Hoya Foliage A
Pictorial guide II Foliage of Hoya Species By Dale Kloppenburg & Dr. Monina Siar
22
Hoya andalensis Kloppenburg 2005
From the type description
23
Leaves opposite small, cupped below, ovate 5.0 x 2.2 cm. mostly uniform, medium flat green color above, below much lighter, most leaf pairs turned to one side of the stem. Petioles curved 1 cm. or less long, not grooved above, very small rudimentary glands at attachment to the blade. Measurements of above leaves: apex blunt obtuse, base cuneata. Length: 4.5 - 4.75 cm. Nerve angle 50˚. Width: 1.75 - 2.00 cm.
Hoya australis "Santos #1" Top view
24
Measurements from leaves above: broadly ovate, base broadly obtuse, apex short apiculate. Length: 10.7 cm. Width: 7.0 cm. Petiole: 2.0 cm. long 0.3 cm. diameter. grooved above. Bottom view
25
Petiole: 2.2 cm. long. 0.2 cm. in diameter. Vein/midrib angle: 25º
Hoya australis subsp. tenuipes Forster & Liddle 1978 Specimen from West of Luganville, Esperitu Santo, Vanuatu 2001 Top view
26
Measurements from leaves above: broadly ovate, base cordate, apex short blunt. Length: 12.4 cm.. Width: 9.6 cm. 7-8 pairs of veins pinnate, anastomosing, divided before reaching the margins, 2 basal nerves. Bottom View
27
Petiole: curved 4.4 cm. long, 0.2 cm. in diameter, terete. Vein/midrib angle:34° - 42º.
28
Hoya bilobata Schlechter 1906
From they type description well leaved; leaves briefly petioled broadly elliptical or somewhat orbicular, obtuse, glabrous, leathery textured, 1.7-2.2 cm. long, 1.3-1.8 cm. long in the middle, petiole about 0.3 cm. long, fleshy; Measurements of above leaves: ovate apex and base obtuse, leaves uniform. Petiole 0.25- 0.4 cm. long, grooved above. Vein/midrib angle: 50-70° Length: 1.5 – 1.7 cm. Width: 1.15 – 1.20 cm.
29
Hoya brevialata Kleijn & Donklaar
From the type description Leaves suborbicular to elliptic, light-green, yellowish to bright red when very exposed. Petioles 0.5-1 cm. long. Measurements from leaves above: ovate, apex and base obtuse, nervation pinnate.
30
Petiole: 2.3 – 2.s cm. long, 0.15 cm. in diameter. Vein/midrib angle:20º. Length: 4.8 – 4.9 cm. apex rounded, blunt base narrowed. Width: 2.1 – 2.3 cm. Nerves nearly obscure in living material. Actually pinnate with 5-7 nerves per side, branched and rebranched. Petioles are grooved above.
31
Leaves from plant grown in Fresno California, USA
Length 5.2 cm. – 4.6 cm., widest 3.0 cm. 0 3.4 cm. Pinnate venation. Vein/midrib angle: 46°
32
Peduncle: curved 0.5 cm. long, 0.2 cm. in diameter
Length 5.2 cm. – 4.6 cm., widest 3.0 cm. 0 3.4 cm. Pinnate venation. Vein/midrib angle: 46° Peduncle: curved 0.5 cm. long, 0.2 cm. in diameter
33
Hoya sp. burtoniae Kloppenburg
This is not that species but claimed to be so by Burton.
34
From they type description Leaves beautifully colored, opposite, remote, thick or leathery with revolute margins, narrowly oblong lanceolate or oblanceolate, rigid, velvety, densely granular underside a little pubescent and granular 2.5 cm – 12 cm. long above the middle 3 cm. wide, base acuminate with apex obtuse or subacute midrib prominent, lateral nerves not prominent, petiole 0.8 – 2.5 cm. long. Measurements from above leaves (non typical): veins alternate – opposite, pinnate Petiole: 1.0 cm. long 0.2 cm in diameter. Vein/midrib angle: 37º. Length: 5.0 – 5.4 cm. Width: 1.9 – 2.0 cm.
35
Hoya browniana Koorders 1911. Type (BO)
36
From the type description Leaves opposite (but not always, rarely one aborts becoming pseudo-alternate), oblong, glabrous, fleshy, dried led-grey, with the base acute, with the apex acute-acuminate, distinctly triple-nerved, 14-20 cm. long and 6 - 7½ cm. wide. Petioles 2 - 2½ cm. long, thickened, glabrous. It appears this species is palmately nerved (nerves arise at the base). The petioles appear as if they may be grooved but I am not certain.
37
Hoya buotii Kloppenburg 2002
From the type description. No description only a photo of the foliage. The typical leaf is elliptic-lanceolate with a long acuminate apex, base narrowly rounded. Blade is 10 cm. long x 2.5 cm. wide near the base. The petiole is grooved above, 1 cm. long x 0.09 cm. in diameter. Nerves are pinnate 7 or so on each side, at from a 30º to nearly perpendicular angle to the midrib; anastomosing. Measurements of above leaves: Petiole: 1.0 1.2 cm. long, 0.16 cm. in diameter Length: 6.9 – 11.0 cm. Width 2.0 – 2.8 cm.
