are leathery; and the stem abounds in white, perhaps acrid, juice. The contrast between the purple of the petals and the ivory white of the parts of fructification renders it highly beautiful.” This species is certainly new, unless it should be the Asclepias Sussuela of Roxburgh, a Moluccan plant, said to have flowers nearly three inches in diameter; but that botanist cites, without any doubt, the Corona Ariadnes of Rumphius, which has flowers only as large as a shilling (denarius), and therefor cannot be the species now described. Neither can this be the Hoya speciosa of Decaisne, which has the flowers velvety inside, and only one inch and three-quarters across; nor the Hoya grandiflora of Blume, which has leaves wooly beneath. Those glorious species are still to be imported, one from Java, the other from Amboina, and either would form an invaluable addition to our gardens.

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Translation: Stems twining, petioles thick tomentose, leaves leathery narrowly oblong, ciliate, glabrous, without veins, apiculate, recurved apex with the base rounded or slightly cordate, peduncles tomentose, hanging down smaller than the leaves, multiflowered umbels, sepals ovate obtuse tomentose, lobes of the corolla large triangular, star shaped outspread ciliate inside smooth and with the throat tomentose, leaflets of the staminal corona compressed, bilobed, interior lobes acute somewhat tooth-like outer ovate obtuse. In Repertorium Botanices Systematicae (1846) 743 (493). Under Additamentia. Pag. 493. No. CXXV. 1. Hoya imperialis Lindl. Bot. Regist. (New Ser.) 1846. And. Ad t. 68.( Caule volubilis, petiolis costaque tomentosis; foliis coriaceis anguste oblongis ciliatis glabris aveniis apiculatis, apice recurvis, basi-rotundatis vel leviter cordatis; pedunculatis tomentosis pedunculis, folia brevioribus; umbellis multifloris; sepalis ovatis obtusis tomentosis; corollae maxime lobis triangularibus steliatim patulis ciliatis, intus laevibus, fauce tomentosa; coronae stamineae phyllis compressis bilobis, lobo interiore acuto subdentato, postico ovato obtuso. Cresent in insula Borneo. Translation: Stems twining, petioles and midrib tomentose; leaves leathery narrowly oblong ciliate, glabrous veinless apiculate, apex recurved, base rounded or slightly cordate; peduncles pendulous and tomentose, leaf short; umbels many-flowered; sepals ovate obtuse tomentose; lobes of the corolla large triangular star-like outspread ciliate, inside smooth, throat tomentose; leaflets of the staminal corona bilobed compressed interior lobes acute somewhat tooth-like, the back ovate obtuse. Lives in the island of Borneo. In Botanical Magazine (1848) t. 4397. (Curtis’s) Hooker. Hoya imperialis. Imperial Hoya. Nat. Ord. Asclepiadeae,—Pentandria Digynia. Gen. Char. Calyx, brevis pentaphylluls. Corolla rotata plus minusve alte 5-fida, laciniis planis v. reflexis, aestivatione valvata. Corona staminea 5-phylla, foliolis depressis patentibus v. plus minusve gynostegio verticaliter adnatis, carnosis angulo interiore in dentem antherae incumbentem producto. Gynostegium breve. Antherae membrana terminatae Massae pollinis basi affixae, oblongae, comprtssae, connivientes, saepius margine pellucidae. Stigma muticum cum papilla media obtusa v. subapiculatum. Folliculi laeves v. appendiculis instructi, subpolypteri. Semina comosa.—Frutices v. suffructices Indica v. Moluccani, rarissime Africani, volubiles, scandentes aut decumbentes, foliis carnosis v. coriaceis v. membranaceis, floribus umbellatis umbellis extra-axillribus saepius multifloris. Dcne. Hoya imperialis; volubilis, ramis petiolis pedicellisque pubesccntibus, foliis (spithamaeis et ultra) obovato-lanceolatis coriaceis brevissime acuminatis subobscure parallelim venosis, pedunculo foliis longiore flexuoso-pendulo umbella pluriflora, floribus maximis purpureo-fuscis nitidis, corollae lobis cordato-triaugularibus marginibus axillas versus praecpue reflexis fauce elevata libera, coronae stamineae albae foliolis in medio arcte approximatis compressis carnosis bilobis lobo exteriore crasso dorso plano, interiore dentiformi lobum est. aequante. Hoya imperialis Lindl. Bot. Reg. 1846, sub. fol. 68. Hoya Sussuela. Roxb. Fl. Ind. v. 2. p. 31 ? Dr. Lindley does not say too much of this plant when he remarks, in the place above quoted, “this is the most noble climbing plant we have ever seen:" and this was spoken

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of the dried specimens we believe, aided by flowers preserved in spirits. With greater truth may it be said of the living plant, now that we have had the pleasure to see it exhibited in full flower at the Horticultural Fete in the Regent's Park Gardens (June, 1848), where it obtained the highest prize given for new plants, and again in the stove of the fortunate possessors of this rarity, Messrs. Lucombe, Pince, and Co., at their Exeter Nursery. A cluster of its flowers is indeed one of the most striking objects we have ever seen; the leaves too are large and handsome. It is a Hoya with glossy fleshy leaves of a deep purple chestnut colour, having, the expanded flowers full three inches in diameter! rendered more conspicuous by the ivory-white of the central column of fructification. It was detected in Borneo by H. Low, Jun., Esq., who sent living plants to the Clapton Nursery, where it was purchased by Messrs. Lucombe, Pince, and Co. "Hoya imperialis requires a strong rich soil in order properly to bring out its numerous large thick flower-trusses, which are produced from different parts of its twining stem. We have used a compost of equal parts of loans, rotten leaves, and peat, with some flakes of dry half-decayed dung intermixed, and a liberal supply of sand and broken crocks blended with the whole. The plant which we exhibited was trained round a low circular trellis, not exceeding three feet in height, and independently of the expanded truss from which the drawing was made, there were several others in different stages upon it. We have also one planted in a corner of the stove, which is twining round n single wire over the path, and upon this there are trusses of flowers ready to expand, having eleven flowers on each. This I think will be found to be the best method of treating it, for with its long pendant bunches of large flowers, overhead, it is a most striking object. Each individual flower lasts a very long time in bloom, and is highly fragrant in the evening and all the night. I send you herewith some of the dried juice of Hoya imperialis, it hardens almost directly after being taken from the plant, and flows so copiously from the wound that I cannot help thinking it may be found available for some useful purpose."—Pince. I think this will prove to be the Asclepias Sussuela of Roxburgh, from the Moluccas, who has erred in quoting Rumphius' Sussuela," and whose specific name is consequently untenable. His character, brief as it is, sufficiently accords, and he describes the corolla as '`nearly three inches in diameter." It seems to be a free flowering and fragrant. Descr.: A climber of quick growth, with rounded, downy, dark green branches. Leaves opposite, on short, terete, thick, downy footstalks, from six to nine inches long, obovato-lanceolate, acuminate, coriaceous, thick, firm, slightly convex above, but even (not waved), with obscure; patent, rather distant, parallel nerves, dark green above, paler beneath, and downy on the costa. Peduncle extra-axillary, much longer than the leaves, terete, downy; flexuose and pendent, terminated by a very large umbel of from nine to fourteen fragrant flowers! Each three inches in diameter. Pedicels downy. Calyx of five, downy, very obtuse, oval sepals. Corolla rotate, pale purplish, downy, within rich purple-brown glossy, pale in the centre: segments spreading, cordato-triangular, the faux elevated, loose around the column. Staminal crown large, projecting, ivory fleshy, glossy: follicles two-lobed; outer lobe large, oval, compressed, flattened on the back, rather obtuse; the inner forming a sharp erect tooth, as long as the outer lobe. Translation: General Characteristics. Calyx short 5 leaved. Corolla rotate plus or minus high 5-fid, flaps flat or reflexed, with valvate aestivation. Staminal corona of 5 leaflets, leaflets depressed flat and plus or minus adnate to the varticular gynostegium, fleshy,