38
Hoya camphorifolia Warburg 1904
From the type description pedicels (I believe this was meant to be petiole) 7 mm long 1.5 mm thick, leaves parchment-like glabrous ovate-elliptic or elliptic 6 - 8 cm long 3 - 3.5 cm wide tip 39
somewhat acuminate acute base rounded, veins on both sides 2 - 3 ascending (including 2 - 3 basal) not at all reaching the tips, tertiary nerve reticulations distinct. Measurements of above leaves: Base narrowly rounded, apex acute, nerves 3 basal, triplinerved anastomosing. Pinnate nerves, anastomosing, 2 distinct pairs, nerves lighter colored above. dark below. Petiole: 1.0 - 1.5 cm. long 0.2 cm. in diameter. Vein/midrib angle: 21°. Length: 8.1 cm. – 8.7 cm. Width: 2.9 cm. – 3.4 cm.
40
Hoya carnosa R. Brown 1802
From the type description Leaves ovate-oblong fleshy.
41
Measurements of above leaves: Base rounded, apex acuminate to acute, nervation pinnate. Nerves opposite- alternate 7-8 pairs. Vein/midrib angle: 48º. Petiole: 1.5 -1.7 cm. long, 0.1 cm. in diameter. Grooved above Length: 8.3 – 8.6 cm. Width: 4.0 – 4.2 cm.
42
Hoya davidcummingii Kloppenburg 1995
43
From the type description Petiole: twisted (with slight curve) round, not grooved, with a small gland at upper blade attachment. 1 cm. long by 0.18 cm. in diameter. Leaf Blade: spreading lanceolate-elliptic, upper surface deep green even olivaceous, glabrous with fine granulation’s, edges rounded and slightly thickened, lighter green below: apex mostly acute turning down, base cuneate, overall thickened and coriaceous, veins not visible but pinnate, 5-7.5 cm. long, widest about the middle 2-2.5 cm. Measurements of above leaves: Petiole: 1.4 cm. long, 0.15 cm. in diameter. Vein/midrib angle: 45°. Length: 5.9 – 7.5 cm. Width: 2.1 – 2.5 cm. Nerves faint, pinnate/
44
Hoya densifolia Turczaninow 1848
From the type description 45
leaves somewhat sessile membranaceous densely imbricate elliptic, with short cordate sinus, slightly acute penninerved reticulate glabrous. Measurements of above leaved: Base obtuse to cordate, apex acute, nerves pinnate; alternate opposite. Petiole very short, 0.2 cm. in diameter. Vein/midrib angle: 42º. Length: 5.9 – 6.5 cm. Width: 2.9 – 3.0 cm.
46
Hoya dolichosparte Schlechter 1916
47
From the type description leaves oblong-lanceolate obtuse acuminate above (dried) reticulate, below somewhat veinless (5-7 inches long 1 ½ -2 wide) petioles short. Measurements of above leaves: Base somewhat obtuse, apex acute, nerves triplinerved. Petiole: 1.6 cm. long, 0.2 cm. in diameter. Looks grooved above. Vein/midrib angle: 2230°. Length: 9.8 cm. – 10. 5 cm. Width: 5.0 – 5.3 cm.
48
Hoya finlaysonii Wight 1834
49
From the type description leaves oblong-lanceolate obtuse acuminate above (dried) reticulate, below somewhat veinless (5-7 inches long 1 ½ -2 wide) petioles short.
Measurements from leaves above: Apex sharply acute , base narrowed to cuspidate, nervation pinnate anastomosing, but with 3 basal nerves. Petiole: 1.3 – 1.7 cm., long 0.3 cm in diameter. Length: 11.0 cm. – 11.2 cm. Width: 3.4 – 3.8 cm.
50
Hoya heuschkeliana Kloppenburg 1989
51
From the type description Leaves without nerves, petiolate, ovate-elliptic obtuse, base wedge-shaped, fleshy, on the back puberulous both sides punctate (pustulous or granulate) 2 to 5 mm long, in the middle 0.5 to 2 cm. wide, above convex. Petioles curved about 0.2 cm. long.
Measurements from leaves above: broadly ovate, apex broadly rounded, base cuneate, netted pinnate nervation. Petiole: 0.4 – 0.5 cm. long, 0.10 cm. in diameter. Vein/midrib angle: 50º. Length: 4.3 – 4.5 cm. Width: 1.8 cm.
52
Hoya juannguoana Kloppenburg Dorsal surface
53
Ventral surface 54
From the type description A strong dangling, creeping glabrous vine with opposite leaves, obovate to ovatelanceolate, base cuneate, edges entire, with apiculate acute apex. Blades usually somewhat cupped, waxy medium green upper surface, lighter beneath. Pinnate venation 3 veins on either side of midrib not readily visible (in contrast to those of H. finlaysonii Wight) 4 ½” – 6” long 2- 2 ½” widest near the middle or beyond.
Measurements from leaves above: lanceolate, apex narrowing acute-acuminate, base narrow cuneate, nervation pinnate, anastomosing. Petiole: 2.5 cm. long, 0.4 cm. in diameter: grooved above. Vein/midrib angle: 33°. Length: 16.0 cm. Width: 5.3 cm.
55
Hoya kerrii Craib 1911
56
From the type description Leaves opposite, obovate or orbicular, apex bilobed lobes rounded, base broadly wedgeshaped or somewhat rounded 4 to 12 cm. long, 5 to 9.5 cm wide, thickly fleshy, both surfaces glabrous, margins turner under; petiole 5 to 15 mm. long, 5 mm. thick.
Measurements from leaves above: broadly rounded apex cordate, base narrowed, nervation pinnate. Petiole: 1.7 – 2 cm. long, 0.5 cm. in diameter. Vein/midrib angle: 35-42° Length: 9.4 – 10.5 cm. Width: 6.6 – 7.6 cm.
57
Hoya lacunosa Blume
58
From the type description leaves veinless ovate acuminate glabrous laciness above beneath convex, Measurements from leaves above: oblanceolate, apex acute – blunt rounded, base cuniate, pinnate nervation sub anastomosing ca. 9 pairs. Petiole: 0.7 – 1.0 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 27º Length: 4.8 – 6.2 cm. Width: 1.6 – 1.7 cm.
59
Hoya leytensis Elmer ex Burton 1938
60
easurements from leaves above: Variable in shape (of two types). Type 1: apex obtuse, base broad. Type 2: apex narrowly rounded base more acuminate. Nervation obscure but pinnate Petiole 0.8 – 1.0 cm. long, 0.1 – 0.2 cm. in diameter. Vein/midrib angle: 26° Length: Type 1: 2.7 -3.0 cm. Type 2: 3.8 – 5.2 cm. Widest: Type 1: 1.3 cm. Type 2: 1.8 – 2.0 cm.
61
From the type description From Elmer’s description: Leaves coriaceous and subglaucous, more or less persistent, glabrous, opposite or in subwhorles at the ends of abortive branchlets, quite variable in size and shape, entire, narrowly oblong to rotund, most of them are from 2-3.5 cm. long and from 6-10 mm. wide across the middle or widest portion, the rotund ones measure 10 by 12 mm., the normal or the oblong blades equally tapering toward both ends, obtuse to subacute or even rounded for the smaller rotund to elliptic blades which are in the minority, flat, grayish to brown on my dry specimen; midrib quite fine or obscure, plainer on the lower surface, lateral nerves none, only the upper side marked with faint transverse lines; petioles 3-5 mm. long or much shorter, dark brown, crinkle rugose when dry, leaving orbicular flat scares upon falling. From Burton’s description: Leaves coriaceous and subglaucose, glabrous, quite variable in size and shape, entire, narrowly oblong to round, most 2- 3.5 cm. long, 0.6-1.0 cm. wide. The rotund leaves 1.0 by 1.2 cm., the oblong blades tapering at both ends, obtuse to subacute, the round ones of course rounded, flat, midrib quite fine and obscure, more visible on the lower surface, lateral veins none, upper surface marked with fine transverse lines. Petioles 0.3-0.5 cm. long or much shorter, leaving orbicular scars when falling.
62
Hoya maxima (Karsten) Koorders #16204 (BO)
63
From the Type Description No description of foliage.
64
Hoya meliflua var. fraterna Green 1995 Top view
65
Bottom view
66
From the type description Some leaves may attain 40 cm. in length and 15 cm. in width. In contrast, the type usually has leaves that are 10 cm. long by 5 cm. in width. Nerves not easy to discern even in the dried specimens but pinnate, about 6-7 from either side of the midrib at least a 60º angle. Petiole fleshy, curved
Measurements from leaves above: long lanceolate, apex blunt acuminate, base narrowing to cuneate. Nervation obscure pinnate. Petiole: 1.6 – 2.7 cm. long, 0.3 – 0.6 cm. in diameter. Vein/midrib angle: 42-47°. Length: 16.0 - 16.7 cm. Width: 4.6 – 4.7 cm.
67
Hoya merrillii Schlechter 1904 Type: Merrill 2218. Pola, Mindoro Is., Philippines.
68
From the type description
69
Leaves far apart, fleshy; leaves outspread, ovate acuminate, fleshy, both sides glabrous, with 5 prominent nerves when dry, 6-9 cm long, below the middle 5-6.5 cm wide; petiole short fleshy, 1.5-2 cm long.
Measurements from leaves above: There are 3 prominent nerves including the midrib and 2 others not as prominent. There are also a number of looping nerves from the midrib above the palmate ones. The petiole is especially large, 2.0 cm. long here and 0.5 cm. in diameter, corky and not grooved above. Petiole: 1.8 – 2.0 cm. long, 0.5 -0.7 cm. in diameter. Vein/midrib angle: 30º. Length: 11.4 – 12.3 cm. Width: 7.0 – 7.7 cm.
Hoya micrantha Hooker f.
70
From the type description
71
Leaves 2 1/2 -4 by 1 1/4 - 1 1/2 in., midrib and nerves not visible; petiole very short.
Measurements from leaves above: Lanceolate, apex narrow acuminate, base broad cuneate, nerves pinnate. Petiole: 0.7 -0.8 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 30°. Length: 5.5 – 6.7 cm. Width: 1.8 - 2.0 cm.