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inner angle produced into a tooth incumbent upon the anthers. Gynostegium short. Anther membrane terminating with pollen masses affixed at the base, oblong, compressed, contacting, often with the margines transparent. Stigma pointed with papilla in the center, obtuse or somewhat apiculate. Follicles smooth or provided with small appendages, somewhat many winged. Seeds comose. Shrubby or somewhat shrubby from India or the Moluccas rare in Africa, twining climber or prostrate, leaves fleshy or leathery or membranaceous, flowers in umbels, umbels from the axils often multiflowered Decaisne. Hoya imperialis, twining, stems petioles, peduncles pubescent, (leaves very broad and large) obovate-lanceolate leathery briefly acuminate acute veins parallel somewhat obscure with many flowered Umbels, flowers large purple-brown, shinny, lobes of the corolla cordate triangular and the axillary margines especially turned under (revolute) free of the elevated throat, leaflets of the white staminal corona in the center confined compressed together, fleshy, bilobed, exterior lobes thick back flat, interior lobe toothlike, not equal. In Flower Garden 12 (1848) 164. (Paxton’s) Hoya imperialis. This is another handsome plant, introduced by Mr. Low, two years since from Borneo, and exhibited by Messrs. Pince and Co, Nurserymen, Exeter, in flower for the first time, at the June Fete of the Royal Botanical Society. The plant has a fine and noble habit, with handsome foliage. The flowers, six in number radiate at the end of a short stem about eight inches long, and are of a brownish purple colour, with a large raised pure white star-like substance in center; they are glossy and wax-like, about two inches in In Fleur des Serres 4 (1848) 393-394. A copy of Botanical Magazine but in French. In Annales de la Societe Royal D’Agriculture et de Botanique 4 (1848) 413-414. Hoya imperialis. Lindl. (Hoya imperial) Classe Pentandrie. Order Monogynie. Famille naturelle. Asclepiadacees. Tribe Pergulariees. (Voir pour la description du genre, tome II, p. 401 de ces Annales.) Car. Spec. H. imperialis. Lindl. Caule volubili, petiolis costanque tomentosis, foliis coriaceis anguste oblongis ciliatis petiolis costaque tomentosis; foliis coriaceis anguste oblongis ciliatis glabris aveniis apiculatis, apice recurvis, basi-rotundatis vel leviter cordatis; pedunculatis tomentosis pendulis, folia brevioribus; umbellis multifloris; sepalis ovatis obtusis tomentosis; corollae maxime lobis triangularibus steliatim patulis ciliatis, intus laevibus, fauce tomentosa; coronae stamineae phyllis compressis bilobis, lobo interiore acuto subdentato, postico ovato obtuso. Car. spec. H. imperial. Lindl. (Same as Botanical Magazine, preceding, however in French). Nous avons deja, a deux reprises, parle de cette plante remarquable, la premiere fois, en rendant compte de l'article de M.. Lindley, insere sans planche dans le dernier volume du Botanical register 1846, (N° 68 ); la seconde fois, en citant l'article de sir William Hooker sur le meme vegetal, (Annales, tom. III, p. 2t, tom. IV, p. 341.) Cet Hoya est une des plus belles creations vegetales qu'on puisse voir. Roxburg le connaissait deja sous le nom d'Asclepias sussuela et l’avait rencontre aux Moluques. Rumph ne l'avait pas vu, puisqu’il etait aveugle, mais il l'a decrit sous le nom de Corona Ariadnes. Ce nom de couronne d'Ariane meriterait de lui etre conserve, car on ne peut pas trouver une plante, dont une branche fleurie forme mieux a elle seule une veritable

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couronne des plus elegantes et des plus somptueuses. M. Lowe, juoior, la decouvrit de nouveau a Borneo et l'envoya en Angleterre. M. Lindley decrivit d'abord cette espece au moyen d'exemplaires fleuris, conserves dans l’esprit de vin, mais sir William Hooker fut plus heurex; il put faire sa deseription d'apres nature, car la plante parut en fleur a l'exposition du Regents-park, en juin 1848. Sa beaute lui fit decerner incontinent et sans conteste la premiere medaille pour les planles nouvelles en fleur. La figure ci-jointe donne une idee de la richesse de cette vegetation. Une tige forte en forme de liane, cylindrique, verte, charnue mais resistante; les feuilles opposees, oblongues et coriaces, d’un beau vert; de longs pedoncules termines par des ombelles de huit a dix fleurs, celles-ci grandes, en forme d'etoiles pourpres et la couronne staminale dun blanc un peu jaune. Le derriere ou le dessous des petales au lieu d'etre pourpre est d'un jaune un peu fauve, ce qui ajoute encore a la beaute du coloris, deja si riche de cette belle plante. Les details de cette couronne en rapport avec la description donnee plus haut, sont figures separement. Nous rappellerons ici, que le 12 janvier 1846, M. Lowe decrivit dans une lettre datee de Sarawak, comment il parvint a retrouver le Russuela de Roxburg. Il fait connaitre que ce fut sur le territoire de Gumbang, en parcourant les collines, qu'il vit pour la premiere fois cette asclepiadee dont un seul individu grimpait sur un arbre mort et pendait au-dessus d'une riviere. Il etait precisment en fleur. Dans le pays natal, M. Lowe remarque qne la couronne staminale etait d'un blanc d’ivoire pur. Sir William Hooker l'a fait peindre jaunatre. Il est probable que c'est la necessite d'ombrer le blanc, qui aura fait adopter cette teinte et que naturellement ces organes sont blancs. Culture. Une terre fort riche en principes alibiles, du loam ou terre franche argileuse, du terreau de feuilles, de la terre de bruyere sablonneuse, du fumier animal, les premiers reunis par tiers et la tout melange avec un peu de sable. blanc, siliceux et aere par des poteries cassees; tel est le sol dans lequel cette asclepiadee se developpe a merveille. On la place dans une serre chaude ou la pleine terre lui convient encore mieux. Le soir et la nuit, la fleur exhale un parfum des plus exquis. La reproduction se fait par boutures en bache chaude. Translation: (Latin). Stems twining, petioles and midrib tomentose; leaves leathery, narrowly oblong ciliate, glabrous without veins, apiculate, apex recurved, base rounded or slightly cordate; peduncles tomentose hanging down and leaf short, many flowered umbel; sepals ovate obtuse tomentose, lobes of the large corolla triangular, starlike outspread ciliate, inside smooth, throat tomentose, leaflets of the staminal corona compressed bilobed, interior lobe acute somewhat tooth-like, outside ovate obtuse. (French) We already have, has two retaking, speak of this remarkable plant, the first time, while giving account of the article of M. Lindley, insert without board in the last volume of the Botanical register 1846, (no. 68); the second time, while mentioning the article of sir William Hooker on the same plant, (Annals, tom. III, p. 2t, tom. IV, p. 341.) This Hoya is one of the most beautiful plant creations that one can see. Roxburg the connaissait already under the name of Asclepias sussuela and had him meeting to the Moluccas. Rumph had not seen him, since he was blind, but he described it under the name of Corona Ariadnes. This name of Ariadne crown would deserve to be it preserved, because one is not able to find a plant, of which a branch in bloom shape better has it alone a real crown of the more elegant and of most sumptuous. Mr. Lowe, junior,