72
Hoya mindorensis Schlechter 1906
73
From the type description Loosely leaved, with the leaves outspread or spreading, oblanceolate-elliptic, shortly acuminate, glabrous, leathery textured 9-12 cm. long, above the middle 3.5-4.5 cm. wide, with the petiole thick, somewhat terete (circular), 2-2.5 cm. long,
Measurements from leaves above: lanceolate to sub-oblanceolate, apex acute, base obtuse to broadly cuneate, pinnate nervation opposite-alternate. Petiole: 1.3 – 1.5 cm. long, 0.25 cm in diameter. Vein/midrib angle: 50-53º. Length: 9.0 – 12.0 cm. Width: 3.0 – 3.1 cm.
74
Hoya myrmecopa Kleijn & Donklaar
75
From the type description Leaves 2.5--7.5 cm long, ovate to elliptic, acute to acuminate, venation obscure. Petioles 0.5--1.5 cm long.
Measurements from leaves above: Ovate-elliptic, Apex obtuse, base cuneate, nervation pinnate anastomosing. Petiole: 1.0 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 75-120° Length: 5.7 – 6.2 cm. Width: 2.4 – 2.6 cm.
76
Hoya nicholsoniae Mueller 1866
77
From the type description
Glabrous, leaves ovate acuminate, somewhat fleshy, flat, 5-nerved, reticulate-veined; petiole often longer;
Measurements from leaves above: ovate, apex acuminate-acute. base narrow obtuse, with 3 palmate base nerves, above pinnate netted. Petiole: 1.4 – 1.5 cm. long, 0.02 cm. in diameter. Vein/midrib angle: 20° Length: 8.9 – 9.4 cm. Width: 3.0 – 3.8 cm.
78
Hoya nummularioides Constantin 1912
From the Type description: . Leaves fleshy, small, rounded or oval, but with the apex obtuse and apiculate, very pubescent on both surfaces; nerves not visible; blades 9-16 mm., by 12-25 mm. long; petiole pubescent, 2-5 mm. long. 79
Hoya obovata Decaisne 1844
80
From the type description Leaves obovate apexes rounded or rarely emarginate, bases thick attenuate, towards the petioles a callused gland arising from the limb, margins reflexed, both sides glabrous; peduncle short, petiole shorter;
Measurements from leaves above: Ovate with apex slightly indented, nervation pinnate, petioles grooved above. Petiole: 1.7 – 2.0 cm. long, 0.15 – 0.20 cm. in diameter. Vein/midrib angle: 54-58º. Length: 7. 4 – 7.5 cm. Width: 6.5 – 6.7 cm.
81
Hoya obscura Elmer ex Merrill ex Burton 1906
82
Dorsal surface.
83
Ventral surface:
84
From the type description Elmer’s description: Leaves scattered along all the branches, opposite, falling glabrous, smooth and glossy especially on the upper side, entire, diverse in size but not in shape, flat, elliptic to obovately oblong in the longer ones, the smaller subelliptic blades 2 by 3 cm, the others up to 10 cm long and nearly 4 cm wide across the middle or a little above this, gradually tapering to the acute to subacuminate apex, the smaller ones are broadly obtuse at the base, the largest leaves cuneate, sometimes slenderly cuneate, all gradations between these extremes, similarly curing yellowish brown to grayish on both sides; midrib not prominent, very shallowly depressed on the upper face, straight from the base to the apex; lateral nerves faint, 3 to 5 on each half of the leaf blade, quite oblique especially the basal ones, obscurely forked and more or less anastomosing at their ends, reticulations none; petioles 5 to 10 mm long or even up to 1.5 cm in length, likewise glabrous, dirty brown and rugose when dry, caniculate on the upper side. Burton’s description: leaves opposite, glabrous, smooth, size variable, clearly elliptical or obovate or obovateoblong, with leaves longer, small leaves somewhat elliptic 2 -3 cm., others up to 10 cm. long and 4 cm. wide at the middle or smaller above, gradually narrowing to a subacute apex. smaller leaves obtuse at the base, large leaves mostly cuneate, occasionally narrowly altogether gradually cuneate; midrib not prominent, lateral nerves 3-5 on both sides of the costa, basal nerves entirely oblique, at the margins obscure and plus or minus anastomosing; petiole 5-10 mm long or even as 1.5 cm long), glabrous, channeled above.
Measurements from leaves above: Broadly ovate, apex blunt- obtuse, base broadly cuneate. nervation 3 basal and above pinnate. This might be termed triplinerved. Petiole: 1.7 – 2.0 cm. long, knobby, 0.25 cm. in diameter. Vein/midrib angle: 14°. Length: 6.2 – 6.5 cm. Width: 3.4 – 3.6 cm.
85
Hoya obscurinervia Merrill 1923 Isotype #9819 (A)
86
From the type description
Leaves opposite, fleshy, when dry leathery, pale, elliptic to oblong-elliptic, somewhat thin acuminate, bases for the most part acute, obscurely 3 or 5 plinerved, about 10 cm long, 4 to 4.5 cm wide, nerves thin, reticulations irregular; petiole thick, 2 to 2.5 cm long. This species has also been placed into synonymy with H. pottsii Traill so see that file also.
87
Hoya oleoides Schlechter 1913 Type #1970 (B)
88
From the type description
Leaves erect spreading or outspread, petiolate, elliptic obtusely acuminate, leathery, both sides glabrous, petiole short, somewhat thick, above lightly grooved. Leaves 5.5-7.4 cm. long, in the middle 2.3-3.5 cm. wide. Petioles bare, about 0.5 cm. long.
It is difficult to tell the nervation from the type sheet above but I believe it is pinnate with looping anastomosing nervation.
89
Hoya oligantha Schlechter 1913 Type #20176 (B)
90
From the type description
Densely leaved, round, minutely puberulous. Leaves erect-spreading or outspread, shortly petioled, narrowly lanceolate, minutely apiculate or somewhat acute, base rounded, leathery texture, both sides glabrous, petiole lightly grooved above, glabrous, very short. Leaves 2.3-3 cm. long, above the ground (base) 0.5-0.8 cm. wide, petioles very short 1.5-2.5 mm. long.
91
Hoya oreostemma Schlechter 1913 Type #17688 (B)
92
From the type description Loosely leaved, round, glabrous. Leaves spreading or outspread, petiolate. lanceolate elliptic or elliptic, acuminate, base rounded, leathery texture, both sides glabrous, lucid above (shiny), below opaque, petiole glabrous, above slightly grooved. Leaves 6.5-17 cm. long, near the middle 1.5-4.3 cm. wide. Petiole 0.3-0.8 cm. long, bare.
93
Hoya pachyclada Kerr
From the type description
94
Leaves obovate, bases cuneate or somewhat rounded apexes very shortly and obtusely acuminate 7.8 to 10.5 cm. long, 5.2 to 6 cm. wide, thickly leathery, both surfaces having a few sprinkled short hairs, dried costa and nerves on both sides obscure but seen with transmitted light, lateral nerves 2 pairs and basal (or almost at the base) arising acutely, arching and ascending almost reaching apex; petioles thick, above concave, minutely puberulous, 6-12 mm. long.
Measurements from leaves above: Obovate, apex blunt acuminate – obtuse, base broad cuneate. Nervation pinnate. Petiole: 1.1 – 2.0 cm. long 0.35 – 0.4 cm. in diameter. Vein/midrib angle: 37º. Length: 4.9 – 8.0 cm. Width: 2.0 – 4.9 cm.
95
Hoya palawanica (Schltr.) Kloppenburg 1990 Type #834 (B)
96
Sheet #8974 (PNH)
97
From the type description
Leaves spreading or outspread, petiolate lanceolate acuminate, texture leathery, with both surfaces glabrous 13.2-14.5 cm. long, in the middle 3.8-4.2 cm. wide, deciduous with 5 nerves, inner ones nearly reaching the apex with distinct reticulations. Petiole about 1.8 cm. long, 0.6 cm. in diameter, glabrous, grooved above. Peduncle thin, axillary about 0.4 cm. long.
98
Hoya pallilimba Kleijn & Donklaar
99
From the type description
Leaves 3--8 cm long, ovate, acute, without obvious venation, margins on basal half of the leaves thin, pale green. Stems thin, hirsute when young, glabrescent. Petioles 0.5--1 cm. Peduncles more than 5 cm long, hirsute to glabrescent. Umbels concave, positively geotropic, c. 15--30 flowers. Flowers corona yellowish, inner coronal lobe points almost vertically upwards, outer coronal lobes have bilobed extensions with tips that point slightly upwards, corolla dull whitish-red, hirsute on the inside, strongly recurved. Pollinaria caudicles with large wings, retinaculum with very wide groove in the middle.
Measurements from leaves above: ovate, apex acute to blunt, base obtuse, nervation pinnate. Petiole: 0.4 cm. long. 0.1 – 0.2 cm. in diameter. Vein/midrib angle: 45°. Length: 4.7 - 3.8 cm. Width: 2.6 – 2.7 cm.
100
Hoya papuana Schlechter 1913 Type #14400 (B)
101
From the type description
Loosely leafed; leaves lanceolate-elliptic, long acuminate, both sides glabrous, texture thinly leathery, 6 to 8 cm long, below the middle 1 to 2.3 cm wide, petiole short, glabrous, 0.5 - 0.7 cm long. Normally Schlechter mentioned if the petiole was grooved above, so I will assume that it is not grooved.
102
Hoya parvifolia Schlechter 1908 Type #13307 (B)
103
From the type description
Loosely leaved, having aerial roots; with the leaves shortly petiolate oblong or oblongelliptic, very obtuse, both surfaces densely and shortly hispid, 1.2-1.7 cm. long, 0.8-1 cm. wide, petiole hispid about 0.3 cm. long. Again I will assume the petioles are not grooved above.
104
Hoya patella Schlechter 1913 Type #16375 (B)
105
From the type description
Loosely leaved, round, minutely and sparsely pubescent. Leaves spreading or outspread, briefly petiolate, oblong-elliptic, somewhat obtusely acuminate, below pubescent, petiole pubescent, above slightly grooved. Petioles 0.5-0.7 cm. long. Leaves 6.5-9 cm. long, in the middle 2.7-3.4 cm. wide.
106
Hoya pauciflora Wight 1848
107
From the type description
Leaves approximated, fleshy, glabrous, narrow lanceolate, blunt pointed;
Measurements from leaves above: Leaves long narrow, apex acute to somewhat blunt, base cuneate, 3 basal nerves otherwise pinnate. Pedicel: 0.7 – 0.8 cm. long, 0.15 cm. in diameter. Vein/midrib angle: 12º. Length: 0.70 – 8.9 cm. Width: 1.0 – 1.2 cm.