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discovered it has Borneo again and sent it to England. Mr. Lindley first described this species from bloom, preserved in spirits while living, but sir William Hooker was more happy; he, made his deseription of it after nature, because the plant appeared in flower at the exhibition of the Regent-park, in June 1848. Its beauty made him award incontinent and without dispute premiere medal for the new plants in flower. The enclosed page gives an idea of the wealth of this vegetation. A strong stem in shape of liana, cylindrical, green, fleshy but resistant; the opposite, oblong leaves and leathery of a beautiful green; of long stalked finish by umbels of eight to ten big flowers, in the shape of crimson stars and the staminal crown white dun a little yellow. Below on the underside of the petals instead crimson detre are yellow or a little fawn, which adds to the beauty of the coloration again, already so rich for this beautiful plant. Details of this crown had report with the higher description data, are separately faces. We will recall here, that January 12, 1846, Mr. Lowe described in a letter dated from Sarawak, how he arrived has recover the Russuela of Roxburg. He makes know that it was in the territory of Gumbang, in wandering hills, that it lives for the first time this asclepiadee of which only one individual climbed on a dead tree and hung above a river. It was prominently in flower. In the native country, Mr. Lowe notices that the staminal crown was a white or pure ivory. Sir William Hooker made his painted yellow. It is likely that it is a necessary shade of white, that will have made adopt this hue and that naturally these organs are white. Culture. An earth rich strong part in principles alibiles, of the loam or clay straightforward earth, of the compost of leaves, of the gritty heather earth, of the animal manure, the mixture united by thirds and then all mixed with a little sand,. white, silica and for aeration with broken poteries; such is the soil in which this Asclepiads develops is itself a marvel. One places it in a hot greenhouse or the full earth is even again better. In the evening and the night, the flower exhales a most charming perfume. Reproduction makes himself by cuttings in a hot awning. In Transactions of the Royal Horticulture Society 5 (1850) 80. 4. Hoya imperialis. Lindley, in Botanical Register, 1846, under plate 68. Presented to the Society by Messrs. Veitch and Son, of Exeter in 1848. There are two varieties in cultivation of this noble plant: one with long, flat, sharp-pointed leaves, figured in Botanical Magazine, t. 4397; and the present, with shorter, blunter, and wavy leaves, which is what was originally described. Both have large flowers, greenish on the outside and stained with deep purple all over the inside of the corolla, the coronet remaining straw-coloured, and they are among the finest of the stove-twiners in cultivation. If the purple of the corolla were more brilliant, they would be almost unrivaled. A strong climbing plant, growing freely in a mixture of sandy peat and leaf-mold if placed in a strong moist heat. It is easily increased by cuttings in the usual way, and the flowers freely at different times all summer and autumn. A fine shrub for places where there is plenty of room up the rafters, in the stove, or it may be trained round a trellis in a pot. In Magazine of Botany 15 (1851) 214. Paxton. Hoya imperialis, Imperial Hoya Dr. Lindley does not say too much of this plant when he remarks, in the "Bot. Reg.," 1846,

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fol. 68, that this is the most noble climbing plant we hare ever seen." A cluster of its flowers is indeed one of the most striking objects; the leaves, too, are largo and handsome. It is a Hoya with glossy, fleshy leaves, of a deep, purplish chestnut colour, having the expanded flowers full three inches in diameter! rendered more conspicuous by the ivory white of the central column of fructification. It was discovered in Borneo by H.. Low, jun., Esq., who sent living plants to the Clapton Nursery, where it was purchased by Messrs. Lucombe, Pince, and Co. It requires a strong rich soil, in order properly to bring out its numerous large thick flower-trusses, which are produced from different parts of its twining stem. We have used a compost of equal parts of loam, rotten leaves, and peat, with some flakes of dry half-decayed dung intermixed, and a liberal supply of sand and broken crocks blended with the whole. Each individual flower lasts a very long time in bloom, and is highly fragrant in the evading and all the night.(Bot. Mag., 4397. In the Cottage Gardener 8 (1852) 50. D. Beaton. Hoya imperialis (The Imperial Hoya) This is indeed, a noble plant, worthy of the utmost care and skill of the cultivator! Just and look at the dimensions of one flower now laying before us. It is full three inches across, and the umbel from whence it was taken had ten of the these large flowers upon it. The diameter of the umbel measures eight inches; the stem of the umbel is seven inches long; and the stem of each separate flower is four inches; altogether forming one of the finest umbels of flowers ever seen. The colour of the flower, indeed, is not so bright as a Scarlet Anemone, yet it is very pleasing. The calyx is five-parted, green, and very small for so large a flower. the corolla is also five-parted, forming a star-like appearance, each part is triangular-shaped and turned back a little on the edges; the colour is a reddish chocolate, shaded off in the centre with creamy white, the nectaries, five in number, are large and of a yellowish white. Between each there is a small dark coloured spot. the whole flower is highly polished and glossy, like ivory or wax. The leaves are large as the leaves of the common laurel. They grow in pairs, are of a pale lovely green, and covered with silky down. Now, we would ask, is not this faithful description the picture of a fine, first rate, desirable plant? We think it is; and our readers who have never seen it, and have the means of growing it, ought immediately to procure it, the price being now reasonable. 7s. 6d. Culture.— This fine plant was discovered by Mr. Low, jun., growing wild in the woods of Borneo, in perhaps the hottest climate in the world, consequently it requires a warmer stove in this country than the preceding species. The soil it thrives well in with us is a compost of peat, loam, and leaf mold, made very sandy and well drained. It will thrive better if there is the convenience of a bark-bed to plunge the pot in, in which the plant is growing. In is that situation it will grow rapidly, and flower soon. Our plant is only two years old, and it has at present two umbels in flower, and several others showing; and no doubt would have been much larger had it had a bark-bed to stimulate its growth still more. However, it will encourage those who have not the convenience of a bark-bed to grow it in to cultivate so noble a plant. It strikes easily: for incipient roots may be observed appearing on the stem of the plant. Short shoots with two leaves make the best cuttings: place them singly in thumb pots chiefly of sand, under a hand glass, and a fortnight they will be rooted, and may then be hardened off gradually and repotted. As it is a climbing plant it may either be trained to a trellis, or, which is the best method, may

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be trained along the roof of the stove. In this way it shows off its fine flowers to greatest advantage. It requires moderate supplies of water even in summer, but in winter very little will be sufficient. In Annales Botanices Systematicae 3 (1852). G. W. Walpers. H. imperialis Lindl. — Wlprs. Repert. VI. 751. No. 1. – Hook. Bot. Mag. Tab. 4397. — Van Houtte, Flore des Serres IV tab. 393. 394. — Morren, Ann de la soc. D’agricult. De Grand IV. 413. tab. 226. — Lind. Jour. Of the hort. Soc. V. 81. C. lc. In Tuinbouw Flora 1 (1853) 69-70. W. H. DeVriese. 8. Hoya imperialis Lindl. Het epitheton imperialis duidt genoogzaam aan, dat deze plant van eene uistekende en zeldzame schoonheid moet zijn. Deze Hoya is eene der schoonste van dit geslacht. Zij is afkomstig van Borneo, en is van dear gezonden door Low. In l848 werd deze plant bekroond met den grooten prijs, voor nieuwe planten, bij gelegenheid van de tentoonstelling in Regent-Park. Bij een uitnemend schoon gebladerte, heeft deze Hoya bloemen van 8 cent. in middellijn die eene bruine paarse kleur hebben, terwijl het kroontje, ‘t welk in het midden van do bloem zich bevindt, wit is. De bloemen staan op schermen en worden gedragen door eenen laden, afhlangenden steel. Des avonds en gedurende den nacht geven zij eenen zeer aangenamen geur. De kultuur heeft plaats in boomgrond (bois pourri), bladaarde, mest. Men bindt de planten langs kolommen of standaards, en in horizontale rigting, ten einde de bloemsehermen kunnen afhangen. Vochtigheid en warmte zijn de voornaarden van de kultuur dezer prachtige gewassen. (Afb. Lindl., Bot. Reg. 68. 1846. Hook. Bot. Mag. T. 4. 4397. Fl. D. serr. 1848. 393-4.) Translation: (by Albert Hofman) It Hoya imperialis duidt clearly indicates, that this plant must have a distinctive and rare beauty. This hoya is one of the most beautiful of the genus. It comes from Borneo, sent from there by Low. In l848 this plant was awarded with the first prize for new plants at an exhibition in Regent-Park. This has exceedingly beautiful foliage, and flowers of 8 cm. diameter, with a brown-purple color, whereas the little crown, sitting in the middle of the flower is white. These flowers are in umbels and are supported by a long depending stalk. In the evening and during the night they produce a pleasant odor. The culture takes place in the tree soil, leaf mold, manure. One should tie these plants along columns and standards in a horizontal direction so the umbels ban hang down. Humidity and warmth are the cultural conditions for these gorgeous plants. In Gartenflora (1855) 282. b) Hoya imperialis Lindl. var. rauschii. (Hierzu Tafel 130.) Asclepiadeae. Die prächtige Hoya imperialis mit ihren mächtigen braunen Blumelldolden-blühet gegenwärtig unter der sorgsamen Pflege des Hrn. kraft, Gärtners bein Hrn. Rausch in Schaffhausen, in 2 ver schiedenen Formen. Von diesen ist die erstere, die ächte Hoya imperialis, wie, sie das Botanical Magazine und nach diesem alle andern Journale abgebildet. Die Blumenkrone ist hier brnunpurpur, mit breit-oval-deltaförnigen Lappen, die so lang als breit sind. Die zweite Varintät erhielt Her. Rausch von Mackoy in Lüttich, als Hoya imperialis. Sie hat eine lackroth gefärbte viel hellere Blumenkrone, deren Lappen lanzettlich-deltaförmig, also länger als breit sind. Da Herr Rausch uns die Freiheit genommen, sie nach ihm zu nennen. ' Gehört als Schlingpflanze zu den schönsten Zierden des Orchideenhauses. Kultur.

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Grtfl. II. Band S. 350. Translation: The splendid Hoya imperialis, with its immense brown flower umbels, at present flowers under the attentive care of Herr. Kraft, gardener with Herr Rausch at Schaffhausen (Switzerland) in 2 different forms. Of these the first is the true Hoya imperialis, as portrayed by the Botanical Magazine and after that all other journals. The corolla is in this case brown-purple, with broadly deltoid lobes, which are as long as they are broad. Mr. Rausch obtained the second variety from Mackoy at Lüttich, a Hoya imperialis. It has a lacquered-red coloured much paler corolla, its lobes lanceolatedeltoid, which are therefore longer than broad. Since Herr Rausch sent this form to us, so we have taken the liberty of naming it after him. As a vine it is one of the most beautiful ornaments of the orchid house. In The Cottage Gardener and Country Gentlemen's Companion (1856) 78. Hoya imperialis. Presented to the Society by Messers. Veitch and Son, of Exeter, in 1848. There are two varieties in cultivation of this noble plant: one with long, flat, sharp-pointed leaves, figured in Botanical Magazine, t. 4397; and the present, with shorter , blunter, and wavy leaves, which is what was originally described. Both have large flowers, greenish on the outside and stained with deep purple all over the inside of the corolla, the coronet remaining straw-coloured, and they are among the finest of the stove-twiners in cultivation. If the purple of the corolla were more brilliant, they would be almost unrivaled. A strong climbing plant, growing freely in a mixture of sandy peat and leaf-mold if placed in a strong moist heat. It is easily increased by cuttings in the usual way, and the flowers freely at different times all summer and autumn. A fine shrub for places where there is plenty of room up the rafters, in the stove, or it may be trained round a trellis in a pot. (Horticultural Society’s Journal.) In Flora Indiae Bataviae 2 (1857) 527. F.A. W. Miquel. 37. Hoya imperialis Lindl. Caulis volubiles, petioli cum costs tomentosi, folia e basi rotundata vel leviter cordata anguste oblonga apiculo recurvo terminate, coriacea, avenia, glabra, ciliata, pedunculi tomentosi penduli folio breviores, umbellae multiflorae, sepala ovate obtuse tomentosa, corollae maximae laciniae triangulates stamineae patulae ciliatae, intus laeves, fauce tomeutosa, coronae stamineae phylla compressa sursum acuta subdentata, postice ovata obtuse. Lindl. Bot. Reg. 1846 adn. Tab. 68.Hook. bot. Mag. tab. 4397. Van Houtte Fl. D Serr. IV. Tab. 393, 394. Morr. Ann. De Gand IV. P. 413, tab. 226. Lindl. Jour. Hortic. Soc. V. p. 84 cum icone. Borneo. Translation: Stems twining, petioles along with the midrib tomentose, leaves at the base rounded or slightly cordate narrowly oblong with the apical end recurved, leathery, no veins, glabrous, ciliate, peduncle tomentose, short at the foot of the blade, multiflowered umbel, sepals ovate obtuse, tomentose. Leaflets of the large corolla staminal triangular, outspread ciliate, inside smooth, throat tomentose, leaflets of the staminal corona compressed upwards somewhat dentate, rearwards ovate obtuse. In Manual of Cultivated Plants. L. H. Bailey. H. imperialis, Lindl. Lofty climber, with puberulent stems and foliage: lvs. elliptic or 1inear-oblong, obtuse but with a short point: fls.