108
Hoya paziae Kloppenburg 1990 Type #5618 (B)
109
#32474 (PNH)
110
Isotype #5618 (US)
111
From the type description
Bene foliates; foliis erecto-patentibus breviter petiolaris ellipticus vel obtusus cum acumine glabris, lucidis, texture tenuiter coriaceis, 6-7 cm. longis, infra medium 2.5-3 cm. latis. Well foliaged, leaves erect spreading, shortly petioled, elliptic or obtuse very glabrous, shiny, of leathery texture, 6-7 cm. long, in the middle 2.5 - 3 cm. wide.
112
Hoya phyllura Schwartz 1931 Type # 339 (K)
113
From the type description
Leaves opposite, blade oblong-elliptic or very slightly ovate-oblong, bases abruptly narrowed, apex long and acute caudate-acuminate, glabrous in the dry state coriaceous shining, nerves very prominent, nervation pinnate veins 3 to 4 on each side disappearing very near apex, curving and anastomosing, nerves narrowly reticulate. Leaf margins entire and very lightly revolute, 8 to 16 cm long, 3.2 to 5.2 cm wide, acumen 1.5 to 2 cm long, in the living state lighter green beneath than above. Petioles robust 1 to 1.7 cm long.
114
Hoya piestolepis Schlechter 1913 Type #19376 (B)
115
From the type description
Loosely leaved, round, glabrous. Leaves spreading or outspread, petiolate oblong or elliptic, acuminate, base briefly cordate, texture leathery, both sides glabrous. Leaves 12-15 cm. long, in the middle 5.5-7 cm. wide. Petiole 1-1.5 cm. long.
116
Hoya pimenteliana Kloppenburg
117
From the type description
Leaf: 6-10 cm long; 2.5-3 cm wide near the middle, apex acute to slightly apiculate, base cuneate to slightly rounded, gland at base, pinnate venation obscure on either surface, Upper surface very finely velvety and deep green, lower surface lighter green, dull with numerous lighter punctations. Leaves rather thick, but not succulent, somewhat leathery.
Measurements from the leaves above: Broadly elliptic, apex blunt, base rounded to broadly cuneate, pinnate nervation. Pedicel: 0.4 -0.5 cm. long, 0.10 cm. in diameter. Vein/midrib angle: 32°. Length: 5.5 – 7.1 cm. Width: 1.3 – 2.1 cm.
118
Hoya plicata King & Gamble
119
From the type description Leaves fleshy coriaceous when dry; ovate-lanceolate, acuminate at apex, acute at base; smooth and glabrous on both surfaces; margins slightly recurved; 3 in. long, 1 in. broad; midrib very faint and then only when dry; remaining nerves invisible; petiole stout, .25 in. long, flattened. Measurements from the leaves above: elliptic, apex acute, base cuneate, nervation pinnate anastomosing. Peduncle: 1.4 cm. long, 0.20 cm. in diameter, grooved above. Vein/midrib angle: 45º. Length: 6.9 -8.0 cm. Width: 2.9 cm.
120
Hoya pottsii Traill 1830 From Loddings Botanical Cabinet 1833
121
From the type description
The leaves are cordate, sharply acuminate, having occasionally a rusty stairs partly spread over them, beneath of a pale whitish green without veins, above light yellowish green, with three distinct veins of ratter lighter colour than the leaf, from which smaller veins occasionally branch off. The petioles are not particularly thick, and less than half an inch long; they are of the same colour as the stems.
122
Hoya puber Blume 1826
123
From the type description
Leaves (small) veinless oval-oblong acute fleshy puberulous,
Measurements from the leaves above: ovate, apes sub acute, base slightly cuneate. Nervation pinnate. Petiole: 0.5 cm. long, 0.210 cm. in diameter. Vein/midrib angle: 54°. Length: 3.8 – 4.0 cm. Widest: 2.5 – 2.6 cm.
124
Hoya pubicalyx Merrill 1918
125
From the type description
Leaves opposite, fleshy, when dry coriaceous, pale and shining on both surfaces and more or less wrinkled, oblong to oblong-obovate, 10 to 14 cm long, 3 to 6 cm wide, base obtuse, apex distinctly acuminate; lateral nerves pinnately arranged, indistinct, about 6 on each side of the midrib, ascending, scarcely more evident than are the lax, indistinct reticulations; petioles about 1 cm long. Measurements from the leaves above: long elliptic, apex acute, base cuneate, nervation pinnate with midrib prominent. Petiole: 1.0 – 1.4 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 32-35º. Length: 14.4 – 15.0 cm. Width: 4.4 – 4.5 cm.
126
Hoya pubifera Elmer #16971 Type (BO)
127
From the type description
Leaves opposite, fleshy, when dry coriaceous, pale and shining on both surfaces and more or less wrinkled, oblong to oblong-obovate, 10 to 14 cm long, 3 to 6 cm wide, base obtuse, apex distinctly acuminate; lateral nerves pinnately arranged, indistinct, about 6 on each side of the midrib, ascending, scarcely more evident than are the lax, indistinct reticulations; petioles about 1 cm long.