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Immense (2-3 in. across), leathery, dull purple, somewhat pubescent near the white crown, the segments triangular-acute: umbels drooping on long peduncles; follicles 9 in. long. E. Indies. M. 4397. F.S. 4:393 -4.—A noble Hoya, requiring very rich soil and a rather high temperature. Although naturally a very tall climber, it can be made to flower in pots when 3 to 4 ft. high. In Annales Botanices Systematicae 5 (1858) 506. G. W. Walpers. Icones addendae. Hoya imperialis Lindl. var. Rauschii. ( Regel Gart. Fl. 1855. P. 282. tab. 132. ( Corolla pallidiore, laciniisque ejus lanceolato-deltoideis. Translation: Corolla pale, lobes the same lanceolate deltoid. In The Garden 18 (1880) 355. “The Hoyas or Honey Plants”. Z. B.. Hoya imperialis is a magnificent species, with leaves from 6 in. to 9 in. in length, and large purplish flowers, each about 3 in. in diametre, produced in umbels of about from nine to fourteen flowers. These are very fragrant at night, and will last a long time in water. In The Flora of British India (1883) 59. J. D. Hooker.29. H. imperialis, Lindl. In Bot. Reg. 1846, t. 68 in note, and in Jour. Hist. Soc. V. 80, with woodcut; leaves 6-9 in. elliptic or linearoblong obtuse apiculate very thick puberulous or glabrate beneath; peduncles and pedicels long stout, sepals rounded-ovate, corolla 2 ½-3 in. diam., follicles stout woody. Hook. Bot. Mag. T. 3397 *, copied in Fl. Serres, iv. T. 393: Ann de Grand. 1848, t. 228. ? H. speciosa, Dcne. In DC. Prodr. Viii 635. ? Asclepias Sussuela, Roxb. Fl. Ind. Ii 31, excl. syn. Malacca, Maingay.(Distrib. Borneo, ? Amboina. A lofty stout climber. Leaves 1 ½- 2 ½ in. diam., rarely acute, shining, base often narrowed and cordate, midrib very stout, nerves quite horizontal; petiole ¼- ½ in. Peduncles 310 in.; pedicles 2-3, and sepals pubescent. Corolla leathery dull purple, puberulous near the corona, lobes triangular acute. Coronal-processes white, laterally compressed erect, turgid, obtuse, inner angle shortly produced into a subulate point. Follicles 9 in. long by 1 ½ in. diam. Straight, lanceolate, finely acuminate, pubescent; pericarp thickly coriaceous, endocarp hard. Seeds ¼ in long. * Numbering error, should be 4397. In The Journal of Horticulture and Cottage Gardener (1883) 254. Hoya imperialis. “E.M.” asks us “Which is the handsomest plant to cultivate and train in a trellised pot?” This is a wide question, and we can only reply that the plant so cultivated and trained which we have never seen surpassed was a Hoya imperialis, and of that plant we here publish a portrait. It was exhibited by Messers. Lucombe, Prince, & Co., of Exeter. This beautiful flowering climber was thus noticed by Dr. Lindley at the time of its first introduction in 1846:— "Imagine a true Hoya, with woolly stems, leaves 6 inches long, and clusters of the most magnificent flowers, forming a diadem of ten rays; each flower fully 3 inches in diameter, and with the delicate texture of the common Hoya carnosa, and you will have some notion of this superb species. In Mr. Lowe’s letter from Sarawak, dated January 12th, 1846, we have the following account of its discovery:— “On the next day, when in the territory of the Gumbang Dyaks, I found another curious plant, belonging to Asclepiads; it is an epiphytic climber; there was but one individual, growing from the docked part of a tree, also overhanging the river. The

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flowers are large and in umbels; the leaves are leathery; and the stem abounds in a white, perhaps acrid, juice. The contrast between the purple of the petals and the ivory white of the parts of fructification renders it highly beautiful. It requires a strong rich soil to sustain fully its numerous large flower-trusses, which are producer throughout the lengths of its twining stem. At Kew they used a compost of equal parts loam, decayed leaves, and peat intermixed with sand and broken crocks mixed liberally throughout. Each flower lasts a long time without fading, and is highly fragrant throughout the evening and night. ((Botanical Register (Botanical Magazine.)) In The Illustrated Dictionary of Gardening 2 (1884) 156. G. Nicholson. H. imperialis (imperial).* fl. Reddish-brown, very large, about 3in. across, waxy; umbels 8in. to 9in. in diameter, and eight to ten-flowered. June. L. 6in. long, slightly tomentose, light green. Borneo, 1847. A very noble plant. (B. M. 4397.) In Sukkulenten (1892) 16. Rumpler & Schumann. 3. Hoya imperialis Lindl. Kaiserliche Wachsblume. Die langen verästelten, stielrunden Triebe sind in der Jugend filzig. Die Blätter werden über spannenlang, sie sind oblong-lanzettlich, kurz zugespitzt, fleischig und kurz gestielt. Die lang gestielte, lockere, hängende Dolde besteht aus 8—10 Blüten, deren Blumenkrone dunkelbraun ist, während bie wachsartige Corona weisz ist. Translation: Imperial wax-flower. The long branched, instincts stalk-round is small in the youth. The leaves become long over time, they are oblong-lanceolate, sharpened shortly, fleshy and shortly petiloled. That long petiole, seduces, hanging umbel has from 8-10 blooms, its corolla is dark brown, while the waxy Corona is white. Flora of the Malay Peninsula 4 (1903) 565. King & Gamble. 5. Hoya imperialis, Lindl. In Bot. Reg. (1846) under t. 68, in note, and in Journ. Hort. Soc. V. 80 with woodcut (1850). A stout climbing shrub, rooting on trees and probably epiphytic; branches rounded, dark green, puberulous, puncticulate and with papery bark when dry, reaching 5 in. in diam. Leaves fleshy coriaceous, membranous when dry; elliptic or elliptic-oblong, obtuse or shortly acuminate or apiculate at apex, rounded at base or slightly cordate; both surfaces shining when dry, glabrous above, puberulous or glabrate beneath; margins slightly recurred; 6 to 9 in. long 1.5 to 2.5 in. broad; midrib stout and broad, pubescent on both Sides; main nerves many, more than 12, but difficult to enumerate, nearly horizontal, at first straight, then branching anastomosing; petiole .25 to .5 in. long., stout, pubescent. Umbels many-flowered (about 10), on short, often strongly tubercular and cushioned; rachises at the ends of stout pubescent axillary peduncles 3 to 10 in. long; pedicels stout, pubescent, 2 to 3 in. long; buds large, rounded above, produced below in 5 short recurved angles; flowers with corolla dark purplish-brown above, cinnamomeous beneath, the corona white or pale yellow Calyx membranous, puberulous without, scurry within; lobes ovate-ciliate, 2 in. long; scales minute, triangular. Corolla leathery, 2 to 3 in. in diam., the puberulous tube a little shorter than the ovate-triangular acute lobes, which are at first much incurred, afterwards spreading. Corona of 5 laterally compressed processes; the lower lobes erect, obtuse, deeply 2-cleft and pubescent in the clefts the upper-lobes also erect-subulate-acuminate. Staminalcolumn short, broad; anther-cells broad; appendages membranous, broadly winged, acuminate; pollen-masses flattened, oblong, truncate at both ends, .075 in. long, attached

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by short slender caudicles to the thick, trigonous, horny, brown, pollen-carriers. Style-apex pentagonal, the angles rounded below, ridged above, the ridge joining in an acuminate cone. Follicle I, lanceolate-acuminate, 8 to 9 in. long,. 1 in. broad; pericarp leathery, opening flat, at first puberulous; afterwards glabrous, striate when dry. Seeds obovate, flask-shaped, .2 in. long, trigonous above, truncate at tip and bearing a silky white coma 2 in. long; testa corky, pale brown, albumen copious; cotyledons thick, ovate, cordate at base, .1 in. long; radicle .075 in. long, thick. Hook. Bot. Mag. t. 3397*,-copied In Fl des Serres IV. tt. 393, 394 ; Ann. de Gand (1848). t. 228, Jour. Hort. Ser. 3. XXV. 485; Miq. Fl. Ind. Bat. II. 527; Hook. f. Fl. Br. Ind. IV 59. Hoya splendens, Maingay MS. Asclepias Sussuela, Roxb. Fl. Ind. II 31 (excl. syn.)? Malacca: Maingay (K.D.) 1135.(Distrib. Borneo (Beccari 4001, Motley 1041).. *correct is t. 4397. In Flora of the Malay Peninsula 2 (1923) 399. H. N. Ridley. (16) H. imperialis Lindl. Bot. Reg. Under t. 68; Bot. Mag. t. 3397*; King, l.c. 564. A shrubby climber. Leaves fleshy, elliptic or oblong, short acuminate base rounded; nerves horizontal, numerous; 6 to 9 in. long, 1.25 to 2.5 in. wide; petioles .25 to .5 in. long. Umbels of about 10 flowers on a short rachis; pedicels 3 in. long; peduncles 3 to 10 in. long. Calyx-lobes .2 in. long. Corolla spreading, brown purple, 3 in. across, whitish outside, tube shorter, lobes ovatetriangular acute. Corona white, lower lobes erect, obtuse, 2-cleft; upper lobes subulate acuminate. Follicles 1 lanceolate acuminate, 8 to 9 in. long, 1 in. through. Hab. Very rare, growing in mangroves in Borneo; easily known by its very large purple flowers. Malacca (Maingay) Distrib. Borneo. corolla-lobes spreading, star-like. Flowers over 1 in. across, deep purple……… (16) H. imperialis. * numbering mistake, should be 4397. (Repeating Hooker’s and King’s mistake) In Plantae Elmerianae (UC Press) “Borneenses” 15 (1929) 257-258. E. D. Merrill. Hoya R. Brown. Hoya imperialis Lindl. in Bot. Reg., n.s. 9; sub pl. 68. 1846. No 20629, near Tawao. Suffrutescent and rambling over dense thickets near tidal streams, the flowers 2 to 3 inches across when spread, the calyx and corolla outside green, the latter dull red inside. Endemic? Lindley's species was described from Bornean material, and the specimen cited apparently agrees with the original description and with the illustration in Curtis's Botanical Magazine, pl. 4397. In my Enumeration of Bornean Plants (p. 506), I reduced it to Hoya sussuela (Roxb.) Merr. Interpret. Herb. Amb. 438. 1917, which may be the correct disposition of it; unfortunately no Moluccan material is available to me at this time for purposes of comparison. In The Cyclopedia of Gardening (1930) 1613. L. H. Bailey. H. imperialis, Lindl. Lofty climber, with puberulent stems and foliage: lvs. Elliptic or linear-oblong, obtuse but with a short point; fls. Immense (2-3 in across), leathery, dull purple, somewhat pubescent near the white crown, the segms. Triangular-acute: umbels drooping on long peduncles: follicles 9 in. long. E. Indies. B. M. 4397. F.S. 4:394. R.H. 1900:576. J. H. III 55:443. G. 7:607.(A noble hoya, requiring very rich soil and rather high temperature. Although naturally a very tall climber, it can be made to flower in pots when 3 to 4 ft high.