128
Hoya pulchella Schlechter 1913 Type #20103 (B)
129
From the type description
Leaves erect, spreading or outspread, petiolate, small, ovate, obtuse, slightly leathery, both sides glabrous, petiole minutely pubescent, above slightly grooved. Leaves 1.7 - 2.2 cm. long, below the middle 0.9 - 1.3 cm. wide, petioles 0.3 -0.5 cm. long, shortly pubescent. Inflorescences shaped like umbels, 4 - 6 flowered, on a very slender, finely pubescent peduncle, 4.5 - 8 cm. long. Pedicels threadlike, thin, bare, 2 -2.7 cm. long.
130
Hoya pusilla Rintz 1978
131
From the type description
Leaves very similar to those of H. lacunosa; fleshy, oblanceolate with acuminate bases and thickened margins; up to 8cm long by 3cm wide.
Measurements from the leaves above: lanceolate-elliptic, apex tapered subacute, base obtuse, nervation pinnate-netted. Petiole 1.0 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 34-44°. Length: 6.0 – 6.2 cm. Width: 2.0 – 2.5 cm.
132
Hoya recurvula Kloppenburg
133
From the type description
Well leaved with pairs of lanceolate foliage, 8.0 - 12.0 cm. long by ca. 3.5 cm. wide, held mostly upright along the stem; flat to slightly curved, apex acute. Petiole terete, at first bronze-green becoming corky with age, curved, no groove above, 2.5 cm. long, about 0.5 cm. in diameter. Measurements from the leaves above: narrowly lanceolate, apex acute sub-apiculate, base cuneate, nerves 3 basal, upward very branched and broken up, netted. Petiole: 1.0 – 1.5 cm. long, 0.1 – 0.4 cm. in diameter, grooved above. Vein/midrib angle: 23º. Length: 6.9 – 12.0 cm. Width: 1.5 – 2.3 cm.
134
Hoya ridleyi King & Gamble 1903 as H. acuta Haworth 1821 Synonym H. parasitica, pallida etc.
135
From the type description Acute leaved—leaves ovate lanceolate acuminate.
Measurements from leaves above: 3 basal nerves, acute angled, pinnate 4 pairs anastomosing. Base cuneata – broadly cuneata, apex tapering acute to apiculate. . Petiole: 2-2.7 cm. long, grooved above. Vein/midrib angle: 27-42°. Leaf length: 10.0 – 14.5 cm. Leaf width: 3.5 – 4.2 cm.
136
Hoya rigida King & Gamble 1908Dorsal surface. Ventral surface.
137
From the description
Leaves ovate or ovate-elliptic, bases rounded or broadly cuneate, apex long acutely acuminate, margins narrowly revolute, 12 to 14 cm. long, 4.5 to 6.3 cm. wide, when dry stiffly leathery, very hard, above black-brown, beneath dark, both sides opaque from the base 5-pli-nerved, nerves on both sides flat-convex, transverse nerves between prominent; petioles very thick, above flat-convex, 1.2 to 2 cm. long.
Measurements from the leaves above: large elliptic apex acute to blunt acute, base cuneate, nervation 3 basal, looping toward apex (anastomosing), petioles large and prominent. Petiole: 4.7 cm. long, 0.7 – 0.8 cm. in diameter. Vein/midrib angle: 17-27º. Length: 13.2 – 14.7 cm. Width: 5.0 – 5.3 cm.
138
Hoya shephardell Hort.
139
Measurements from the leaves above: narrowly elliptic to oblanceolate, apex acute, base cuneate, 2 short basal nerves with prominent midrib, pinnate anastomosing. Petiole: 0.8 – 1.0 cm. long, 0.1 – 0.3 cm. in diameter. Vein/midrib angle: 36-53°. Length: 9.3 – 12.0 cm. Width” 1.7 - 1.9 cm.
140
Hoya sp. abouata From Sak Die Sir Nursery, Thailand
Dorsal surface
141
Ventral surface.
142
Measurements of the (1) leaf above: broadly ovate with short apiculate acute apex. base subcordate, pinnate nervation with prominent midrib. (Clone has rind virus). Petiole: 1.1 cm. long 0.2 cm. in diameter. Vein/midrib angle: 44º. Length: 11.9 cm. Width: 6.7 cm.
143
Hoya sp. DH 54 Palawan, Philippines
144
Measurements of the leaves above: Elliptic, apex blunt, base cuneate, prominent midrib, usually petiole and most of midrib red-purple colored, nervation pinnate with a paid of basal nerves. Petiole: 0.6 – 1.0 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 22-30°. Length: 6.2 – 8.0 cm. Width: 1.7 – 3.2 cm.
145
Hoya sp. IPPS 18888
146
Measurements of the leaves above: Oval to ovate, apex blunt, base obtuse, nervation pinnate. Petiole: 0.6 – 0.7 cm. long, 0.1 – 0.2 cm. in diameter. Vein/midrib angle: small 46-53º lg. 37-46°. Length: 3.9 – 6.9 cm. Width: 2.8 – 2.9 cm.