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In A Handbook of Succulent Plants (1954). (Reprint 1960). H. Jacobsen Hoya imperialis Lindl. E. India …. Freely growing trailing shrub; stalk, petioles, leaf ribs and pedicels covered with felt; l. tough-leathery, elongate, minutely acuminate, recurved at the tip, rounded at the base, or slightly cordate, smooth; f. umbels pendent. fl. 5-7cm across, dark purple, greenish outside. Beautiful, freely growing species, suitable for planting in warm house. Hoya imperialis var. rauschii. Hort. Variety with lighter varnished-red flowers. In Dictionary of the Royal Horticulture (1965) 1st Edition 1951. H. imperialis. Tall climber with downy stems and l. elliptical or linear-oblong obtuse with a short point, 6 to 9" long, slightly hairy. fl. Reddish-brown or dull purple, about 3 in. across, waxy, 10-fid. drooping on long peduncles. Borneo. 1847. B.M. 4397. Needs rich soil and a rather high temperature. In Exotica III (1976). A. B. Graf. H. imperialis (Malakka, Borneo) “Honey Plant”; tall robust climber with felty stems and elliptical, leathery shiny leaves 15-22 cm. long lightly downy, margines wavy; waxy flwrs. Reddish brown, about 8 cm. across, the crown cream-white in pendent clusters, needs rich soil and high temperatures. In Complete Handbook of Cacti and Succulent Plants. Clive Innes H. imperialis, Lindl. A trailing species with stems, leaf ribs and stalks covered with felt. Leaves very tough, elliptic, somewhat downy. Flowers very beautiful on pendent umbels, darkish purple with greenish centered. In Exotic Plant Manual. A. B. Graf. H. imperialis. Same as in Exotica III (1976) below. In Exotica III (1978) 1631. A. B. Graf. Hoya imperialis (Borneo), tall climber with downy stems and elliptic shiny leaves lightly downy, margines wavy; flowers with cream corona in reddish-brown, waxy flowers to 7 cm. across. In Malayan Nature Journal 30 (1978) 503. “The Peninsular Malaysian Species of Hoya” R. E. Rintz. Fig. 18. Hoya imperialis Lindl. a) section of stem; b) corona in side view; c) corona in top view; d) flower in median section; e) twin-pollinia. From F. C. Fogh in 19.3.1941, liquid collection. 14) Hoya imperialis Lindley, Bot. Reg. t. 68 (1846). Type: Borneo (?) (not seen).-FIG. 18. Distinguishing Features. STEMS very thick and smooth. LEAVES fleshy, oblong with obtuse or shallowly cordate bases; up to 16cm long by 5 - 6cm wide; PEDUNCLE reflexed, rigid (?), 10 - 12cm long. UMBEL positively-geotropic, convex with flexnous, uniform pedicels c 8cm long; 1-10 flowers. COROLLA lobes spreading, fleshy,: very finely pubescent inside; 7 -8cm diam; deep red. CORONA blunt and massive, upper lobe with a conical process; entirely yellow. CORPUSCLE wide, clavate. FOLLICLE c. 23cm long by 2.5cm diam. Ecology: Recorded from mangrove and lowland forests in Johore, Melaka, Perak and Selangor; not common. Distribution: Borneo.

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In Hortus III (1979). & In (1976) 574. H. imperialis, Lindl. Puberulant-stemmed, climber, lvs. Obovate to oblanceolate to 9 inches long, peduncles longer than the leaves, umbels pendulous, loosely few-fld; corolla up to 3” across, lobes purplish but darker towards the base, corona whitish. Borneo. In Ashingtonia 3 (1980) 5/6. 2. Hoya imperialis Lindl. Bot. Reg. (1846), sub. t 68 This has for long been recognized as possessing the largest flower in the genus—no other species comes anywhere near squalling its stature! Like so many other well-known plants with long-standing and accepted names, the title is open to question. In one of the more recent and popular publications on wild plants, Dr. Brian Morley in his Wild Flowers of the World suggests that this should be synonymous to Hoya sussuela: (Roxb.) Merr., a plant discovered by Sir Hugh Low, in 1847 ant subsequently introduced into Europe by him in the mid-19th century. The specific title 'imperialis' has remained constant for many years, and if indeed its popular and dignified epithet is to be sunk into the synonymy of Hoya sussuela, it will probably be a long and equally unwelcomed process, even if valid reasons are on its side. The plant is one of noble and superb appearance, features which emphasize the value of its title Hoya imperialis, and the author certainly had excellent perception. Whichever title is acceptable—the habitat is more or less the same. They are equatorial plants, they could hardly be more so—from parts of Malaysia, Borneo and the Moluccan islands, now part and parcel of Indonesia, and probably this area, with all its numerous islands affords scope for an even larger distribution than has already been disclosed. Incidentally, the late Professor Jacobsen in his 'Lexicon' gives the habitat solely as Peninsula India. It is a robust, vigorous species of climbing habit with somewhat tomentose branches and stems. Leaves are fresh or pale green, leathery, lightly pubescent, elliptic in shape with slightly wavy margins and slender, almost pointed tips, and inclined to be cordate at the base, and smooth, in all 1 6-22cm long. Flowers are long-lived, fragrant and borne in large pendulous umbels. frequently 20-24cm in diameter, each flower being about 7cm across, and as many as 8-14 individual warlike flowers in each cluster. The corolla lobes are fleshy and thick, dark purple or reddish; on the inner surface, and somewhat greenish ore greenish-yellow externally, each lobe with a pronounced pointed apex. The corona segments are cream-coloured and prominent, and these alternate with the corolla lobes. Flowering is mainly in June an July. It does not readily produce seed in cultivation, and it would seem preferable to hand pollinate to try and ensure success In this respect. The seed capsule is in the form of a large 'horn' enclosing numerous brownish seeds, each affixed to a whitish parachute. In cultivation a moist, sunny position is necessary—sun is an essential to flowering! Soil must be completely lime-free-the richer the mixture, the better. Temperatures must not drop below 15°C for safety, even higher is preferable. Propagation is from seeds or by stem cuttings which root fairly easily—but its a matter of first finding someone with a plant and willing to divide it. In Asklepios (1992) 24-26. P. I. Forster & A. J. Liddle. Asclepiad Profile no.. 2. Hoya imperialis Lindl. by Paul I. Forster and David J. Liddle 1) Queensland Herbarium, Meiers Road, Indooroopilly, Old 4068, Australia.