147
Hoya sp. Mt. Apo
148
Measurements of the leaves above: elliptic, apex blunt, base broadly cuneate, nervation pinnate. Petiole: 0.5 cm. long, 0.1 cm. in diameter, grooved above. Vein/midrib angle: 36-42º. Length: 4.5 – 5.0 cm. Width: 1.8 – 2.0 cm.
149
Hoya sp. Mt. Apo Acanthostemma
Measurements of the leaves above: elliptic, apex acute, base obtuse, nervation pinnate. Leaves uniform. Petiole: 0.5 cm. long 0.1 cm. in diameter. Vein/midrib angle: 40-48°. Length: 3.3 – 3.5 cm. Width: 1.6 cm. 150
Hoya sp. Nagtabon Hill, Palawan
151
Measurements of the leaves above: elliptic, apex sub-apiculate, sun-acute, base obtuse, nervation pinnate. Petiole: 1.2 cm. – 1.5 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 33-36º. Length: 8.6 – 0.2 cm. Width: 3.0 – 3.5 cm.
152
Hoya sp. triplinerved
153
Measurements of the leaves above: elliptic-lanceolate, apex acute apiculate, base broadly cuneate, nervation triplinerved anastomosing. Petiole 1.0 – 2.1 cm. long, 0.4 cm. in diameter. Vein/midrib angle: 23-26°. Length: 10.5 – 12.8 cm. Width: 3.3 – 4.0 cm.
154
Hoya sp. Road back to Bada, Sulawesi
155
Measurements of the leaves above: elliptic to lanceolate, apex acute, base obtuse, nervation pinnate anastomosing. Petiole: 0.7 – 0.9 cm. long, 0.2 cm. in diameter. Vein/midrib angle: 28-32º. Length: 6.2 – 6.4 cm. long. Width: 2.6 – 2.7 cm.
156
Hoya sp. Simas Orchids, Jakarta, Indonesia
157
Measurements of the leaves above: lanceolate, apex acute, base obtuse, Nervation palmate. Petiole: 2.2 cm. long, 0.3 cm. in diameter. Vein/midrib angle: 28-32°. Length: 10.8 – 11.0 cm. Width: 4.0 – 4.5 cm. below the middle.
158
Hoya stoneana Kloppenburg & Siar 2006
159
From the type description: Foliage linear lanceolate 5-5,8 cm. long, 1.5-2,5 cn, at the widest, waxy dark green, on both surfaces, lighter on the underside. Dorsal surface with a few scattered hair cells and more along the edges, ventral surface covered with clear curved hair cells. Apex sub-acute bur mostly obtuse. Petiole: green colored, pubescent, mostly 1 cm. long and grooved above. Vein/midrib angle: 19º
160
Hoya vitellinoides Brink f. Dorsal
161
Ventral surface 162
From the type description
Leaves oblong base shortly obtuse and acutely acuminate on both surfaces very sparsely and very shortly pilose thickly coriaceous, penninerved, lateral nerves 4 to 6 parts on both surfaces conspicuously transverse all over by irregular veins without (?) trichomes 14 to 16 cm long, 7 to 7.5 cm wide, petiole thick 7.5 to 10 mm long.
Measurements of the leaves above: elliptic, apex acute apiculate, base cuneate, nervation pinnate anastomosing and prominent as is midrib. Petiole: 1.0 – 1.2 cm. long, 0.4 cm. in diameter. Vein/midrib angle: 37-46º. Length: 10.0 – 14.0 cm. Width: 5.0 – 5.5 cm.
163
Hoya wayetii Kloppenburg 1993 Type : Maximo Wayet #1989 (CAHUP) Baguio, Benguet Prov. Luzon, Philippines
164
From the type description
Petiole: 0.5-1.5 cm long, terete, thick, glabrous. Not evidently grooved, deep green. Leaf Blade: 5-14.5 cm. long x 1.5-2.7 cm. wide, thick, succulent, convex to nearly flat on the upper glabrous surface, triangular below and lighter green covered with granulose lighter colored punctations. Upper surface deep green with deeper green edging especially in younger leaves. Some rusty tones may be exhibited in newly formed leaves. Blades tapering to both ends, especially narrow toward the petiole which merges with the blade, leaf gland very small almost non-existent.
Measurements of the leaves above: thick v-shaped with dark green edge, apex acute, base narrow cuneate, nervation pinnate. Petiole: 0.8 – 1.2 cm. long, 0.1 cm. in diameter. Vein/midrib angle: 27-52°. Length: 7.7 – 8.8 cm. Width: 1.4 – 1.5 cm.
165
Hoya wibergiae Kloppenburg Dorsal surface
166
Ventral surface 167
From the type description
Leaves opposite tend to be long narrowly elliptic curved along length apex acute to apiculate 6.5 cm. - 11.0 cm. long x ca. 4.2 cm wide, but some smaller and broader, triplinerved with faint anastomosing, only primary veins and midrib visible below surfaces glabrous, margins sharp and slightly curved under, blade not flat. Leaf upper surface with splotchy silvering, petiole terete, corky.
Measurements of the leaves above: elliptic, apex apiculate acute, base obtuse, nervation triplinerved anastomosing. Petiole: 0.7 cm. long 0.4 cm. in diameter. Vein/midrib angle: 16-19º. Length: 13.8 – 13.9 cm. Width: 4.4 – 4.7 cm.
168