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2) P.O. Box 794, Mareeba, Old 4880, Australia. Hoya imperialis Lindl., Bot. Reg. 32: after t. 68 in note (1846); Lindl. in Hook., Bot. Mag. 74: t. 4397 (1848); Lindl., J. Hort. Soc. 5: 81 (1850); J. D. Hook., Fl. Brit. India 4:59 (1885); Rintz, Malay. Nat. J. 30: 501, t.18 (1978). Type: Borneo. Gumbang Dyaks. (not located by Rintz 1978). Epiphytic liane, latex white. Stems cylindrical, up to 8 m long and 5 mm diameter; internodes up to 15 cm long, with dense indumentum when young, scattered indumentum when old. Leaves petiolate lamina elliptic-obovate, up to 16 cm long and 6 cm wide, fleshy, discolorous; upper surface dark green venation obscure, glabrous or with scattered indumentum on edge; lower surface pale green, with 1517 secondary veins faintly visible, tertiary venation obscure; tip acute; base rounded, cuneate, cordate; petiole up to 11 mm long and 3 mm diameter, ± flattened on top, with sparse indumentum; colleters 1 or 2 at lamina base. Inflorescence pendant, umber convex. Cyme umbelliform to racemiform, up to 13 cm long; peduncle up to 13 cm long, 4-5 mm diameter, with scattered indumentum; bracts ovate, 0.60.8 mm long, 0.6-0.8 mm wide, with scattered indumentum. Flowers c. 2 cm long, 5-6 cm diameter; pedicels 42- 50 mm long, c. 3 mm diameter with scattered indumentum. Sepals lanceolate-ovate, 6-7 mm long, 4.5-5 mm wide, glabrous, ciliate; colleters 1 at each sinus base. Corolla flattened campanulate, pink-red; tube 14-15 mm long, 25-27 mm diameter, glabrous; lobes triangular, 25-26 mm long, 18-20 mm wide, incurved at anthesis, glabrous. Staminal corona c. 1 cm long, 1.8 cm diameter, yellow; lobes c. 6 mm long at outer edge, 10 mm long at inner edge, 4.8 mm wide, 8 mm broad, outer edge rounded and incurred, top incurved and spathulate. Staminal column c. 8 mm long and 10 mm diameter, anther appendages lanceolate-triangular, 2-2.3 mm long, c. 2 mm wide; alar fissure c. 3 mm long. Style-head depressed- globose, c. 5 mm diameter. Pollinarium 2.7-2.8 mm long,1.6-1.7 mm wide; pollinia narrow-oblong, 1.9-2.1 mm long, 0.68-0.7 mm wide, with pellucid germinating mouth on outer edge, corpusculum ovate, 0.81-0.86 mm long, 0.75-0.8 mm wide; caudicles c. 0.23 mm long and 0.14 mm wide, unwinged. Fruit and seed not seen. Fig. 2. Specimens Examined: Malaysia: Malacca, 1867-1868, Maingay 1135 (L); Mersing, Apr.1934, Doux s.n. (SING); Senebal, Aug.1929, Mahmood 15530(SING); Port Dickson, Apr.1924, Handover s.n. (SING). Borneo: Pasin Pandjang, Oct. 1964, Ebener 1170 (L); Bern, Mt. Njapa on Kelai River, Oct.1964, Kostermans21314 (BO); Mt. Kinabalu, ENE Dallas, Mt. spur, Dec. 1931, Clemens 27640 (BO); Tawao, Oct. 1922 - Mar. 1923, Elmer20629(BO). Sumatra: Belawan River, May 1929, Lo'rzing 15743(BO, L). Cultivated: Emerald Creek, Mareeba, LiddlelML222 ( BRI ) . Distribution and Habitat: Restricted to northern Malaysia where it grows in mangroves and lowland rainforests (Rintz 1978). Notes: The original place of publication for this species is somewhat unusual in that the diagnosis and brief discussion is on an unnumbered page directly following t.68. At this stage we have not pursued the problem of whether or not a type specimen actually exists for this species; however, it may be inferred from Rintz (1978) that one is not extant at BM, K or L, and a lectotype may have to be chosen from one of the plates published by Lindley. H. imperialis evidently created a sensation when it was first displayed in flower. Hooker (1848) states Exhibited in full flower at the Horticultural Fete in the Regent's Park Gardens (June, 1848), where it obtained the highest prize given for new plants. It is

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still one of the most beautiful of the large flowered Hoyas, and is a lot easier to flower than some other species like those in the section Eriostemma. Cultivation: This species is easily grown if it is given a well-watered compost, with some organic matter, and plenty of strong, filtered light. It will not tolerate cold in conjunction with compost wetness. Hooker (1848) recommended a compost of equal parts of loam, rotten leaves and peat, with some flakes of dry half-decayed dung intermixed, and a liberal supply of sand and broken crocks blended with the whole. Little seems to have changed since 1848! Acknowledgments: We are grateful to the Director of the Rijksherbarium (L) for loan of material, and the Directors/Curators of Herbarium Bogoriense (BO) and Singapore Botanic Gardens (SING) for enabling us to examine holdings at those institutions. References: Hooker, W. J. (1848). Hoya imperialis Curtis's Botanical Magazine 74: t. 4397. Rintz, R. E. (1978). The Peninsular Malaysian species of Hoya (Asclepiadaceae). Malayan Nature Journal 30: 467-522. In Tropica 4 (1992) 1020 A. B. Graf. Hoya imperialis (Borneo) “Honey plant”; tall climber with silky stems and elliptic shiny leaves lightly downy, margines waxy; flowers with cream corona in reddish-brown, waxy flowers to 7 cm across. Tropical. In Philippine Hoya Species 3rd Ed. (1996) 83-84. R. D. Kloppenburg. Hoya imperialis Lindley in Botanical Register 1846, plate 68. Type Borneo, (evidently from dried material and flower in spirits). A noble climbing plant rooting and rambling along tree trunks and high in coastal trees but also found rooted in the forest litter. It seems to prefer mangrove locations at least in the Philippines on both coasts of the Island of Palawan. Discovered on Palawan inn 1993 by Ted Green and David Cumming. It is a strong grower with large woolly stems , leaves 6 inches long, elliptic-oblong, obtuse or shortly acuminate, undulant deep green, lighter green below, rounded at the base to subcordate, nerves pinnate, many somewhat obscure almost horizontal to the midrib then branching and anastomosing, almost identical in leaf form, color and shape to the trees in which it prefers to grow. Umbels of 6-12 flowers 2.5-3” in diameter flat, with stout pedicels 5-7 cm. long, from equally stout peduncles 7-20 cm. long, both pubescent. Flowers waxy various shades from pale to deep purplish brown with cream to pale yellow coronas. Calyx membranaceous, lobes ovate-ciliate. Corolla lobes spreading, fleshy inside finely pubescent. Corona blunt and massive, inner lobe with a horned process back from apex. References: Rep. Bot. Sys. (1846-1847) 743; Curtis Botanical Magazine 1848:4397 Fl. Des Ser. 6 (1848) 310;; Flower Garden Mag. Paxton 12 (1848) 164; Trans. RHS 5 (1850) 80; The Cottage Gardener 1852:50; Tuinbouw Fl. 1 (1853) 69; Gartenflora 1855: 282; Flora Ind. Bat. 2 (1856) 527; Fl. v. Ned. Ind. 1 (1856); Jour. of Hort./Cott. Gdn. 1863:254; The Garden 18 (1880) 354; Fl. Br. Ind. 4 (1883) 59; Jour. Royal Asiatic Soc. Bengal Br. 75 #2 (1908) 563; Fl Malay Pen. 2 (1923) 369-; Plant. Elm. Borne. 15 (1929) 257; Ann. la Soc. RA et Bot. 4 (1848) 413; Malay Nat. Jour. 30 (1978) 503: Ashingtonia 3 (1980) 5/6; Mag. Bot. 6:214. Herbarium Sheets: Elmer #20629, 1923; Ramos #1230, 1930; Beccari #4001; Maingay #1135, Motley #1041; #20629 near Tawao. All from Borneo. Meanin: imperialis — L. imperialis, Of the empire, or emperor, imperial. Of high

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stature.

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